Aname marae, Harvey, Frances S. B., Framenau, Volker W., Wojcieszek, Janine M., Rix, Michael G. & Harvey, Mark S., 2012

Harvey, Frances S. B., Framenau, Volker W., Wojcieszek, Janine M., Rix, Michael G. & Harvey, Mark S., 2012, Molecular and morphological characterisation of new species in the trapdoor spider genus Aname (Araneae: Mygalomorphae: Nemesiidae) from the Pilbara bioregion of Western Australia, Zootaxa 3383, pp. 15-38 : 35-37

publication ID	https://doi.org/ 10.5281/zenodo.215484
DOI	https://doi.org/10.5281/zenodo.6168290
persistent identifier	https://treatment.plazi.org/id/039587C5-A449-DA26-5CBF-AB9AFF78FE23
treatment provided by	Plazi
scientific name	Aname marae
status	sp. nov.

Treatment

Aname marae n. sp.

Figs 8 View FIGURES 5 – 8 , 9 View FIGURE 9 , 43–51 View FIGURES 43 – 51

Type material. AUSTRALIA: Western Australia: holotype male, Tom Price powerlines, 4 km NW. of Tom Price, 22°41’10”S, 117°44’56”E, 6 September 2009, dug from burrow, D. Kamien ( WAM T98424 DNA). Paratypes. 3 males, same data as holotype except 5 September 2009, dug from burrow, D. Kamien ( WAM T98427 DNA, T 112543, T 112544); 3 males, same data except 22°40’56”S, 117°45’06”E, 5 September 2009, dug from burrow, D. Kamien ( WAM T98428, T98432 DNA, T98435 DNA).

Other material examined. AUSTRALIA: Western Australia: 2 males, Tom Price powerlines, 5 km NW. of Tom Price, 22°40’36”S, 117°44’19”E, 4 September 2008, dug from burrow, D. Kamien ( WAM T98423 DNA, T98425 DNA); 2 males, Tom Price powerlines, 6.1 km NW. of Tom Price, 22°40’08”S, 117°44’53”E, 4 September 2009, dug from burrow, D. Kamien ( WAM T98430 DNA, T 112542); 1 male, Tom Price powerlines, 8 km NW. of Tom Price, 22°39’17”S, 117°42’59”E, 6 September 2009, dug from burrow, D. Kamien ( WAM T98436 DNA); 5 males, same data except 22°39’28”S, 117°43’01”E, 6 September 2009, dug from burrow, D. Kamien ( WAM T98426 DNA, T98429 DNA, T98431 DNA, T98433 DNA, T98434 DNA).

Etymology. This species is named for Māra Blosfelds, the senior author’s mother.

Diagnosis. Males of A. marae resemble A. barrema Raven, 1985 , A. distincta Raven, 1985 , A. inimica Raven, 1985 and A. ellenae in the shape of the bulb which tapers gradually to the embolus ( Figs 50, 51 View FIGURES 43 – 51 ), but differ in the shape of the embolus which is broadly flattened at the base ( Fig. 51 View FIGURES 43 – 51 ). In addition, A. marae males lack the retrolateral patch of short setae on the pedipalpal tibia ( Figs 50, 51 View FIGURES 43 – 51 ) that are characteristic for A. ellenae ( Figs 41, 42 View FIGURES 34 – 42 ) and also have fewer maxillary cuspules (<100 in comparison to> 100) which do not extend to the maxillary heel ( Fig. 47 View FIGURES 43 – 51 ). Females of A. marae are unknown.

Description. Adult male (based on holotype WAM T98424): medium-sized nemesiid spider (total length 13.4).

Colour: carapace reddish-brown with somewhat darker cephalic region ( Fig. 43 View FIGURES 43 – 51 ), eye region dark brown ( Fig. 46 View FIGURES 43 – 51 ); sternum light brown, darker towards margins, sigilla light brown and labium brown ( Fig. 45 View FIGURES 43 – 51 ); abdomen dorsally dark greyish-brown ( Fig. 44 View FIGURES 43 – 51 ), ventrally light brown; chelicerae very dark brown; pedipalp segments uniformly brown ( Fig. 50 View FIGURES 43 – 51 ); leg I dark reddish-brown with metatarsi and tarsi brown ( Fig. 48 View FIGURES 43 – 51 ), legs II-IV brown.

Carapace: 7.2 long, 5.6 wide; eye group ( Fig. 46 View FIGURES 43 – 51 ) 1.4 wide, 0.7 long; fovea slightly procurved ( Fig. 43 View FIGURES 43 – 51 ).

Sternum and labium: 4.4 long, 2.7 wide; 2 pairs of very indistinct elongated sigilla in posterior half ( Fig. 45 View FIGURES 43 – 51 ); labium wider than long, slightly indented anteriorly ( Fig. 47 View FIGURES 43 – 51 ).

Maxillae: with <100 pin-like cuspules, not extending to heel ( Fig. 47 View FIGURES 43 – 51 ).

Chelicerae: without rastellum, promargin with 11 irregularly sized teeth, retromargin with 3 small teeth proximally.

Abdomen: 6.7 long, 3.7 wide ( Fig. 44 View FIGURES 43 – 51 ). Four spinnerets, terminal segment of posterior lateral spinnerets digitiform.

Pedipalp: length of femur 3.8, patella 2.2, tibia 2.5, tarsus 1.7. Femur with 1 disto-prolateral spine, patella with 2 prolateral spines. Tibia with 2 long ventral spines, and 4 disto-prolateral; with distinct disto-ventral depression without setae (accommodates bulb and embolus) ( Fig. 50 View FIGURES 43 – 51 ); tarsus terminally blunt ( Fig. 51 View FIGURES 43 – 51 ); bulb elongated, globular, tapering into short embolus ( Figs 50, 51 View FIGURES 43 – 51 ), embolus ca. 0.8 long, curved and flattened at base, straight and tapering distally ( Figs 50, 51 View FIGURES 43 – 51 ).

Legs: femur I with 2 dorsal spines and 2 disto-prolateral spines, patella with 2 prolateral spines, metatarsi and tarsi without spines; tarsi and apical half of metatarsi of leg I and II ventrally with entire dense scopulae, tarsi of legs III and IV with divided very weak scopulae. Tibia I with large spur and megaspine, metatarsi incrassate ( Fig. 49 View FIGURES 43 – 51 ). Tarsi, metatarsi and tibiae with numerous dorsal trichobothria of variable length. Paired tarsal claws of legs with two rows of 8–12 teeth each; third tarsal claw reduced and very small.

Leg measurements: length of legs IV>I>II>III. Leg I: femur 5.8, patella 3.5, tibia 4.1, metatarsus 4.2, tarsus 2.5, total = 20.1. Dorsal proximal width of patella I = 1.2, tibial index = 0.16, leg I formula = 2.79. Leg II: femur 5.2, patella 2.8, tibia 3.6, metatarsus 3.8, tarsus 2.3, total = 17.7. Dorsal proximal width of patella II = 0.9, tibial index = 0.14; leg II formula = 2.46. Leg III: femur 4.4, patella 2.5, tibia 2.7, metatarsus 3.6, tarsus 2.1, total = 15.3. Dorsal proximal width of patella III = 1.0, tibial index = 0.19; leg III formula = 2.13. Leg IV: femur 5.8, patella 3.0, tibia 4.2, metatarsus 5.0, tarsus 2.5, total = 20.5. Dorsal proximal width of patella IV = 1.1, tibial index = 0.15; leg III formula = 2.85.

Variation: total length 14.6–18.1, carapace 6.1–7.6 long, 4.7–5.8 wide (n = 10).

Adult female: unknown.

Distribution. Aname marae has only been found near the township of Tom Price in the central Pilbara region of Western Australia ( Fig. 8 View FIGURES 5 – 8 ).

Remarks. As for A. ellenae (see above), most males of A. marae were dug from burrows before the beginning of the wet season (in September) and may have waited there for the first seasonal rainfalls before searching for females.