Olegia szokolyensis ( Strausz, 1960 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 87-88

Olegia szokolyensis ( Strausz, 1960) nov. comb.

Figs 24D 1 –D 2 , 5

* Clavatula schreibersi szokolyensis nov. var. — Strausz 1960: 351, pl. 20, figs 2, 5.

Clavatula schreibersi szokolyensis Strausz, 1960 — Strausz 1966: 410, pl. 16, figs 9–10, text-fig. 182.

Clavatula (Clavatula) interrupta sophiae (R. Hoernes et M. Auinger, 1879) — Švagrovský 1981b: 156, pl. 48, fig. 1 [non ‘Clavatula’ sophiae (Hoernes & Auinger, 1879) ]

Type material. Holotype: M.9, Szokolya ( Hungary), SL: 14.5 mm, MD: 5.7 mm, illustrated in Strausz (1960, pl. 20, figs 2, 5), stored in the Geological and Geophysical Institute of Hungary (Budapest), figs 24D 1 –D 2.

Revised description. Shell small, moderately stout fusiform, with weakly gradate conical spire and high last whorl; apical angle 44°. Protoconch not preserved. Teleoconch of at least eight whorls. Early teleoconch whorls flat-sided (strongly abraded) with smooth subsutural spiral cord and strongly beaded suprasutural cord. Later teleoconch whorls with narrow, indistinctly beaded subsutural collar, weakly concave mid-portion bearing axial riblets, suprasutural cord bearing large, rounded tubercles. Suture narrowly incised, undulating, running below abapical row of tubercles. Last whorl ~ 65% of total height, subcylindrical; subsutural collar broad, swollen, bearing 12 large rounded tubercles. Subsutural ramp weakly concave, delimited by tubercular shoulder cord, weakly convex below, weakly constricted at base. Base delimited by weakly tubercular peribasal cord; poorly delimited and developed siphonal fasciole. Sculpture of two tubercular primary spiral cords. Adapical cord delimiting shoulder bearing large low tubercles, second cord at mid-whorl delimiting periphery with smaller tubercles, peribasal and perifasciolar cords not developed; suggestion of several weaker cords over base and fasciole. Aperture moderately wide, pyriform. Outer lip not preserved. Anal sinus moderately deep and wide, asymmetrically U-shaped, with apex mid-ramp. Siphonal canal moderately long. Columella weakly excavated, straight, oblique, smooth. Columellar and parietal callus weakly thickened, adherent, forming narrow pseudumbilical chink.

Discussion. Olegia szokolyensis was established by Strausz (1960) as subspecies of ‘ Clavatula ’ schreibersi ( Hörnes, 1854) , but the absence of spiral sculpture in O. szokolyensis excludes a close relationship between the two species. It is difficult to characterize this species as it is represented by a single, abraded specimen. It differs from all of its congeners in having only the spiral cords at the shoulder and the periphery well developed, bearing large, rounded tubercles, and a weakly developed and subobsoletely tubercular peribasal cord. The perifasciolar cord is not developed. The strong subsutural and peribasal cords make it most similar to O. doderleini ( Hörnes, 1854) , but in that species the mid-whorl portion of the last whorl is more inflated and, as mentioned above, it has a stronger peribasal and a perifasciolar cord developed, the former very weak and the later absent in O. szokolyensis . Apart from these shell differences O. szokolyensis derives from open marine Badenian deposits, whereas O. doderleini is an endemic species of the middle Sarmatian, known from hypersaline environments.

Paleoenvironment. The fauna from Szokolya ( Hungary) corresponds to that of the basinal clays of the Baden Formation in the Vienna Basin, which formed in middle to outer neritic environments (Krenner et al. 2021).

Distribution in Central Paratethys. Badenian (middle Miocene): Pannonian Basin: Szokolya ( Hungary) ( Strausz 1966), Vienna Basin: Devín ( Slovakia) ( Švagrovský 1981b).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 2. Descriptive terms used in descriptions and measurements to shell morphology, whorl profiles and anal sinus. SL: shell length, MD: maximum diameter, AA: apical angle, LWH: last whorl height, AH: aperture height.AL: aperture length.AW: aperture width.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 24 A1–A3. Olegia agathae (Hoernes & Auinger, 1891), lectotype, NHMW 1999z0098/0001, Niederkreuzstetten (Austria). B1–B3. Olegia agathae (Hoernes & Auinger, 1891), paralectotype, NHMW 1861/0050/0088a, Niederkreuzstetten (Austria). C. Olegia agathae (Hoernes & Auinger, 1891), paralectotype, NHMW 1861/0050/0088b, Niederkreuzstetten (Austria). E1–E2. Olegia curta (Boda 1959), holotype, Sz.1432, Várpalota (Hungary). D1–D2. Olegia szokolyensis (Strausz, 1960), holotype, M.9, Szokolya (Hungary). F1–F3. Olegia doderleini (Hörnes, 1854), lectotype, NHMW 1866/0001/0550, Wiesen (Austria). G1–G2. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria).