Glaresis pardoalcaidei Martinez, Pereira, and Vulcano

Gordon, Robert D. & Hanley, Guy A., 2014, Systematic revision of American Glaresidae (Coleoptera: Scarabaeoidea), Insecta Mundi 2014 (333), pp. 1-91 : 7-8

Glaresis pardoalcaidei Martinez, Pereira, and Vulcano

Glaresis pardoalcaidei Martinez, Pereira, and Vulcano 1961:69 .

Description. Male. Length 5.3 mm, width 3.0 mm; body form blocky, robust, slightly widened from elytral base to apical 1/3 ( Fig. 1A). Color reddish brown. Head with clypeal surface and frons densely rugose, microtuberculate, with scattered large, round tubercles, setae long, nearly erect ( Fig. 1B); vertex with short, pronounced basomedian carina, surface rugose, with small round tubercles. Clypeal apex weakly emarginate, medially thickened, slightly angled posteriorly, with small, evenly spaced, apically acute tubercles, appearing dentate, lateral angles oblique, acute ( Fig.1B). Mandible pair symmetrical; mesal tooth strong; lateral prominence obsolete, outer margin evenly rounded. Pronotum without fovea except for small, anteromedian and posteromedian foveae evident; surface densely rugose, microtubeculate, with small, uniform, round, setae-bearing tubercles, setae slightly decumbent, much longer than tubercle ( Fig.1 E); lateral and posterior margins bordered, crenulate, anterior border smooth, not bordered; posterolateral angle produced, flanged. Elytra with surface dull, densely rugose; all striae wide, without carinae, with widely, evenly spaced, small tubercles bearing long, slightly decumbent setae; intervals with small, deep, round punctures ( Fig. 1D). Metasternum long, dull, densely microreticulate, median surface not microtuberculate, without larger tubercles, laterally both microtuberculate and with small, round tubercles bearing long, nearly erect setae, median area flat, without median carina ( Fig. 1C), metasternal groove strong, deep, both sides abruptly ridged. Lateral protibial teeth short, broadly rounded, unevenly spaced, basal teeth close together ( Fig. 1E). Mesotibia with 5 short, closely spaced spines on outer margin, slightly projecting at apex ( Fig. 1C). Posterior metatrochanteral margin serrate with several small teeth, lateral angle produced, strongly curved inward, apically acute ( Fig. 1G); posterosuperior surface of metatrochanter without teeth. Metafemoral surface with widely scattered, small, round, setae-bearing tubercles, tubercles arranged in horizontal row posteromedially, microreticulate; width to length ratio 1.0:1.6, without flange on anterior margin; posterosuperior margin without teeth. Metatibia narrow, abruptly widened before apex; surface smooth, shiny, without posteromedian lateral projection, lateral margin with row of closely spaced, tooth-like tubercles bearing short, stout setae, medially with very thin tubercles forming slightly irregular row from near base to apex ( Fig. 1G, H). Apical margin of 5th abdominal ventrite broadly rounded. Genitalia long, basal piece slightly longer than parameres, proximal end curved; median lobe shorter than parameres, wider at middle than a paramere, curved upward in apical 1/3, apex bifid; parameres weakly curved in lateral view, strongly narrowed in apical 1/ 3, slightly curved on inner margin, apex bluntly rounded ( Fig. 1F).

Female. Metasternum without large, median tubercles; metatrochanter slightly angled outward apically, without curved tooth; apex of 5th abdominal ventrite same as in male.

Variation. Length 4.3 to 6.3 mm, width 2.4 to 3.4 mm. Clypeal apex with thickened ridge narrow or wide, clypeal surface extremely rugose to moderately so; median area of metasternum varies from convex, flat, or with depression, metasternal surface varies from densely, smoothly rugose medially to densely, roughly rugose, angled outward apically, without curved tooth.

Type locality. South America, Argentina, “ciudad de San Miguel del Tucumán, Puente del río. Sah.”

Type depository. Unknown.

Temporal distribution. November-January.

Geographical distribution. Martinez et al. (1961) had a large number of specimens at hand when he described this species. He listed the holotype and 118 paratypes from localities in Argentina. Bolivia, Brasil, and Paraguay. Specimens available to us fall within the range of the Martinez paratypes with the

following exceptions. ARGENTINA. Cordoba: “Obispo Tre?” La Pampa: Santa Rosa ; Santiago del Estero: Santiago del Estero. BOLIVIA: Santa Cruz: Buena Vista, F & F Hotel. BRASIL: Sao Paulo: Piracicaba. ( FSCA) ( OSUC) ( USNM) .

Remarks. This is the largest known species of Glaresidae further distinguished by the blocky, robust body form; flanged posterolateral pronotal angle; sexual dimorphism; and narrow metatibia with smooth, shiny surface and lack of armature on lateral margin.