Glaresis, Erichson, 1848

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 31-32

Key to species of Glaresis - Mendica Group

1. Mesotibial apex strongly projecting laterally, projection apically acute ( Fig. 34C ); metatibia abruptly widened before apex; Mexico, Central America; subgroup C ................................... 12

— Mesotibial apex weakly projecting laterally, projection rounded ( Fig. 29D ) (except G. confusa , G. falli , and G. texana with apex strongly projecting, apically acute); metatibia gradually widened from base to apex ( Fig. 19G ); Mexico, Canada, United States; subgroups A, B ..................... 2

2(1). Metasternal groove shallow, at least inner border of groove tapered ( Fig. 19D ); subgroup A .. 3

— Metasternal groove deep, distinct, sides abrupt, ridged ( Fig. 27D ); subgroup B ....................... 7

3(2). Outer margin of metatibia with series of small, regular teeth from base to apex, without large posteromedian lateral projection ( Fig. 21F ); Texas ................................... G. texana Gordon

— Outer margin of metatibia with distinct posteromedian lateral projecton ( Fig. 20F ); not known from Texas .................................................................................................................................. 4

4(3). Clypeal apex with large, widely spaced tubercles medially ( Fig. 23B ); metatibial posteromedian lateral projection large, apically bifid ( Fig. 23F ) ............................................... G. falli , n. sp.

— Clypeal apex with small, evenly spaced tubercles throughout ( Fig. 22B ); metatibial posteromedian lateral projection small, tooth—like ( Fig. 22F ) ........................................................................ 5

5(4). Tubercles on head surface small, sparse ( Fig. 22B ); apex of mesotibia strongly projecting, apically acute; Alberta, Colorado, Utah ................................................................... G. confusa Brown

— Tubercles on head large, dense ( Fig. 19B ); apex of mesotibia not strongly projecting, apically rounded ....................................................................................................................................... 6

6(5). Posterosuperior metatrochanteral surface with 2 teeth ( Fig. 19C ); median lobe of male genitalia wide, apex broadly rounded ( Fig. 19E ) ..................................................... G. california , n. sp.

— Posteriosuperior metatrochanteral surface with single tooth ( Fig. 20C ); median lobe of male genitalia slender, apically acute ( Fig. 20E ) ............................................. G. howdeni Gordon

7(2). Posterosuperior metatrochanteral surface with 2 teeth ( Fig. 24G ) ............................................ 8

— Posterosuperior metatrochanteral surface with single tooth ( Fig. 27G ) .................................. 10

8(7). Metasternal sculpture with carinae straight, not irregular, pronotal carinae distinctly spaced, not dense; male genitalia with basal lobe slender, narrowed in basal 1/3, apex rounded ( Fig. 26F ); southern California ............................................................................ G. bautista , n. sp.

— Metasternal sculpture with carinae irregular, dense; pronotal carinae closely spaced; male genitalia not as described for G. bautista ; not known from southern California ................................... 9

9(8). Metatasternal sculpture irregular, forming reticulate pattern extended from apex to metacoxal cavity of metasternum ( Fig. 24D ); apical trochanteral margin with small teeth, teeth often not visible; male genitalia with basal lobe broad, apically rounded ( Fig. 24E ); Arizona, California, Mexico............................................................................................................ G. mendica Horn

— Metasternal sculpture irregular, not forming reticulate pattern, not extended to metacoxal cavity ( Fig. 25D ) ; apical trochanteral margin with several distinct teeth; male genitalia with basal lobe narrow, apically acute ( Fig. 25E ); California, Idaho, Nevada, Wyoming ........................... ............................................................................................................. G. paramendica , n. sp.

10(7). Posteromedian lateral projection of metatibia large, pronounced ( Fig. 29F ); surface sculpture of frons and vertex composed of dense raised, irregular ridges coalesced into a reticulate pattern ( Fig. 29B ); Baja California Sur ................................................................. G. australis . n. sp.

— Posteromedian lateral projection of metatibia small, not pronounced ( Fig.20F ); surface sculpture of frons and vertex reduced, if ridges present then not forming a reticulate pattern; United States......................................................................................................................................... 11

11(10). Metasternum tumid, protuberant, with heavy, somewhat reticulate surface sculpture laterally ( Fig. 30D ); southeast Texas ........................................................................... G. tumida , n. sp.

— Metasternum flat or medially concave, surface sculpture comparatively light, not forming reticulate pattern; not known from southeast Texas .............................................................................. 12

12(11). Posterosuperior metafemoral margin with single small tooth ( Fig. 28H ) ; posterior metatrochanteral margin serrate with several small teeth; median lobe of male genitalia wide, not “pinched” medially ( Fig. 28F ); New Mexico, west Texas .............. G. caenulenta , n. sp.

— Posterosuperior metafemoral margin without tooth, or occasionally with single, tiny, indistinct tooth ; posterior metatrochanteral margin with 3 small teeth ( Fig. 27G ); median lobe of male genitalia slender, “pinched” medially ( Fig. 17E ); Arizona ....................... G. gordoni Warner

13(1). Tubercles in median emargination of clypeal apex large, pronounced, tooth—like ( Fig. 34B ); metasternal groove widened from base to apex, apex extremely wide, nearly reaching metipimeron ( Fig. 34D ); Costa Rica.......................................................................... G. costaricensis , n. sp.

— Tubercles in median emargination of clypeal apex small, tubercles irregular but more or less equal in size across apical margin ( Fig. 33B ); metasternal groove not extremely widened, apex not approaching metepimeron ................................................................................................. 14

14(13). Metatibia with posterolateral projection ( Fig. 33H ); southern Mexico.................................... 15

— Metatibia without posterolateral projection, lateral flange usually serrate with small teeth ( Fig. 35F ); not known from southern Mexico.................................................................................. 16

15(14). Metasternal surface lacking pronounced carinae arranged in reticulate pattern; surface of metafemur finely rugose, appearing smooth ( Fig. 33G ); Zacateca, Mexico................................ ...................................................................................................................... G. zacateca , n. sp.

— Metasternal surface with pronounced carinae arranged in reticulate pattern ( Fig. 32D ); surface of metafemur coarsely microreticulate; Puebla, Mexico.............................. G. donaldi , n. sp.

16(14). Posterior margin of metatrochanter with series of large tubercles, second tubercle to trochanteral margin pronounced, tooth—like ( Fig. 31F ); surface of clypeus, apical portion of frons with large, dense tubercles ( Fig. 31B ); Guatemala.............................................. G. dentata , n. sp.

— Posterior margin of metatrochanter smooth, without tubercles ( Fig. 35F ); surface of clypeus and anteromedian portion of frons densely, finely rugose, with small, sparse tubercles ( Fig. 35B ); Mexico, Coahuila ........................................................................................... G. limbata , n. sp.

Figures

Figures 17A-17G. Glaresis inducta Horn. A) Habitus view. B) Clypeus and head surface. C) pronotal surface. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg and trochanter, ventral view.

Figures 19A-19G. Glaresis california, new species. A) Habitus view. B) Clypeus and head surface. C) Trochanter with 2 teeth. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.

Figures 20A-20G. Glaresis howdeni Gordon. A) Habitus view. B) Clypeus and head surface. C) Trochanter with single tooth. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.

Figures 21A-21G. Glaresis texana Gordon. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytra. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.

Figures 22A-22G. Glaresis confusa Brown. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. 22D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.

Figures 23A-23G. Glaresis falli, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.

Figures 24A-24G. Glaresis mendica Horn. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg. ventral view.

Figures 25A-25G. Glaresis paramendica, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Metasternum with metasternal groove. E) Male genitalia. F) Metaleg and trochanter, dorsal view. G) Metaleg and trochanter, ventral view.

Figures 26A-26G. Glaresis bautista, new species. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum with metasternal groove and mesotibia. E) Male genitalia. F) Metaleg and trochanter, dorsal view. G) Metaleg and trochanter, ventral view.

Figures 27A-27G. Glaresis gordoni Warner. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum with metasternal groove and mesotibia. E) Male genitalia. F) Metaleg and trochanter, dorsal view. G) Metaleg and trochanter, ventral view.

Figures 28A-28H. Glaresis caenulenta, new species. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytra. D) Metasternum with metasternal groove and mesotibia. E) Elytral striae. F) Male genitalia. G) Metaleg, dorsal view. H) Metaleg, ventral view.

Figures 29A-29G. Glaresis australis, new species. A) Habitus view. B) Clypeus and head surface. C) Pronotal surface. D) Metasternum with metasternal groove and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.

Figures 30A-30G. Glaresis tumida, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Metasternum and mesotibia. E) Male genitalia. F) Metaleg, dorsal view. G) Metaleg, ventral view.

Figures 31A-31G. Glaresis dentata, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Metasternum and mesotibia. E) Metafemur and trochanter, dorsal view. F) Metafemur and trochanter, ventral view. G) Metatibia, ventral view.

Figures 32A-32F. Glaresis donaldi, new species.A) Habitus view. B) Clypeus and head surface. C) Pronotal surface and elytral striae. D) Metasternum and mesotibia. E) Elytral striae. F) Metaleg and trochanter with teeth, dorsal view.

Figures 33A-33H. Glaresis zacateca, new species. A) Habitus view. B) Clypeus and head surface. C) Elytral striae. D) Pronotal surface and elytra. E) Metasternum. F) Male genitalia. G) Metalegs and trochanter. H) Metatibia, dorsal view.

Figures 34A-34F. Glaresis costaricensis, new species. A) Habitus view. B) Clypeus and head surface. C) Mesotibia. D) Metasternum. E) Metafemur and trochanter, dorsal view. F) Metafemur and trochanter, ventral view.

Figures 35A-35F. Glaresis limbata, new species. A) Habitus view. B) Clypeus and head surface. C) Metatrochanter with teeth indicated. D) Metasternum. E) Pronotal surface and elytra. F) Metaleg, dorsal view.