Dima Charpentier, 1825

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 5-6

Dima Charpentier, 1825

Dima Charpentier, 1825: 191 . = Celox Schaufuss, 1862: 201 .

Type species: Dima elateroides Charpentier, 1825: 191 . = Celox dima Schaufuss, 1862: 202 .

Diagnosis. The current generic concept of Dima includes morphologically diverse species from the Iberian Peninsula , the Balkan Peninsula , the Himalayas, East and South East Asia. Species from the Balkan Peninsula, including the type species Dima elateroides , can be recognized by the following combination of characters: body medium-sized, robust, more or less oval, 8.0– 17.1 mm long, 2.3–2.6 times as long as wide, pale brown to blackishbrown ( Figs 1–118 ), antennae weakly serrate, surpassing the pronotal posterior angles by about 1–4 antennomeres, antennomere III shortest (or subequal to antennomere II), antennomeres IV–X 1.5–2.1 times as long as wide, pronotum 1.3–1.6 times as wide as long, moderately convex dorsally, with anterior angles obtuse and posterior angles prominent, acute, more or less distinctly produced outward, tarsomeres III–IV lobate ventrally, and a typically elongate aedeagus with narrow paramera, and apical lobe bearing a more or less developed tooth ( Figs 172–221 ). Species of Dima from the Iberian Peninsula differ in having the antennae with short, bulbous, subtriangular antennomeres, and the Asian species in having much longer antennae, often reaching beyond metacoxae, and in the differently shaped male genitalia ( Schimmel 1996, Schimmel & Platia 2008, and personal observations).

Figures

FIGURES 1 – 14. Habitus images of Dima species. Dima arndti Platia, 2013, 1 — holotype, male, 2 — paratype, female, 3 — nontype specimen, male, 4 — non-type specimen, female; 5 — Dima sp. from Crete, male; D. assingi Schimmel & Platia, 2008, 6 — holotype, male, 7 — holotype of D. peloponnensis Schimmel & Platia, 2008, male, 8 — non-type male from Kilini Mts., 9 — nontype female from Kilini Mts.; D. bruhai sp. nov., 10 — holotype, male, 11 — paratype, female, 12 — paratype, male, 13 — paratype, male; D. dalmatina Küster, 1844, 14 — syntype, male, lateral view. Not to scale.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.

FIGURES 30 – 44. Habitus images of Dima species. 30 — D. fialai sp. nov., paratype, female; D. florinensis Platia, 2012, 31 — holotype, female, 32 — non-type male from Varnous Mts., 33 — non-type female from Varnous Mts., 34 — non-type female from Galičica Mts.; D. fthiotidensis Schimmel & Platia, 2008, 35 — paratype, female, 36 — holotype of D. fokidensis Schimmel & Platia, 2008, female, 37 — holotype of D. scutellaris Platia, 2010, female, 38 — non-type male from Kallidromo Mts., 39 — nontype male from Vardoussia Mts., 40 — non-type female from Vardoussia Mts., 41 — non-type female from Vardoussia Mts.; D. giachinoi Schimmel & Platia 2008, 42 — paratype, female, 43 — non-type specimen, male, 44 — non-type specimen, female. Not to scale.

FIGURES 45 – 59. Habitus images of Dima species. 45 — Dima hirtipennis Platia, 2011, holotype, female, 46 — Dima sp. from Mani; D. hladilorum Schimmel, 1987, 47 — paratype, female, 48 — non-type specimen, male, 49 — non-type specimen, female; D. kozufensis sp. nov., 50 — holotype, male, 51 — paratype, female, 52 — paratype, female, paratype of D. lebenbaueri Schimmel & Platia, 2008; D. lebenbaueri Schimmel & Platia 2008, 53 — holotype, male, 54 — paratype, female, 55 — non-type specimen, male, 56 — non-type specimen, female; D. macedonica Schimmel, 1993, 57 — holotype, male, 58 — non-type male from Galičica, 59 — non-type female from Galičica Mts. Not to scale.

FIGURES 60 – 74. Habitus images of Dima species. Dima macedonica Schimmel, 1993, 60 — non-type male from Ehloec, 61 — female paratype of D. hladilorum Schimmel, 1987 from Galičica Mts.; D. marvani Mertlik & Dusanek, 2006, 62 — holotype, male, 63 — female paratype from Lovčen Mts., 64 — female paratype from Rumija Mts., 65 — non-type male from Rumija Mts.; 66 — D. moraveci Schimmel & Platia 2008, holotype, female; D. neumanni Platia, 2013, 67 — holotype, male, 68 — non-type male from Erimanthos Mts., 69 — non-type female from Erimanthos Mts., 70 — non-type male from Aroania Mts., 71 — non-type female from Aroania Mts., 72 — non-type female from Aroania Mts., 73 — non-type female from Panachaiko Mts., 74 — non-type female from Panachaiko Mts. Not to scale.

FIGURES 75 – 89. Habitus images of Dima species. Dima olympica Meschnigg, 1934, 75 — non-type specimen, male, 76 — non-type specimen, female; D. orientalis sp. nov., 77 — holotype, male, 78 — paratype, female, 79 — paratype, female; 80 — Dima sp. from Xanthi, Greece; D. parnonensis Schimmel & Platia 2008, 81 — holotype, female, 82 — non-type specimen, male, 83 — non-type specimen, female, 84 — non-type specimen, male; D. pecoudi Fleutiaux, 1944, 85 — non-type specimen, male, 86 — non-type specimen, female, 87 — paratype of D. evritaniensis, male, 88 — non-type specimen, teratological male; 89 — D. pelikani sp. nov., holotype, male. Not to scale.

FIGURES 90 – 104. Habitus images of Dima species. Dima pelikani sp. nov., 90 — paratype, female, 91 — paratype, male; D. pelionensis sp. nov., 92 — holotype, male, 93 — paratype, female, 94 — paratype, female; 95 — paratype, female, paratype of D. fokidensis; D. raineri Wurst, 1997, 96 — holotype, male, 97 — non-type specimen, male, 98 — non-type specimen, female, 99 — holotype of D. pindosensis Schimmel & Platia, 2008, male, 100 — male paratype of D. macedonica from Mt. Tomorr, Albania; 101 — Dima sp. from Librazhd, Albania; D. schimmeli Platia & Gudenzi, 2009, 102 — holotype, male, 103 — non-type specimen, male, 104 — non-type specimen, female. Not to scale.

FIGURES 105 – 118. Habitus images of Dima species. Dima schnitteri Platia, 2013, 105 — holotype, male, 106 — paratype, female, 107 — non-type specimen, male, 108 — non-type specimen, female; D. vailatii Schimmel & Platia 2008, 109 — holotype, male, 110 — paratype, female, 111 — non-type specimen, male, 112 — non-type specimen, female, 113 — holotype of D. riesei Platia, 2010, male; D. vonickai sp. nov., 114 — holotype, male, 115 — paratype, female; D. zbuzeki sp. nov., 116 — holotype, male, 117 — paratype, female, 118 — paratype, male. Not to scale.

FIGURES 172 – 181. Aedeagus images of Dima species. 172 — D. arndti Platia, 2013, holotype; 173 — D. arndti, non-type specimen; 174 — Dima sp. from Crete; 175 — D. assingi Schimmel & Platia, 2008, holotype; 176 — D. assingi, non-type specimen from Kilini Mts.; 177 — D. peloponnensis Schimmel & Platia 2008, holotype; 178 — D. bruhai sp. nov., holotype; 179 — D. dalmatina Küster, 1844, non-type specimen from Sutorman, Montenegro; 180 — D. dalmatina, non-type specimen from Vosec, Croatia; 181 — D. dusaneki sp. nov., holotype. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.

FIGURES 192 – 201. Aedeagus images of Dima species. 192 — D. lebenbaueri Schimmel & Platia 2008, holotype; 193 — D. lebenbaueri, non-type specimen; 194 — D. macedonica Schimmel, 1993, holotype; 195 — D. macedonica, paratype; 196 — D. macedonica, non-type specimen from Galičica Mts.; 197 — D. macedonica, non-type specimen from Ehloec; 198 — D. marvani Mertlik & Dusanek, 2006, holotype; 199 — D. neumanni Platia, 2013, holotype; 200 — D. neumanni, non-type specimen from Erimanthos Mts.; 201 — D. neumanni, non-type specimen from Aroania Mts. Not to scale.

FIGURES 202 – 211. Aedeagus images of Dima species. 202 — D. olympica Meschnigg, 1934, non-type specimen; 203 — D. orientalis sp. nov., holotype; 204 — D. parnonensis Schimmel & Platia 2008, non-type specimen; 205 — D. parnonensis, nontype specimen; 206 — D. pecoudi Fleutiaux, 1944, paratype of D. evritaniensis; 207 — D. pecoudi, non-type specimen; 208 — D. pelikani sp. nov., holotype; 209 — D. pelionensis sp. nov., paratype; 210 — D. raineri Wurst, 1997, holotype; 211 — D. raineri, non-type specimen. Not to scale.

FIGURES 212 – 221. Aedeagus images of Dima species. 212 — D. raineri Wurst, 1997, paratype of D. macedonica; 213 — D. raineri, holotype of D. pindosensis Schimmel & Platia, 2008; 214 — D. schimmeli Platia & Gudenzi, 2009, holotype; 215 — D. schimmeli, non-type specimen; 216 — D. schnitteri Platia, 2013, holotype; 217 — D. vailatii Schimmel & Platia, 2008, holotype; 218 — D. vailatii, non-type specimen; 219 — D. vailatii, holotype of D. riesei Platia, 2010; 220 — D. vonickai sp. nov., holotype; 221 — D. zbuzeki sp. nov., holotype. Not to scale.