Perrona ilonae, Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 106-107

Perrona ilonae nov. nom.

Figs 31F 1 –F 2 , 4Q , 5

Clavatula styriaca Auinger — Friedberg 1912: 198, pl. 12, fig. 7 [non Perrona styriaca ( Hilber, 1879) ].

* Clavatula vindobonensis nodosa n. subsp. — Csepreghy-Meznerics 1954: 141, pl. 7, figs 8–9, 11–12 [non ‘Clavatula’ nodosa ( Bellardi, 1847) ].

Pleurotoma n.f.— Bauer 1900: 25, pl. 1, figs 5a–c.

Clavatula styriaca (Hilb.) — Kókay 1966: plate captions, pl. 9, fig. 15 [non Perrona styriaca ( Hilber, 1879) ].

Clavatula jouanneti [sic] vindobonensis Partsch (in Hörnes), (1845) 1856— Strausz 1966: 400, pl. 14, figs 10–11, 14 [non Perrona vindobonensis (Quenstedt, 1884) ].

Clavatula interrupta Br. aff. sophiae (Hoern. et Auing.) — Kókay 1966: 63, pl. 9, fig. 16 [non ‘ Clavatula ’ sophiae ( Hoernes & Auinger, 1891) ].

Clavatula veronicae (Hoern. et Auing.) — Kókay 1966: 64, pl. 9, fig. 17 [non ‘ Clavatula ’ veronicae ( Hoernes & Auinger, 1891) ].

Clavatula (Perrona) vindobonensis nodosa Mez. — Kókay 1966: plate captions, pl. 9, fig. 18 [non ‘Clavatula’ nodosa Bellardi, 1847 ].

? Clavatula gothica Mayer, 1873 — Steininger 1973: 437, pl. 8, fig. 5a [non ‘Clavatula’ gothica Bellardi, 1877 ].

Clavatula vindobonensis nodosa Meznerics 1954 — Bohn-Havas 1973: 1064 [non ‘Clavatula’ nodosa Bellardi, 1847 ].

Clavatula (Perrona) vindobonensis nodosa Csepreghy-Meznerics, 1954 — Pálfy et al. 2008: 105 [non ‘Clavatula’ nodosa Bellardi, 1847 ].

non Perrona (Perrona) vindobonensis nodosa (Csepreghy-Meznerics) — Nikolov 1994: 59, pl. 6, figs 14–15 [= Perrona descendens ( Hilber, 1879) ].

Type material. Holotype: M.61.4367, Szentkút, Mátraverebély ( Hungary), SL: 22.0 mm, MD: 8.1 mm, holotype of Clavatula vindobonensis nodosa Csepreghy-Meznerics, 1954 , illustrated in Csepreghy-Meznerics (1954, pl. 7, figs 8–9), stored in the Hungarian Natural History Museum, Budapest ( Hungary); figs 31F 1 –F 2, 4Q.

Type locality. Mátraverebély ( Hungary), Pannonian Basin.

Type stratum. Shallow marine sand silt of the Sámsonháza Formation ( Dulai & Moissette 2007).

Age. Early/middle Badenian (Langhian).

Etymology. In honor of Ilona Csepreghyné Meznerics (1906–1977), pioneer of Paratethyan mollusc paleontology.

Revised description. Shell medium-sized, moderately slender fusiform with weakly gradate spire and subcylindrical whorls; apical angle 36°. Protoconch not preserved. Teleoconch of nine whorls. Early teleoconch whorls flat-sided with two narrow adapical spiral cords with opisthocline, comma-shaped axial riblets. Suprasutural spiral cord indistinctly beaded by abapical tips of riblets. Sculpture changing on fourth and fifth whorl; subsutural cord swelling into rounded, sharply-delimited, weakly tuberculose subsutural collar, axial riblets at mid-whorl and suprasutural row of beads disappear, five narrow, flattened spiral cords develop below collar, separated by deep, narrow grooves; grooves becoming shallower towards abapical suture. Suture superficial, slightly undulating. Last whorl ~ 65% of total height; subsutural collar broad, bearing round to pointed tubercles, sharply delimited. Broad subsutural ramp weakly concave, poorly delimited by low-set, rounded shoulder, base poorly delimited, moderately constricted. Siphonal fasciole weakly swollen, twisted. Sculpture of narrow, flattened cords separated by deep grooves on ramp, obsolete at mid-whorl, strengthening again below shoulder, extending over base and fasciole. Aperture narrowly pyriform. Outer lip not preserved. Anal sinus wide, moderately deep, asymmetrically U-shaped, with apex at mid-ramp. Siphonal canal moderately long and wide, straight, with shallowly notched tip. Columella obscured by sediment.

Discussion. This species was established by Csepreghy-Meznerics (1954) as Clavatula vindobonensis nodosa . This name, however, is a secondary homonym of the Italian Pleurotoma nodosa Bellardi, 1847 , which is currently placed in Clavatula by Ferrero Mortara et al. (1981) and Zunino & Pavia (2009). Therefore, we propose Perrona ilonae as new name for the Hungarian species.

The holotype is a relatively slender specimen with rounded beads. Additional specimens from Herend ( Hungary), illustrated by Strausz (1966) and Kókay (1966) are slightly broader than the holotype and document considerable variability of sculpture with pointed to spinose nodes at the subsutural collar. All specimens display the characteristic spiral grooves below the subsutural collar. Perrona ilonae is highly reminiscent of Perrona vindobonensis (Quenstedt, 1884) and at first sight it could be an extreme morphotype. Typical specimens of P. ilonae bear prominent, pointed beads on the subsutural collar whereas Perrona vindobonensis lacks beads or spines. In P. ilonae the spiral cords are separated by narrow, deeply incised grooves, whereas P. vindobonensis has wider and shallow interspaces. Both species might also have been ecologically separated. Perrona ilonae lived on coastal mudflats, whereas P. vindobonensis preferred sandy bottoms with seagrass.

Perrona coppii ( Bellardi, 1877) , from the Tortonian of Stazzano ( Italy), differs by its beaded basal and peribasal spiral cords (see Ferrero Mortara et al. 1981, pl. 14, figs 8a–b).

Paleoenvironment. Kókay (1966) described this species from coastal mudflats. Similar conditions are indicated at St. Florian ( Austria), where Perrona ilonae co-occurs with potamidid gastropods.

Distribution in Central Paratethys. Badenian (middle Miocene): Ukrainian Fore-Carpathian Basin: Faszczówka ( Ukraine)( Friedberg 1912); Pannonian Basin: Herend, Mátraverebély, Szentkút ( Hungary) (CsepreghyMeznerics 1954; Kókay 1966; Pálfy et al. 2008).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 4A. ‘Clavatula’ justinae (Hoernes & Auinger, 1891), NHMW ex1949/0005/0110b, Buituri (Romania). B. ‘Clavatula’ mariae (Hoernes & Auinger, 1891), lectotype, NHMW 1849/0013/0009, Mold (Austria). C. ‘Clavatula’ olgae (Hoernes & Auinger, 1891), lectotype, NHMW 1862/0001/0552, Buituri (Romania). D. ‘Clavatula’ orientoromana Báldi, 1960, holotype, M.60.145, Szokolya (Hungary). E. ‘Clavatula’ juliae (Hoernes & Auinger 1891), paralectotype, NHMW 1949/0005/0043c, Lăpugiu de Sus (Romania). F. ‘Clavatula’ apolloniae (Hoernes & Auinger, 1891), NHMW 1868/0001/0474a, Lăpugiu de Sus (Romania). G. Neoperrona harzhauseri (Kovács & Vicián, 2021), NHMW 2019/0151/019a, Letkés (Hungary). H. Neoperrona zoltanorum nov. sp., holotype, NHMW 2019/0151/0108, Letkés (Hungary). I. Scaevatula sidoniae (Hoernes & Auinger, 1891), NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). J. Pusionella palatina Strausz, 1954, NHMW 2021/0159/0003, Várpalota (Hungary). K. Pusionella hofmanni nov. sp., NHMW 1867/0019/0136, holotype, Coşteiu de Sus (Romania). L. Perrona barbarae (Hoernes & Auinger, 1891) NHMW 1949/0005/0103a, Rückersdorf (Austria). M. Perrona descendens (Hilber, 1879), NHMW 1846/0049/0017, St. Florian (Austria). N. Perrona emmae (Hoernes & Auinger 1891), NHMW 2021/0127/0005, Enzesfeld (Austria), NHMW 1970/1396/1449, Baden (Austria). O. Perrona ernae (Šuklje, 1929), NHMW 2021/0156/0001, Weinsteig (Austria). P. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). Q. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary). R. Perrona lydiae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0112, Gainfarn (Austria). S. Perrona oliviae (Hoernes & Auinger 1891), paralectotype, NHMW 1873/0026/0027, Lăpugiu de Sus (Romania). T. Perrona rosaliae (Hoernes & Auinger 1891), lectotype, NHMW 1999z0098/0002, Vienna/Pötzleinsdorf (Austria). U. Perrona sabinae (Hoernes & Auinger 1891), paralectotype, NHMW 2021/0154/0001, Lăpugiu de Sus (Romania). V. Perrona styriaca (Hilber, 1879), lectotype, NHMW 1846/0037/0309, Pöls (Austria). W. Perrona ursulae (Hoernes & Auinger, 1891), paralectotype, NHMW 1846/0001/1294, Buituri (Romania). X. Perrona vindobonensis (Quenstedt, 1884), paralectotype, NHMW 1997z0178/1467, Gainfarn (Austria), Y. Perrona wanzenboecki nov. nom., NHMW 2021/0158/0001, neotype, St. Veit an der Triesting (Austria). Z. Tomellana dulaii nov. sp., NHMW 2021/0148/0002, holotype, Nemeşeşti (Romania). Aa. Tomellana aueri nov. sp., NHMW 2013/0078/0436, Baden (Austria). Ab. Striopusionella szekelyhidiae (Kovács & Vicián, 2021), holotype, PAL 2021.19.1, Letkés (Hungary). Ac. Scaevatula sidoniae (Hoernes & Auinger, 1891). NHMW 1855/0043/0045, holotype, Lăpugiu de Sus (Romania). Scale bar is 5 mm for all specimens except for Perrona barbarae and protoconch of Scaevatula sidoniae.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 31 A1–A3. Perrona descendens (Hilber, 1879), lectotype, NHMW 1862/0033/0034, Pöls (Austria). B1–B3. Perrona descendens (Hilber, 1879), paralectotype, NHMW 1949/0005/0101, St. Florian (Austria). C1–C3. Perrona descendens (Hilber, 1879), paralectotype, NHMW 1861/0001/0260, Pöls (Austria). D1–D3. Perrona descendens (Hilber, 1879), paralectotype, NHMW 1861/0001/0260a, Pöls (Austria). E1–E3. Perrona floriana (Hilber, 1879), holotype, NHMW 1858/0020/0012, Guglitz at St. Florian (Austria). F1–F2. Perrona ilonae nov. nom., M.61.4367, Szentkút, Mátraverebély (Hungary).