Ancyronyx

Freitag, Hendrik & Jäch, Manfred A., 2007, The genus Ancyronyx Erichson, 1847 (Coleoptera, Elmidae) in Palawan and Busuanga, (Philippines) with descriptions of six new species, Zootaxa 1590, pp. 37-59 : 40-41

Key to the Philippine species of Ancyronyx

1. CL> 1.9 mm; legs very long (length of elytra: metafemur> 0.5); ovipositor (coxite) short and stout (comp. Fig. 16 d) ......................................................................................................................................... 2

- CL <1.8 mm; legs long to moderately long (length of elytra: metafemur <0.5); ovipositor (coxite) long and slender ( Figs. 11 d; 13d; 14d) ............................................................................................................... 3

2. Pronotum predominantly dark, anteriorly yellowish; elytra with yellowish “X”-shaped, colour pattern. Aedeagal parameres moderately long (ca. 0.5 of entire aedeagus), clearly not reaching opening of ventral sac, with numerous long apical setae. Ovipositor medially without, but apically with few trichoid spines. .......................................................................................................................... A. schillhammeri Jäch, 1994

- Pronotum predominantly yellowish, anteriorly brown; elytra with large yellowish median marking, sometimes extended into bands at the edges ( Fig. 10 ). Aedeagal parameres very long (ca. 0.65 of entire aedeagus), surpassing opening of ventral sac, almost reaching apex of penis, without conspicuous setae ( Fig. 16 a, b). Ovipositor without apical spines, but with one long median spine (at apical margin of valvifer) ( Fig. 16 d) ...................................................................................... A. helgeschneideri Freitag & Jäch sp. n.

3. Elytra with four pairs of elongately oval, yellow patches; total length ca. 1.3 mm. Aedeagus conically tapered; parameres moderately long (ca. 0.6 of entire aedeagus), with few very short apical setae ............ ............................................................................................................................ A. sophiemarie Jäch, 2004

- Elytra with two pairs of yellowish patches or entirely dark ( Figs. 3–9 ). Aedeagus different ..................... 4

4. Elytra with two pairs of large longitudinal elongate yellowish patches ( Fig. 9 ); CL ca. 1.1 mm. Aedeagal parameres moderately long (ca. 0.5 of entire aedeagus), with two very long apical setae (> 0.5 paramere length) ( Fig. 15 a, b) ................................................................................ A. minutulus Freitag & Jäch sp. n.

- Elytra with one subapical pair of longitudinal elongate yellowish patch and one pair of round or transversally elongate, yellowish patches or entirely dark ( Figs. 3–8 ); CL usually> 1.1 mm. Aedeagus different; parameres short (<0.5 of entire aedeagus length), with> 10 long setae or without long setae ( Figs. 11 b, 12b, 13b, 14b) ............................................................................................................................................. 5

5. Pronotum anteriorly with transverse white to yellowish patch or band ( Figs. 7, 8 ); CL <1.35 mm. Aedeagus conically tapered; parameres with> 10 long apical setae ( Fig. 14 a, b) A. minerva Freitag & Jäch sp. n.

- Pronotum entirely dark, without transverse patch or band ( Figs. 3–6 ); CL> 1.35 mm. Aedeagus subapi-

cally more or less widened, not conically tapered; parameres with <10 short to moderately long apical setae ( Figs. 11 a, 12a, 13a)........................................................................................................................... 6 6. [Determination in dark specimens unambiguously possible by sexual characters only]: Subbasal elytral patches more or less equally round, not reaching posterior or lateral elytral margin; legs entirely dark ( Fig. 6 ). Aedeagal median lobe long and moderately slender, subapically slightly widened (ca. 0.3 times as wide as long); ventral sac almost reaching apex ( Fig. 13 a, b) ..................... A. punkti Freitag & Jäch sp. n.

- Subbasal elytral patches transversely elongate (cross-stripe-like), at least reaching elytral margin at shoul- der; legs partly yellowish to pale brown ( Figs. 3, 5 ). Aedeagal median lobe subapically distinctly widened; ventral sac not reaching apical 0.1 of aedeagus ( Figs. 11 a, b; 12a, b) .............................................. 7

7. Femora predominantly yellowish to pale brown (except for articulations), ventral (inner) side without conspicuous setae; basal elytral patches <0.65 times as long as broad ( Fig. 3 ), in dark specimens medially interrupted, reduced or lacking ( Fig. 4 ). Aedeagal median lobe long and slender (ca. 0.2 times as wide as long; Fig. 11 a) ............................................................................................ A. patrolus Freitag & Jäch sp. n.

- Dorsal side of femora entirely dark brown; ventral side with conspicuous setae; basal elytral patches large, ca. 0.8 times as long as broad ( Fig. 5 ). Aedeagal median lobe short and stout (ca. 0.4 times as wide as long; Fig. 12 a) ................................................................................ A. pseudopatrolus Freitag & Jäch sp. n.

Figures

FIGURE 11. Ancyronyx patrolus sp. n., paratypes from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (right half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) female sternite VIII (strioles omitted); h) male sternite VIII (strioles omitted); i) female tergite VIII (setae and strioles of left half omitted).

FIGURE 12. Ancyronyx pseudopatrolus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male ventrite 5 (left half).

FIGURE 13. Ancyronyx punkti sp. n., paratypes from site “ Zettel (59) ”: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (left half); e) female tergite VIII (setae and strioles of left half omitted); f) female ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female sternite VIII (strioles omitted); i) male tergite VIII (setae and structures of left half omitted); j) male ventrite 5 (setae and strioles of left half omitted).

FIGURE 14. Ancyronyx minerva sp. n., ♂ holotype, Ψ paratype from same locality: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) ovipositor in ventral view (left half); e) male ventrite 5 (setae and strioles of left half omitted); f) female ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) male tergite VIII (setae and structures of left half omitted); i) female tergite VIII (setae and strioles of left half omitted); j) female sternite VIII (strioles omitted).

FIGURE 15. Ancyronyx minutulus sp. n., holotype: a, b) aedeagus in ventral and lateral view; c) male sternite IX (strioles omitted); d) male sternite VIII (strioles omitted); e) male tergite VIII (setae and structures of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted).

FIGURE 16. Ancyronyx helgeschneideri sp. n., ♂ paratype from CR 3, Ψ paratype from PR 1: a, b) aedeagus in ventral and lateral view; c) male sternite IX (one paraproct omitted); d) ovipositor in ventral view (left half); e) male tergite VIII (setae and strioles of left half omitted); f) male ventrite 5 (setae and strioles of left half omitted); g) male sternite VIII (strioles omitted); h) female tergite VIII (setae and strioles of left half omitted).

FIGURES 7 – 10. Habitus of 7) Ancyronyx minerva sp. n., paratype Ψ; 8) A. minerva sp. n., paratype Ψ, winged, partly completed; 9) A. minutulus sp. n., holotype ♂, partly completed; 10) A. helgeschneideri sp. n., paratype Ψ.

FIGURES 3 – 6. Habitus of 3) Ancyronyx patrolus sp. n., paratype ♂, usual colouration; 4) A. patrolus sp. n., paratype Ψ, dark coloured individual, partly completed; 5) A. pseudopatrolus sp. n., holotype ♂, partly completed; 6) A. punkti sp. n., paratype ♂, partly completed.