Dima zbuzeki, Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017

Mertlik, Josef, Németh, Tamás & Kundrata, Robin, 2017, Revision of the flightless click-beetle genus Dima Charpentier, 1825 (Coleoptera: Elateridae: Dimini) in the Balkan Peninsula, Zootaxa 4220 (1), pp. 1-63 : 34-35

Dima zbuzeki sp. nov.

Figs 116–118 , 162 , 221 , 242 , 245 .

Type material. Holotype: GREECE: distr. Kardítsa, Agrafa Mts. , 2.7 km SW of Neraida, firrywood, 1518 m (39°16'45.92"N, 21°40'57.60"E), 31.V.2014, ♂, J. Mertlik leg. ( PCJM) . Paratypes: GREECE: distr. Karditsa, Pindos Mts., Neraida env. [Rafina env.] (39°17'31.29"N, 21°41'58.48"E), 4.–8. VI.2012, 2 ♂♂, 1 ♀, Karel Hodek leg. ( PCJM) ; dtto, 3 ♂♂, 3 ♀♀ (PCPB); dtto, 3 ♂♂, 5 ♀♀ (PCKH); distr. Kardítsa, Agrafa Mts. , 2.7 km SW of Neraida, firrywood, 1518 m (39°16'45.92"N, 21°40'57.60"E), 31.V.2014, 15 ♂♂, 5 ♀♀, J. Mertlik leg. ( PCJM) ; dtto, 11 ♂♂, B. Zbuzek leg. (PCBZ); dtto, 6 ♂♂, 3 ♀♀, P. Brůha leg. (PCPB); Thessalia pref., Karditsa, Pindos Mts., Neraida , Abies forest, singled at night from ground, 39°16'45.92"N, 21°40'57.60"E, 1500 m, 16.VI.2014, 1 ♂, 1 ♀, leg. A. Kotán, Gy. Németh, B. Szelenczey & V. Szénási ( HNHM) ; dtto, 5 ♂♂, 2 ♀♀ (PCBS); dtto, 6 ♂♂ (PCRK).

Diagnosis. Dima zbuzeki sp. nov. is similar in the shape, coloration and pubescence of body, punctation of pronotum and shape of apical lobes of paramera to D. hladilorum ( Figs 47–49 , 116–118 , 137 , 162 , 190 , 221 ). These species differ in the scutellum, which is wider, flat, and only slightly emarginate basally in D. zbuzeki sp. nov. and narrower, convex, and strongly emarginate basally in D. hladilorum . This species differs from D. fthiotidensis and D. evritaniensis by the shiny pronotum with sparse punctation ( Figs 126–127, 130–134 , 162 ) and from D. pelionensis by the pubescence of pronotal lateral margin, which is semi-erect along the whole margin length ( Figs 152–153 ). Dima etoliensis is larger, has longer antennae with longer segments (in females, as no males are known for D. etoliensis ) and more parallel-sided elytra ( Fig. 25 ).

Description. Holotype, male. Body small, 9.9 mm long, 4.0 mm wide, moderately shiny pronotum, more matt elytra. Body brown, with paler, reddish-brown legs, antennae, elytral sutura and margins; pubescence yellowish ( Fig. 116 ).

Head including eyes 0.6 times as wide as pronotum, frons with shallow depression medially; punctation dense and coarse, intervals between punctures shiny posterad, rugose in anterior part; pubescence on head semi-erect, more erect in anterior part of frons, directed forwards. Supraantennal carina short, indistinct. Antennae moderately longer surpassing the posterior angles of pronotum of about four segments; antennomeres II and III short, subequal, less than 1.5 times longer than wide, together about 1.2 times longer than antennomere IV, length ratio of antennomeres II–IV 1.1: 1.0: 1.8, antennomeres IV–X gradually widened apically, about 2 times longer than wide, apical antennomere longest, subacute apically; surface of antennomeres covered with moderately long, semi-erect setae.

Pronotum 1.4 times wider than long, widest at middle, moderately convex dorsally; sides slightly and evenly convex, anterior angles obtuse; posterior angles prominent, sharp, slightly produced outward; punctation moderately dense, coarse, punctures on disc equally distributed; intervals between punctures moderately shiny. Pubescence on pronotal disc moderately long, semi-erect; that on pronotal sides semi-erect along the whole length, gradually longer towards posterior angles, longest and erect in posterior angles ( Fig. 162 ). Prosternum finely, moderately sparsely punctate, with short semi-erect setae; prosternal lobe moderately coarsely punctate, with long semi-erect pubescence.

Scutellum sub-pentagonal, flattened, slightly depressed medially, in lateral view slightly above the plane of elytra, widest at middle; frontal margin slightly emarginate, gradually declined; apex widely rounded, punctation sparse, coarse; pubescence fine, moderately long, semi-erect. Elytra sub-parallel, 3.4 times longer than pronotum, moderately convex, widest behind middle; striae well developed in 4/5 of elytral length, interrupted irregularly at apical part with coarse sparse punctures; surface of interstices flattened, slightly matt, densely and finely punctate; punctures suboval, equally distributed, of the same size.

Aedeagus elongate; paramera long, narrow, its apical lobe long, narrow, subapical tooth robust, obtuse ( Fig. 221 ).

Female ( Fig. 117 ). Like male but slightly paler, suboval, antennae shorter (about 2.5 segments surpassing posterior angles of pronotum), pronotum wider, 1.5 times wider than long, widest at middle. The sclerotized spines of bursa copulatrix as in Fig. 242 .

Intraspecific variability. Body length: 9.0– 11.5 mm. Body coloration is light brown in some specimens ( Fig. 118 ).

Distribution. Greece (Agrafa Mts.; Fig. 245 ).

Etymology. The specific epithet is a patronym in honor of Mr. Bořivoj Zbuzek (Praha, Czech Republic), who collected a part of the type series.

Figures

FIGURE 245. The distribution of Dima species in the Central Greece.

FIGURES 105 – 118. Habitus images of Dima species. Dima schnitteri Platia, 2013, 105 — holotype, male, 106 — paratype, female, 107 — non-type specimen, male, 108 — non-type specimen, female; D. vailatii Schimmel & Platia 2008, 109 — holotype, male, 110 — paratype, female, 111 — non-type specimen, male, 112 — non-type specimen, female, 113 — holotype of D. riesei Platia, 2010, male; D. vonickai sp. nov., 114 — holotype, male, 115 — paratype, female; D. zbuzeki sp. nov., 116 — holotype, male, 117 — paratype, female, 118 — paratype, male. Not to scale.

FIGURES 155 – 162. Pronotal details of Dima species. 155 — D. raineri Wurst, 1997, holotype, male; 156 — D. schimmeli Platia & Gudenzi, 2009, female; 157 — D. schnitteri Platia, 2013, female; 158 — D. vailatii Schimmel & Platia 2008, holotype, male; 159 — D. vailatii, male; 160 — D. riesei Platia, 2010, holotype, male; 161 — D. vonickai sp. nov., holotype, male; 162 — D. zbuzeki sp. nov., holotype, male. Not to scale.

FIGURES 212 – 221. Aedeagus images of Dima species. 212 — D. raineri Wurst, 1997, paratype of D. macedonica; 213 — D. raineri, holotype of D. pindosensis Schimmel & Platia, 2008; 214 — D. schimmeli Platia & Gudenzi, 2009, holotype; 215 — D. schimmeli, non-type specimen; 216 — D. schnitteri Platia, 2013, holotype; 217 — D. vailatii Schimmel & Platia, 2008, holotype; 218 — D. vailatii, non-type specimen; 219 — D. vailatii, holotype of D. riesei Platia, 2010; 220 — D. vonickai sp. nov., holotype; 221 — D. zbuzeki sp. nov., holotype. Not to scale.

FIGURES 234 – 242. The sclerotized spines of bursa copulatrix of Dima species. 234 — D. olympica Meschnigg, 1934, nontype specimen; 235 — D. orientalis sp. nov., paratype; 236 — Dima sp. from Xanthi, Greece; 237 — D. pelikani sp. nov., paratype; 238 — D. pelionensis sp. nov., paratype of D. fokidensis Schimmel & Platia, 2008; 239 — D. schimmeli Platia & Gudenzi, 2009, non-type specimen; 240 — D. schnitteri Platia, 2013, paratype; 241 — D. vonickai sp. nov., paratype; 242 — D. zbuzeki sp. nov, paratype. Not to scale.

FIGURES 45 – 59. Habitus images of Dima species. 45 — Dima hirtipennis Platia, 2011, holotype, female, 46 — Dima sp. from Mani; D. hladilorum Schimmel, 1987, 47 — paratype, female, 48 — non-type specimen, male, 49 — non-type specimen, female; D. kozufensis sp. nov., 50 — holotype, male, 51 — paratype, female, 52 — paratype, female, paratype of D. lebenbaueri Schimmel & Platia, 2008; D. lebenbaueri Schimmel & Platia 2008, 53 — holotype, male, 54 — paratype, female, 55 — non-type specimen, male, 56 — non-type specimen, female; D. macedonica Schimmel, 1993, 57 — holotype, male, 58 — non-type male from Galičica, 59 — non-type female from Galičica Mts. Not to scale.

FIGURES 131 – 142. Pronotal details of Dima species. 131 — D. fthiotidensis Schimmel & Platia, 2008, paratype, female; 132 — D. scutellaris Platia, 2010, holotype, female; 133 — D. fthiotidensis, male; 134 — D. fthiotidensis, holotype of D. fokidensis Schimmel & Platia, 2008; 135 — D. giachinoi Schimmel & Platia 2008, female; 136 — D. hirtipennis Platia, 2011, holotype, female; 137 — D. hladilorum Schimmel, 1987, male; 138 — D. kozufensis sp. nov., paratype, female; 139 — Dima lebenbaueri Schimmel & Platia 2008, holotype, male; 140 — Dima lebenbaueri, non-type specimen, female; 141 — D. macedonica Schimmel, 1993, holotype, female, 142 — D. macedonica, female paratype of D. hladilorum from Galičica Mts. Not to scale.

FIGURES 182 – 191. Aedeagus images of Dima species. 182 — D. elateroides Charpentier, 1825, non-type specimen from Serbia; 183 — D. evritaniensis Schimmel & Platia, 2008, non-type specimen from Timfristos Mts.; 184 — D. fialai sp. nov., holotype; 185 — D. florinensis Platia, 2012, non-type specimen; 186 — D. fthiotidensis Schimmel & Platia, 2008, non-type specimen from Kallidromo Mts.; 187 — D. fthiotidensis, non-type specimen from Vardoussia Mts.; 188 — D. giachinoi Schimmel & Platia 2008, non-type specimen; 189 — Dima sp. from Mani; 190 — D. hladilorum Schimmel, 1987, non-type specimen; 191 — D. kozufensis, sp. nov., holotype. Not to scale.

FIGURES 119 – 130. Pronotal details of Dima species. 119 — D. arndti Platia, 2013, male; 120 — D. assingi Schimmel & Platia, 2008, female; 121 — D. bruhai sp. nov., holotype, male; 122 — D. dalmatina Küster, 1844, male; 123 — D. dusaneki sp. nov., holotype, male; 124 — D. elateroides Charpentier, 1825, male; 125 — D. etoliensis Platia, 2012, holotype, female; 126 — D. evritaniensis Schimmel & Platia, 2008, paratype, female; 127 — D. evritaniensis, male; 128 — D. fialai sp. nov., holotype, male; 129 — D. florinensis Platia, 2012, female; 130 — D. fthiotidensis Schimmel & Platia, 2008, male. Not to scale.

FIGURES 143 – 154. Pronotal details of Dima species. 143 — D. macedonica Schimmel, 1993, male; 144 — D. marvani Mertlik & Dusanek, 2006, holotype, male; 145 — D. moraveci Schimmel & Platia, 2008, holotype, female; 146 — D. neumanni Platia, 2013, female; 147 — D. olympica Meschnigg, 1934, male; 148 — D. orientalis sp. nov., holotype, male; 149 — D. parnonensis Schimmel & Platia, 2008, female; 150 — D. pecoudi Fleutiaux, 1944, male; 151 — D. pelikani sp. nov., holotype, male; 152 — D. pelionensis sp. nov., paratype, female, 153 — D. pelionensis sp. nov., paratype, female paratype of D. fokidensis; 154 — D. raineri Wurst, 1997, female. Not to scale.

FIGURES 15 – 29. Habitus images of Dima species. Dima dalmatina Küster, 1844, 15 — syntype, male, 16 — non-type male from Vosac, Croatia, 17 — non-type female from Vosac, Croatia, 18 — non-type male from Rumija Mts., Montenegro, 19 — nontype female from Rumija Mts., Montenegro; D. dusaneki sp. nov., 20 — holotype, male, 21 — paratype, female; D. elateroides Charpentier, 1825, 22 — non-type male from Serbia; 23 — non-type male from Slovenia, 24 — non-type female from Croatia; 25 — D. etoliensis Platia, 2012, holotype, female; D. evritaniensis Schimmel & Platia, 2008, 26 — paratype, female, Panetoliko Mts., 27 — non-type male from Timfristos Mts., 28 — non-type female from Timfristos Mts.; 29 — D. fialai sp. nov., holotype, male. Not to scale.