Olegia agathae ( Hoernes & Auinger, 1891 ) Harzhauser & Landau & Janssen, 2022

Harzhauser, Mathias, Landau, Bernard & Janssen, Ronald, 2022, The Clavatulidae (Gastropoda, Conoidea) of the Miocene Paratethys Sea with considerations on fossil and extant Clavatulidae genera, Zootaxa 5123 (1), pp. 1-172 : 78-79

Olegia agathae ( Hoernes & Auinger, 1891) nov. comb.

Figs 24A 1 –A 3 , B 1 –B 3 , C, 5 , 6

Clavatula Agathae —Hoernes 1891: 131 [nomen nudum].

* Pleurotoma (Clavatula) Agathae nov. form.— Hoernes & Auinger 1891: 352, pl. 46, figs 10a–b.

C [lavatula]. (C [lavatula]) agathae (R. Hörn. et Au.) — Sieber 1958a: 157.

Type material. Lectotype designated herein: NHMW 1999 z0098/0001, Niederkreuzstetten ( Austria), SL: 21.5 mm, MD: 9.0 mm, illustrated in Hoernes & Auinger (1891, pl. 46, fig. 10), figs 24A 1 –A 3 . Paralectotypes: NHMW 1861 /0050/0088a, SL: 16.9 mm , MD: 7.1mm, Niederkreuzstetten ( Austria) ,figs 24B 1 –B 3; NHMW 1861 /0050/0088b, SL: 18.9 mm , MD: 8.2 mm, Niederkreuzstetten ( Austria) , figs 24C; NHMW 1861 /0050/0088, 14 specimens from Niederkreuzstetten ( Austria) .

Revised description. Shell medium-sized, solid, moderately slender fusiform; apical angle ~35°. Protoconch not preserved. Early teleoconch whorls with bipartite sculpture; weak subsutural cord, concave mid-whorl portion with comma-shaped axial riblets, stronger beaded suprasutural cord. Sculpture changing around forth teleoconch whorl; two prominent spiral cords bearing pointed nodes at sutures, mid-whorl deeply concave, with marked central concavity crossed by about five narrow spiral cords. Suture narrowly but deeply incised, weakly undulating. Last whorl ~ 65% of total height; subsutural collar bearing sharp tubercles, subsutural ramp concave, delimited by tubercular shoulder cord, weakly convex below, weakly constricted at base, siphonal fasciole poorly delimited, weakly rounded; sculpture of low, rounded ribs developed below shoulder, crossed by irregular flattened cords, separated by narrow grooves, forming weakly beaded sculpture, peribasal and perifasciolar cords somewhat strengthened. Aperture elongate, relatively narrow; outer lip not thickened, smooth within; anal sinus moderately wide, moderately deep, asymmetrically U-shaped, with apex just below collar; siphonal canal moderately short, broad, shallowly notched at tip. Columella weakly excavated in upper third, straight below, smooth. Columellar and parietal callus thickened, sharply delimited, forming broad callus rim.

Discussion. This species shows some degree of variability in the sculpture below the shoulder on the last whorl. In some specimens the axial ribs are well-developed making the sculpture clearly tubercular (figs 24B 1 –B 3), whereas in others these ribs are poorly developed, resulting in almost flat cords (figs 24A 1 –A 3).

Olegia mandici nov. sp. is comparable in shape and sculpture, but differs in its larger size and its secondary and tertiary spiral cords on the base. Olegia doderleini ( Hörnes, 1854) differs in its gradate spire and Olegia rumana ( Simionescu & Barbu, 1940) has weaker tubercles, is more slender and lacks secondary spiral cords.

It is superficially reminiscent of ‘ Clavatula ’ apolloniae ( Hoernes &Auinger, 1891) and ‘ C. ’ veronicae ( Hoernes & Auinger, 1891) , both form the middle Miocene of the Paratethys Sea, but differs clearly in its more slender shell and the absence of spiral sculpture on the spire whorls. ‘ Clavatula ’ manzonii Bellardi, 1877, from the Tortonian of Stazzano ( Italy), is comparable in shape and develops two spiral rows of nodes along the adapical and abapical suture, but differs in the presence of a central cord of delicate beads (see Bellardi 1877: 163, pl. 5, fig. 24; Ferrero Mortara et al. 1981, pl. 14, figs. 1a–b).

Paleoenvironment. The locality Niederkreuzstetten represents inner neritic environments with sand bottom ( Harzhauser et al. 2002).

Distribution in Central Paratethys. Karpatian (early Miocene): North Alpine-Carpathian Foreland Basin: Niederkreuzstetten ( Austria) ( Hoernes & Auinger 1891).

Figures

FIGURE 1. A. Map of central and south–eastern Europe, representing the area that was partly covered by the Central Paratethys Sea. The white insert corresponds to the area that is restored in the palaeogeographic map below. B. Paleogeography of the Middle Miocene Paratethys Sea showing the most important sedimentary basins (NAFB: North Alpine Foreland Basin, E.-Sopr B: Eisenstadt–Sopron Basin).

FIGURE 3. Early teleoconch whorls of extant and fossil Clavatulidae species. A. Clavatula regia (Röding, 1798), NHMW 1683 6/28, West Africa. B. Clavatula rubrifasciata (Reeve, 1845), NHMW 10203, Gorée (Senegal). C. ‘Clavatula’ diadema (Kiener, 1839), NHMW 1884/0010/0023, Gorée (Senegal). D. ‘Clavatula’ mystica (Reeve, 1843), MNHN-IM-2013-59897, Est de l'ile de Gorée (Senegal). E. ‘Clavatula’ bimarginata (Lamarck, 1822), MNHN-IM-2013-43369, Dakar (Senegal). F. ‘Clavatula’ pyramidata (Kiener, 1839), MNHN-IM-2009-24982, Baie de Pointe-Noire (Republique du Congo). G. Clavatula sorini nov. sp., paratype, NHMW 1874/0025/0046, Lăpugiu de Sus (Romania). H. ‘Clavatula’ romana (Defrance, 1826), NHMW 1999/0098/0004, Modra-Kráľová (Slovakia). I. ‘Clavatula’ interrupta (Brocchi, 1814), NHMW 1861/0033/0420, Buituri (Romania). J. ‘Clavatula’ hirmetzli Kovács & Vicián, 2021, NHMW 1868/0001/0069, Baden-Sooss (Austria). K. ‘Clavatula’ irisae nov. sp., NHMW 2021/0126/0001, holotype, Gainfarn (Austria). L. ‘Clavatula’ sophiae (Hoernes & Auinger, 1891), lectotype, NHMW 2021/0127/0001, Enzesfeld (Austria). M. ‘Clavatula’ veronicae (Hoernes &Auinger, 1891), NHMW 1855/0043/0044b, Lăpugiu de Sus (Romania). N. ‘Clavatula’ santhai Kovács & Vicián, 2021, holotype, HNHM, PAL 2021.20.1, Letkés (Hungary).O. ‘Clavatula’ camillae (Hoernes &Auinger, 1891), lectotype, NHMW 1949/0005/0042, Lăpugiu de Sus (Romania). P. Megaclavatula amaliae (Hoernes & Auinger, 1891), lectotype, NHMW 1949/0005/0049, Lăpugiu de Sus (Romania). Q. Megaclavatula grunerti nov. sp., paratype, NHMW 2021/0127/0003, Enzesfeld (Austria). R. Megaclavatula laevigata (Eichwald, 1830), NHMW 1853/0038/0035a, Korytnica (Poland). S. Megaclavatula pilleri nov. sp., holotype, NHMW 1949/0005/0051a, Baden-Sooss (Austria). T. Megaclavatula polonica (Pusch, 1837), NHMW 1862/0001/0324, Bad Vöslau (Austria).U. Granulatocincta angelae (Hoernes &Auinger, 1891), paralectotype, NHMW 1949/0005/0099, Möllersdorf (Austria). V. Granulatocincta granulatocincta (Münster in Goldfuss, 1841), NHMW 1846/0028/0009, Enzesfeld (Austria). W. Granulatocincta nataliae (Hoernes & Auinger, 1891), NHMW 1855/0035/0021, holotype, Belgrade/Rakovica (Serbia). X. Granulatocincta schreibersi (Hörnes, 1854), paralectotype, NHMW1997z0178/1133, Bad Vöslau (Austria).Y. Granulatocincta callim nov. sp., paratype, NHMW 2021/0128/0001, Bad Sauerbrunn (Austria). Z. Granulatocincta contorta (Švagrovský, 1958), holotype, SNMB Z 3076, Kuzmice (Slovakia), Aa. Granulatocincta theoderichi nov. sp., paratype, NHMW 2016/0026/0320b, Ritzing (Austria). Ab. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria). Ac. Olegia mandici nov. sp., paratype, NHMW 1872/0031/0008a, Bekića vrelo, Čremušnica at Donji Kirin (Croatia). Ae. Olegia dorotheae (Hoernes & Auinger, 1891), paralectotype, NHMW 1949/0005/0040, Kleinebersdorf (Austria). Scale bar is 5 mm for all specimens except for Granulatocincta contorta, Olegia doderleini and O. mandici.

FIGURE 5. Anal sinus morphology of Paratethyan Clavatulidae.

FIGURE 6. Apertures of Paratethyan Clavatulidae (C. = Clavatula, ‘C.’ = ‘Clavatula’, G. = Granulatocincta, M. = Megaclavaltula, O. = Olegia).

FIGURE 24 A1–A3. Olegia agathae (Hoernes & Auinger, 1891), lectotype, NHMW 1999z0098/0001, Niederkreuzstetten (Austria). B1–B3. Olegia agathae (Hoernes & Auinger, 1891), paralectotype, NHMW 1861/0050/0088a, Niederkreuzstetten (Austria). C. Olegia agathae (Hoernes & Auinger, 1891), paralectotype, NHMW 1861/0050/0088b, Niederkreuzstetten (Austria). E1–E2. Olegia curta (Boda 1959), holotype, Sz.1432, Várpalota (Hungary). D1–D2. Olegia szokolyensis (Strausz, 1960), holotype, M.9, Szokolya (Hungary). F1–F3. Olegia doderleini (Hörnes, 1854), lectotype, NHMW 1866/0001/0550, Wiesen (Austria). G1–G2. Olegia doderleini (Hörnes, 1854), paralectotype, NHMW 1858/0015/0147, Wiesen (Austria).