Theopea sauteri Chûjô, 1935

Lee, Chi-Feng & Bezdĕk, Jan, 2018, Revision of the genus Theopea Baly (Coleoptera: Chrysomelidae: Galerucinae) of East Asia: species lacking modified clypeus in males and the T. sauteri species group, Zootaxa 4508 (3), pp. 334-376 : 366-372

Theopea sauteri Chûjô

( Figs 4 D– 4I, 19)

Theopea sauteri Chûjô, 1935a: 169 ; Chûjô, 1938: 152 (Kosempo= Chiasien, 甲仙; Koshun= Henchun, 恆春); Chûjô, 1962: 158 (redescription); Chûjô, 1965: 98 (Rimogan= Fushan, 福山); Kimoto, 1965: 488 (Teraso= Manchoutsun, 滿州村); Kimoto, 1966: 33 (Hoozan= Fengshan, 鳳山); Kimoto, 1969: 58 (numerous records in Taiwan); Wilcox, 1973: 631 (catalogue); Kimoto, 1986: 58 (faunistic records); Kimoto, 1987: 190 (faunistic records); Kimoto, 1989b: 259 (faunistic records); Kimoto, 1991: 16 (faunistic records); Kimoto & Chu, 1996: 87 (catalogue); Kimoto & Takizawa, 1997: 202 (redescription); Beenen, 2010: 489 (catalogue); Lee & Cheng, 2010: 125 (redescription); Takahashi, 2012: 327 (faunistic records).

Theopea sauteri var. minuta Chûjô, 1935a: 171 .

Types. Theopea sauteri . Holotype ♂ ( TARI): “Formosa / Y. Miwa [p, w] // 西村 (= Nishimura, in Taoyuan County) / 24.7.1929 [h, written on the back of the previous label] // Holotype [h, red letters, pink label] // Theopea / sauteri / CHÛJÔ [h] / DET. M. CHUJO [p, w] // 2334 [p, w]” . Paratypes: 1♀ ( TARI), labeled: “ Rimogan (= Fushan, 福山, in Taipei County) / FORMOSA / 12-V-1933 / Col. M. CHUJO [p, w] // Paratype [h, red letters, pink label] // Theopea / sauteri / CHÛJÔ [h] / DET. M. CHUJO [p, w] // 2946 [p, w]” ; 1♀ ( SDEI), labeled: “ Taihorin (= Talin, 大 林, in Chiayi County) / Formosa / H. Sauter, 1911 [p, w] // 7. VII. [p, w] // Paratype [h, red letters, pink label] // Theopea / sauteri / CHÛJÔ [h] / DET. M. CHUJO [p, w]” .

Theopea sauteri var. minuta . Holotype ♂ (TARI): “ Formosa. / Musha (= Wushe, 霧社, in Nantou County), 1919 / V 18–VI 15. / T. Okuni, [p, w] // Holotype [h, red letters, pink label] // Theopea sauteri / CHÛJÔ var. / minuta CHÛJÔ [h] / DET. M. CHUJO [p, w] // 2893 [p, w]”. Paratypes: 1♂, 2♀♀ (TARI), labeled: “ Formosa. / Musha (= Wushe, 霧社, in Nantou County), 1919 / V 18–VI 15. / T. Okuni, [p, w] // Paratype [h, red letters, pink label] // Theopea sauteri / CHÛJÔ var. / minuta CHÛJÔ [h] / DET. M. CHUJO [p, w] // 2949, 2950, or 2951 [p, w]”; 1♂ (SDEI): “ Formosa. / Musha (= Wushe, Ăṻ, in Nantou County), 1919 / V 18–VI 15. / T. Okuni, [p, w] // Paratype [h, red letters, pink label] // Theopea sauteri / CHÛJÔ var. / minuta CHÛJÔ [h] / DET. M. CHUJO [p, w]”. Those diagnostic characters mentioned by Chûjô (1935a) belong to infraspecific variation of Theopea sauteri .

Other specimens examined (n= 941). TAIWAN. Chiayi: 3♂♂ ( TARI), Alishan (阿里山), 2200m, 21.VI.2014, leg. B.- X. Guo ; 1♀ ( TARI), same locality, 29.V.2016, leg. B.- X. Guo ; 1♂, 2♀♀ (1♂, 1♀ in NMNS; 1♀ in HNHM), Chikutoge (= Juchi , 竹崎), 120m, V.1909, leg. Sauter ; 1♀ ( TARI), Mihu trail (迷糊步道), 1100m, 28.IV.2013, leg. W.- C. Liao ; 2♀♀ ( TARI), Tutzuhu trail (杜仔湖步道), 1500m, 1.VI.2014, leg. W.- C. Laio ; Hsinchu: 1♀ ( NMNS), Kuansi (關西), 50m, 23.V.1992, leg. C.Y. Li ; 1♂, 1♀ ( TARI), Lupi (魯壁), 1450m, 19.VII.2008, leg. M.- H. Tsou ; 1♀ ( TARI), Wufeng (五峰), 400m, 14–16.VII.1982, leg. K.C. Chou & C. C. Pan ; Hualien: 1♀ ( EUMJ), Fuyuan (富源), 250m, 9–11.V.1972, leg. M. Sakai ; 1♂ ( NMNS), same locality, 28.V.1997, leg. C. W. & L. B. O’Brien ; 1♀ ( EUMJ), Kibison (奇美村), 50m, 18.IV.1978, leg. K. Murakami ; 2♂♂ ( TARI), Pilu (碧綠), 2150m, 31.V.2011, leg. M.- H. Tsou ; 1♂ ( JBCB) , 1♀ ( NMPC), Tairoko (太魯閣), 950m, 3.VI.2008, leg. F. & L. Kantner ; 1♂ ( TARI), same locality, 3.VI.2012, leg. J.- F. Hsieh ; 1♂ ( TARI), Tayuling (大禹嶺), 2560m, 9– 16.VI.1980, leg. K.S. Lin & B. H. Chen ; 1♂, 1♀ ( NMPC), Yuli ( Ṯffl ), 500m, 7–8.VI.2008, leg. F. & L. Kantner ; Ilan : 1♂ ( HNHM), Chilan (棲蘭), 500m, 14–15.IV.1997, leg. G. Csorba & L. Ronkav ; 8♂♂, 7♀♀ ( HNHM), Fushan Botanical Park (福山植物園), 600m, 8–11.IV.2002, leg. O. Merkl ; 1♀ ( NMNS), same locality, 28.VII.2003, leg. M.L. Chan ; 1♂, 4♀♀ ( TARI), same locality, 8.V.2008, leg. S.- F. Yu ; 3♂♂, 9♀♀ ( TARI), same locality, 2–9.VII.2013, leg. Y.- T. Wang ; 1♂ ( HNHM), Suyuanyakou (思源啞口), 1900m, 6.VI.1997, leg. B. Herczig & L. Ronkay ; 1♂ ( TARI), Taipingshan (太平山), 1950m, 26–28.VII.1983, leg. L.Y. Chou ; 1♀ ( NMNS), same locality, 15.V.1999, leg. C. W. & L. B. O’Brien ; 1♂, 2♀♀ ( TARI), Yulan (玉蘭), 300m, 4.VI.2017, leg. Y.- T. Wang ; Kaohsiung: 1♀ ( TARI), Chungchihkuan (中之觀), 1900m, 13.X.2012, leg. L.- P. Hsu ; 1♀ ( MCSN), Chiasien (= Kosempo , 甲仙), 250m, IV.1908, leg. H. Sauter ; 2♀♀ ( HNHM), same locality, XI.1908, leg. H. Sauter ; 2♀♀ ( MNHUB), Hoozan (= Fengshan , 鳳山), 50m, 10.III., leg. H. Sauter ; 2♀♀ ( MNHUB), same but with “ 10.V. ” ; 1♀ ( MNHUB), same but with “ 10.XI ” ; 1♀ ( TARI), Maolin (茂林), 400m, 6.VII.2011, leg. M.- H. Tsou ; 3♀♀ (1♀ in JBCB; 2♀♀ in NMPC), Meishan (梅山), 950m, 12.VI.2008, leg. F. & L. Kantner ; 4♂♂ ( SEHU), Paoshan (寶山), 23.IV.1978, leg. W.- L. Chen ; 2♀♀ ( SEHU), Shanping (扇平), 750m, 26–28.IV.1978, leg. Y. Komiya ; 1♀ ( SEHU), same locality, 27.IV.1981, leg. S. Tsuyuki ; 2♂♂, 2♀♀ ( HNHM), same locality, 1.IV.2003, leg. L. Pappo & M. Földván ; 1♂ ( TARI), same locality, 22.III.2014, leg. W.- C. Liao ; 1♂, 3♀♀ ( TARI), same locality, 12.IV.2014, leg. W.- C. Liao ; 1♂, 3♀♀ ( TARI), same locality, 11.IV.2015, leg. W.- C. Liao ; 1♂, 1♀ ( TARI), same locality, 2.V.2015, leg. W.- C. Liao ; 1♀ ( TARI), Tengchi (藤枝), 1500m, 2–5.VI.2008, leg. C.- F. Lee ; 1♂ ( TARI), same locality, 26.V.2009, leg. C.- F. Lee ; 1♂ ( TARI), same locality, 12.III.2013, leg. Y.- T. Chung ; 1♂ ( TARI), same locality, 31.III.2013, leg. W.- C. Liao ; 1♂, 1♀ ( TARI), same locality, 13.IV.2013, leg. W.- C. Liao ; 1♂ ( TARI), same locality, 18.IV.2013, leg. Y.- T. Chung ; 1♂ ( TARI), same locality, 8.VI.2013, leg. W.- C. Liao ; 2♀♀ ( TARI), same locality, 10.VIII.2013, leg. W.- C. Liao ; 1♂ ( TARI), Tona trail (多納林道), 450m, 12.III.2013, leg. B.- X. Guo ; 1♀ ( TARI), same locality, 14.V.2016, leg. B.- X. Guo ; Keelung: 5♀♀ ( TARI), Hungtanshan (紅淡山), 210m, 10.V.2008, leg. S.- F. Yu ; Miaoli: 3♂♂, 3♀♀ ( TARI), Hsuehchien (雪見), 1800m, 7.VI.2013, leg. W.- B. Yeh ; 1♂ ( TARI), Luchang (鹿場), 880m, 1.VI.2014, leg. Y.- M. Weng ; 2♀♀ ( NMNS), Mingteshuiku (明德水庫), 50m, 1.V.1992, leg. C.Y. Li ; 1♂, 1♀ ( TARI), Sanyi (三義), 400m, 30.III.2010, W.- T. Liu ; 1♀ ( TARI), Tunglo (銅 鑼), 500m, 25.IV.2008, leg. H.- T. Shih ; Nantou: 1♀ ( TARI), Chingching (清境), 1700m, 27.VII.2013, leg. W.- C. Liao ; 2♂♂ ( NMNS), Chunyang (春陽), 1200m, 25–27.V.1993, leg. C.S. Lin & W. T. Yang ; 1♀ ( NMNS), same but with “ 12.VIII.–8.IX.1998 ” ; 1♂, 1♀ ( NMNS), same but with “ 12.III.–9.IV.2002 ” ; 2♂♂ ( NMNS), same but with “ 9.IV.–7.V.2002 ” ; 1♂, 5♀♀ ( NMNS), same but with “ 7.V.–11.VI.2002 ” ; 1♂, 4♀♀ ( NMNS), same but with “ 11.VI.–9.VII.2002 ” ; 2♀♀ ( NMNS), same but with “ 9.VII.–13.VIII.2002 ” ; 1♀ ( NMNS), same but with “11,VI.– 8.VII.2003 ” ; 1♂, 3♀♀ ( NMNS), same but with “ 8.VII.–5.VIII.2003 ” ; 1♂, 5♀♀ ( NMNS), same but with “ 11.V.– 13.VII.2004 ” ; 1♂, 4♀♀ ( NMNS), same but with “ 13.VII.–10.VIII.2004 ” ; 2♂♂, 6♀♀ ( NMNS), same but with “ 6.VI.–25.VII.2006 ” ; 1♀ ( NMNS), same but with “ 25.VII.–22.VIII.2006 ” ; 1♀ ( NMNS), same but with “ 17.X.– 14.XI.2006 ” ; 1♂ ( NMNS), same but with “ 10.IV.–8.V.2007 ” ; 1♂ ( NMNS), same but with “ 8.V.–6.VI.2007 ” ; 1♂, 3♀♀ ( NMNS), same but with “ 8.VI.–14.VII.2007 ” ; 1♀ ( NMNS), Fenghuangku (鳳凰谷), 700m, 15.IV.1994, leg. M.L. Chan & W. T. Yang ; 4♀♀ ( NMNS), Hoawanta (奧萬大), 1100m, 30.IV.1992, leg. W.T. Yang ; 1♀ ( TARI), Hsitou (溪頭), 1150m, 15.VI.2016, leg. Y.- N. Lee ; 1♀ ( TARI), Huakang (華崗), 2600m, 26.VII.2010, leg. M.- H. Tsou ; 1♀ ( TARI), same locality, 13.IX.2010, leg. C.- F. Lee ; 2♀♀ ( SEHU), Huisun Forest Area (惠蓀林場), 800m, 28.IV.1991, leg. A. Tanaka ; 1♀ ( NMNS), same locality, 21.V.1999, leg. C. W. & L. B. O’Brien ; 4♀♀ ( EUMJ), same locality, 28–30.IV.2010, leg. M. Nishikawa ; 2♂♂ ( TARI), same locality, 3.IV.2014, leg. F.- S. Huang ; 1♂ ( EUMJ), Jihyuehtan (日月潭), 700m, 14.IV.1976, leg. K. Ushijima ; 1♂, 6♀♀ ( SEHU), same localty (= Tehwashe , 德化社), 3.V.1977, leg. Y. Komiya ; 1♂, 4♀♀ ( SEHU), same but with “ 11.V.1977 ” ; 2♀♀ ( EUMJ), same locality, 23.IV.1978, leg. K. Murakami ; 2♀♀ ( SEHU), same locality, 3.V.1978, leg. Y. Komiya ; 2♀♀ ( SEHU), same but with “ 9.V.1978 ” ; 2♂♂, 1♀ ( SEHU), same locality, 30.IV.1981, leg. S. Tsuyuki ; 2♀♀ ( EUMJ), same locality, 27.V.1981, leg. K. Sasagawa ; 1♂ ( SEHU), Kuantaohsi (關刀溪), 800m, 15.V.1984, leg. A. Tanaka ; 1♀ ( SEHU), Kuantaoshan (關刀山), 1200m, 17.V.1985, leg. S. Yoshimatsu ; 3♀♀ ( TARI), same locality, 7.IV.2012, leg. M.- H. Tsou ; 1♂, 1♀ ( SEHU), Lienhuachi (連華池), 650m, 6.V.1977, leg. Y. Komiya ; 6♀♀ ( SEHU), same but with “ 6- 7.V.1978 ” ; 1♂, 13♀♀ ( TARI), same locality, 23–26.V.1980, leg. K.S. Lin & B. H. Chen ; 2♀♀ ( NMNS), same locality, 9.IV.–19.V.1998, leg. C.S. Lin & W. T. Yang ; 5♂♂, 1♀ ( NMNS), same locality, 9–11.IV.2001, leg. C.S. Lin ; 1♂, 3♀♀ ( NMNS), same locality, 9.IV.–2.V.2001, leg. C.S. Lin & W. T. Yang ; 3♀♀ ( NMNS), same but with “ 2.V.–12.VI.2001 ” ; 3♀♀ ( NMNS), same but with “ 8.IV.–6.V.2002 ” ; 1♀ ( NMNS), same but with “ 9.VII.– 12.VII.2002 ” ; 1♀ ( NMNS), same locality, 9.IX.2002, leg. C.S. Lin ; 1♀ ( NMNS), same locality, 4.III.–6.V.2003, leg. C.S. Lin & W. T. Yang ; 1♀ ( NMNS), same but with “ 5.IV.–10.V.2004 ” ; 1♀ ( NMNS), same but with “ 10.V.– 12.VII.2004 ” ; 1♀ ( NMNS), same but with “ 9.IX.–4.X.2004 ” ; 1♀ ( NMNS), same but with “ 7.VIII.–11.IV.2005 ” ; 1♀ ( NMNS), same but with “ 11.IV.–2.V.2005 ” ; 1♂ ( NMNS), same locality, 6.IV.2005, leg. Y.C. Hsuen ; 1♂, 1♀ ( TARI), same locality, 27.III.2008, leg. C.- F. Lee ; 2♂♂, 1♀ ( TARI), same locality, 3.VI.2009, leg. C.- F. Lee ; 2♂♂, 2♀♀ ( TARI), same locality, 10.III.2013, leg. W.- C. Liao ; 1♀ ( SEHU), Lushan ( AEƜ), 1000m, 5.V.1978, leg. Y. Komiya ; 14♂♂, 23♀♀ ( TARI), same locality, 27–31. V.1 980, leg. K.S. Lin & L. Y. Chou ; 6♂♂ ( TARI), Meifeng ( Ñó ), 2130m, 17–22.VI.1979, leg. K.S. Lin & B. H. Chen ; 1♂ ( TARI), same locality, 8.VI.1980, leg. K.S. Lin & B. H. Chen ; 4♂♂ ( TARI), same locality, 24–26.VI.1981, leg. K.S. Lin & W. S. Tang ; 1♂ ( TARI), same locality, 15.VII.1982, leg. S.C. Lin & C. N. Lin ; 1♂ ( NMNS), same locality, 8–9.III.1999, leg. C.S. Lin & W. T. Yang ; 2♂♂ ( NMNS), same but with “ 13.VI.–18.VII.2001 ” ; 1♂ ( NMNS), same but with “ 11.V.–13.VII.2004 ” ; 1♀ ( TARI), same locality, 20.VI.2009, leg. Y.- T. Wang ; 1♂ ( TARI), same locality, 19.VI.2010, leg. C.- F. Lee ; 1♀ ( NMNS), Nanshanchi (ṀƜffi), 800m, 5.VI.1965, leg. B. S. Chang ; 1♀ ( EUMJ), same locality, 19.V.1970, leg. M. Takagi ; 1♂ ( SEHU), same locality, 1.IV.1976, leg. H. Ono ; 3♂♂ ( EUMJ), same locality, 25–26.III.1977, leg. Y. Notsu ; 1♀ ( SEHU), same locality, 21.V.1974, leg. Akiyama ; 1♂, 2♀♀ ( SEHU), same locality, 17.IV.1977, leg. W. Suzuki ; 1♂ ( SEHU), same locality, 25.IV.1977, leg. S. Saito ; 3♂♂, 22♀♀ ( SEHU), same locality, 29–30.IV.1977, leg. Y. Komiya ; 1♀ ( SEHU), same locality, 4.V.1977, leg. W. Suzuki ; 2♂♂, 4♀♀ ( SEHU), same locality, 7–10.V.1977, leg. Y. Komiya ; 1♀ ( SEHU), same locality, 23.VII.1977, leg. K. Kusama ; 3♀♀ ( SEHU), same locality, 2.V.1978, leg. Y. Komiya ; 1♂ ( EUMJ), same locality, 11.V.1978, leg. N. Yashiro ; 2♂♂, 1♀ ( HNHM), same locality (near Mong Gwu, ḄỖ), 20.IV.2002, leg. D. A. Anstine, G. Fábián & O. Merkl ; 1♂ ( NMNS), same locality, 27.III.2003, leg. C.S. Lin ; 4♀♀ ( TARI), same locality, 10.VII.2010, leg. Y.- T. Wang ; 1♀ ( NMNS), Peikang (北港), 400m, 12.IV.2003, leg. M.L Chan ; 3♀♀ ( TARI), Penpuhsi (本部溪), 400m, III.1971, leg. T.C. Maa ; 3♀♀ ( TARI), Shalihsien trail (沙里仙林道), 1000m, 9–11.VI.2013, leg. Y.- T. Wang ; 6♀♀ ( TARI), Shanlinhsi (杉林溪), 1600m, 15.IV.2014, leg. J.- C. Chen ; 1♂ ( TARI), Shenmutsun (神木村), 1500m, 17.V.2010, leg. Y.- T. Wang ; 1♀ ( TARI), Tatachia (塔塔加), 2600m, 9.VI.2009, leg. C.- F. Lee ; 2♀♀ ( TARI), Tungpu (東埔), 1200m, 20–22.VI.1980, leg. C.C. Chen ; 2♂♂ ( TARI), same locality, 28.IV.–2.V.1981, leg. T. Lin & C. J. Lee ; 10♀♀ ( TARI), same locality, 19– 23.VII.1982, leg. L.Y. Chou & T. Lin ; 6♂♂, 4♀♀ ( TARI), same locality, 20–24.VI.1983, leg. K.C. Chou & C. Y. Wong ; 90♂♂, 101♀♀ ( TARI), same locality, 23–27.VII.1984, leg. K.C. Chou & C. H. Yang ; 1♀ ( SMNS), same locality (= Tongpo ), 16–18.VII.1986, leg. H. Takizawa ; 1♀ ( SEHU), same locality, 14.V.1998, leg. A. Tanaka ; 1♀ ( TARI), same locality, 27.VII.2013, leg. J.- C. Chen ; 3♂♂, 2♀♀ ( TARI), Tungfu (同富), 800m, 27.IV.2010, leg. C.- F. Lee ; 1♀ ( HNHM), Wucheng (五城, = Fuhosho ), VI.1908, leg. H. Sauter ; 1♂ ( NMNS), Wushe (霧社), 1150m, 15.VII.1966, leg. B. S. Chang ; 1♂ ( MCSN), same locality (= Musha ), 18.V.–15.VI.1919, leg. T. Okuni, J. Sonan, K. Miy., M. Yosh. ; 1♂, 3♀♀ ( EUMJ), same locality, 24.V.1972, leg. M. Sakai ; 2♀♀ ( SEHU), same locality, 7.V.1975, leg. M. Kubota ; 1♂ ( SEHU), same locality, 12.V.1977, leg. Y. Komiya ; 1♂ ( EUMJ), same locality, 26.IV.1978, leg. K. Murakami ; 5♀♀ ( TARI), same locality, 19–23.VI.1979, leg. K.S. Lin & B. H. Chen ; 3♀♀ ( TARI), same locality, 6–11.V.1981, leg. K.S. Lin & S. C. Lin ; 2♀♀ ( TARI), same locality, 23–28.VI.1981, leg. K.S. Lin & W. S. Tang ; 1♀ ( TARI), same locality, 4.VIII.1981, leg. T. Lin & W. S. Tang ; 2♀♀ ( TARI), same locality, 14.VII.1982, leg. S.C. Lin & C. N. Lin ; 1♂ ( TARI), same locality, 19–22.IV.1983, leg. K.C. Chou & S. P. Huang ; 1♂, 2♀♀, same locality, 7.V.1984, leg. K.C. Chou & C. C. Pan ; 1♀ ( TARI), same locality, 17.VIII.1984, leg. K.C. Chou ; 1♂ ( NMNS), same locality, 30.VI.1998, leg. C.S.Lin & W. T. Yang ; 1♂, 1♀ ( NMNS), same locality, 19.V.1998, leg. C.S. Lin ; 1♀ ( NMNS), same but with “ 9.IV.2002 ” ; 1♀ ( NMNS), same but with “ 7.V.2003 ” ; 1♀ ( NMNS), Yuanfeng (鳶峰), 2750m, 13.VIII.–10.IX.2002, leg. C.S. Lin & W. T. Yang ; 6♂♂, 17♀♀ ( TARI), Yushih ( Njǿ ), 1750m, 4.VIII.1981, leg. T. Lin & W. S. Tang ; 1♀ ( NMNS), same locality, 13.VII.1987, leg. I.C. Hsu ; Pingtung: 6♀♀ ( TARI), Kaoshih (高士), 100m, 30.IV.2011, leg. J.- C. Chen ; 1♂ ( TARI), Lilungshan (里龍山), 900m, 9.IV.2013, leg. J.- C. Chen ; 1♀ ( TARI), same locality, 26.V.2014, leg. Y.- T. Chung ; 1♂, 1♀ ( TARI), Machia (瑪家), 1100m, 25.V.2016, leg. Y.- T. Chung ; 1♀ ( HNHM), Mutan (牡丹), 400m, 7–8.IV.1997, leg. G. Csorba & L. Ronkay ; 1♂ ( TARI), Mutanchihshan (牡丹池山), 500m, 20.II.2007, leg. S.- F. Yu ; 1♂ ( TARI), same locality, 10.III.2011, leg. J.- C. Chen ; 1♂ ( TARI), same locality, 13.III.2012, leg. C.- C. Wang & Y.- C. Lan ; 1♀ ( TARI), Neiwen (內文), 800m, 24.II.2012, leg. J.- C. Chen ; 27♂♂, 43♀♀ ( TARI), Peitawushan (北大武山) Hiking Gate , 1500m, 7–8.IV.2013, leg. Y.- T. Chung ; 1♂, 1♀ ( TARI), same locality, 12.IV.2013, leg. Y.- T. Chung ; 9♂♂, 13♀♀ ( TARI), 22.IV.2014, leg. Y.-T. Chung ; 1♂, 2♀♀ ( TARI), same but with “leg. J.-C. Chen” ; 3♀♀ ( TARI), same locality, 8.V.2014, leg. Y.- T. Chung ; 1♂ ( TARI), same locality, 28.V.2014, leg. Y.- T. Chung ; 2♂♂ ( TARI), same locality, 21.III.2015, leg. Y.- T. Chung ; 1♂ ( NMNS), Shihtzu (獅子), 50m, 18.IV.2005, leg. J.H. Chen ; 2♀♀ ( TARI), Shouka (壽卡), 400m, 22.III.2009, leg. M.- H. Tsou ; 1♂ ( TARI), same locality, 16.IV.2010, leg. U. Ong ; 1♀ ( TARI), same locality, 28.II.2013, leg. W.- C. Liao ; 1♂, 5♀♀ ( TARI), same locality, 28.II.2013, leg. B.- X. Guo ; 1♂ ( TARI), same locality, 26.IV.2013, leg. Y.- T. Chung ; 1♂, 1♀ ( TARI), same locality, 30.IV.2013, leg. Y.- T. Chung ; 1♂ ( TARI), same locality, 17.II.2014, leg. J.- C. Chen ; 1♀ ( TARI), same locality, 7.III.2016, leg. Y.- T. Chung ; 1♀ ( TARI), same locality, 7.V.2016, leg. Y.- T. Chung ; 1♂, 1♀ ( TARI), Shuangliu (雙流), 400m, 14.III.2018, leg. Y.- T. Chung ; 1♂ ( TARI), Suchunghsi (四重溪), 50m, 13.V.2013, leg. Y.- T. Chung ; 1♀ ( TARI), Tahanshan (大漢山), 1000m, 18.VII.2007, leg. M.- H. Tsou ; 3♀♀ ( TARI), same locality, 25.V.2008, leg. C.- F. Lee ; 1♀ ( TARI), same locality, 4.VII.2008, leg. M.- H. Tsou ; 1♀ ( TARI), same locality, 7.IX.2009, leg. J.- C. Chen ; 2♀♀ ( TARI), same locality, 14.VIII.2011, leg. Y.- T. Wang ; 1♀ ( TARI), same locality, 28.IV.2012, leg. M.- H. Tsou ; 2♂♂, 2♀♀ ( TARI), same locality, 6.VI.2012, leg. C.- F. Lee ; 1♀ ( EUMJ), same locality, 7.VII.2012, leg. N. Ohbayashi ; 2♂♂, 1♀ ( TARI), same locality, 19.VII.2012, leg. C.- F. Lee ; 1♂, 1♀ ( TARI), same locality, 16.IV.2013, leg. Y.- T. Chung ; 1♀ ( TARI), same locality, 9.VI.2013, leg. Y.- T. Chung ; 1♂ ( TARI), same locality, 12.VI.2013, leg. Y.- T. Chung ; 1♀ ( TARI), same locality, 3.VII.2013, leg. B.- X. Guo ; 1♂ ( TARI), same locality, 17.VIII.2013, leg. J.- C. Chen ; 1♂, 1♀ ( TARI), same locality, 3.IX.2013, leg. Y.- T. Chung ; 1♀ ( TARI), same but with “leg. J.-C. Chen” ; 1♂, 1♀♀ ( TARI), same locality, 30.V.2014, leg. Y.- T. Chung ; 1♂ ( TARI), same locality, 10.VI.2014, leg. Y.- T. Chung ; 1♂, 2♀♀ ( TARI), Tungyuan (東源), 300m, 27.III.2012, leg. J.- C. Chen ; Taichung: 2♂♂, 1♀ ( TARI), Anmashan (鞍馬山) (= Tahsuehshan , 大雪山), 2275m, 24.IV.2012, leg. C.- F. Lee ; 2♀♀ ( TARI), same locality, 5.VI.2012, leg. J.- C. Chen ; 1♀ ( NMNS), Kukuan (谷關), 730m, 11–12.IV.1986, leg. C.S. Lin ; 1♀ ( TARI), same locality, 1.XI.2009, leg. M.- H. Tsou ; 1♂ ( TARI), Lishan (梨山), 1900m, 26.VI.1979, leg. K.S. Lin & L. Y. Chou ; 3♂♂, 10♀♀ ( TARI), Wuling (武陵), 1900m, 27–29.VI.1979, leg. K.S. Lin & L. Y. Chou ; 1♂ ( NMNS), Wushihkeng (烏石坑), 800m, 16–17.III.1995, leg. W.T. Yang ; 1♀ ( NMNS), same locality, 2 7–28.X.2003, leg. W.T. Yang & K. W. Huang ; 6♂♂, 1♀ ( TARI), same locality, 21.III.2013, leg. C.- F. Lee ; 1♀ ( TARI), same locality, 29.VIII.2014, leg. C.- F. Lee ; Tainan: 3♂♂ ( TARI), Kantoushan (崁頭山), 800m, 23.III.2013, leg. W.- C. Liao ; 4♂♂, 1♀ ( TARI), same locality, 29.III.2014, leg. W.- C. Liao ; 1♀ ( USNM), Kwantzuling (關子嶺), 350m, 26– 28.VI.1980, leg. D. R. Davis ; 1♂ ( TARI), Meiling (梅嶺), 800m, 22.VI.2010, leg. U. Ong ; 2♀♀ ( TARI), same locality, 24.IV.2013, leg. B.- X. Guo ; 1♂, 2♀♀ ( TARI), same locality, 19.IV.2014, leg. W.- C. Liao ; 1♀ ( TARI), same locality, 11.V.2014, leg. B.- X. Guo ; 1♂ ( TARI), Pichien trail (碧尖步道), 800m, 22.III.2010, leg. U. Ong ; Taipei: 2♀♀ ( USNM), Ankung (安康), 50m, 8.VIII.1992, leg. D. G. Furth ; 1♀ ( TARI), Chihshanyan (芝山岩), 50m, 20.III.2014, leg. S.- F. Yu ; 1♂, 3♀♀ ( HNHM), Guanyinshan (觀音山), 500 m, 14–21.IV.2002, leg. G. Fábián ; 1♀ ( TARI), Hsichih (汐止), 150m, 29.IV.2007, leg. H.- J. Chen ; 1♀ ( HNHM), Pinglin (坪林), 600m, 17.IV.2003, leg. L. Papp ; 5♂♂ ( TARI), same locality, 29.III.2008, leg. M.- H. Tsou ; 1♂, 1♀ ( TARI), Shenking (深 坑), 400m, 9.IV.2009, leg. S.- F. Yu ; 1♀ ( TARI), Taipei Zoo (台北市立動物園), 2.V.2007, Y.- C. Yu ; 1♀ ( TARI), Tatungshan (大桶山), 900m, 3.IV.2008, leg. S.- F. Yu ; 4♀♀ ( EUMJ), Wulai (烏來), 800m, 4.VI.1970, leg. Y. Hori ; 3♂♂ ( HNHM), same locality, 7.IV.2002, leg. G. Fábián & O. Merkl ; 1♀ ( TARI), same locality, 5.IV.2007, leg. M.- H. Tsou ; 2♀♀ ( TARI), same locality, 20.VII.2007, leg. H.- J. Chen ; 1♂, 1♀ ( TARI), same locality, 3.IV.2008, leg. H.- J. Chen ; 1♀ ( TARI), same locality, 26.VI.2009, leg. C.- F. Lee ; 1♀ ( TARI), same locality (= Hsinhsien , 信賢), 11.V.2014, leg. H.- J. Chen ; Taitung: 1♂, 1♀ ( TARI), Chipen (知本), 300m, 24.V.2013, leg. J.- C. Chen ; 4♂♂, 1♀ ( TARI), Hsiangyang (向陽), 2300m, 9.V.2013, leg. J.- C. Chen ; 1♂ ( TARI), Lichia trail (利嘉林道), 500m, 12.VI.2010, leg. Y.- L. Lin ; 2♂♂, 1♀ ( TARI), same locality, 15.VII.2014, leg. Y.- T. Chung ; 1♂, 10♀♀ ( TARI), same locality, 16.VII.2014, leg. Y.- T. Wang ; 5♂♂, 5♀♀ ( TARI), Liyuan (栗園), 1800m, 23.VI.2010, leg. M.- H. Tsou ; 1♀ ( TARI), same locality, 19.VI.2013, leg. C.- F. Lee ; 3♀♀ ( TARI), Motien (摩天), 1500m, 23.VI.2010, leg. M.- H. Tsou ; 2♂♂ ( TARI), same locality, 28.VI.2013, leg. J.- C. Chen ; 2♂♂ ( EUMJ), Yenping trail (延平林道), 500m, 3.VII.2012, leg. J. Yamasako ; Taoyuan: 1♂, 2♀♀ ( TARI), Fuhing (復興), 500m, 6.V.1983, leg. K.C. Chou & C. C. Pan ; 1♂ ( TARI), Hsiaowulai (小烏來), 400m, 20.III.2008, leg. S.- F. Yu ; 3♀♀ ( TARI), same locality, 19.IV.2008, leg. S.- F. Yu ; 1♀ ( TARI), Hsuanyuan (萱源), 850m, 1.VI.2010, leg. W.- T. Liu ; 1♀ ( TARI), same locality, 10.V.2010, leg. Y.- L. Lin ; 1♀ ( TARI), same locality, 13.V.2010, leg. S.- F. Yu ; 1♀ ( TARI), Hsuehwunao (雪霧鬧), 1450m, 3.IV.2011, leg. M.- H. Tsou ; 1♀ ( TARI), Lalashan (拉拉山), 2030m, 4.V.2010, leg. S.- F. Yu ; 1♂ ( TARI), Linkou (林口), 250m, 8.IV.2009, leg. H.- H. Han ; 1♀ ( SEHU), Paling (巴陵), 800m, 1.V.1981, leg. S. Tsuyuki ; 2♀♀ ( TARI), same locality, 3–5.V.1983, leg. K.C. Chou & C. C. Pan ; 1♂ ( SEHU), same locality, 13.V.1984, leg. Y. Komiya ; 1♀ ( SEHU), same locality, 11.V.1998, leg. A. Tanaka ; 1♂ ( TARI), same locality, 28.III.2010, leg. M.- H. Tsou ; 1♂ ( SEHU), Suleng (= Hsileng, M ḃ), 1100m, 17.V.1984, leg. Y. Komiya ; 2♂♂, 1♀ ( TARI), 5.V.2010, leg. Y.-L. Lin ; 1♂, 1♀ ( TARI), Tungyanshan (東眼山), 850m, 12.IV.2007, leg. M.- H. Tsou ; 1♀ ( TARI), same locality, 8.VII.2007, leg. S.- F. Yu ; 2♀♀ ( TARI), same locality, 16.VI.2009, leg. S.- F. Yu .

Redescription. Length 5.0– 5.7 mm, width 1.9–2.2 mm. General color ( Figs 4 D–4F) metallic blue; mouth parts brown or dark brown; legs and antennae pale yellow, sometimes femora, apices of tibiae and tarsi darkened. Vertex with reticulate microsculpture. Antennae filiform in males ( Fig. 19A ), antennomeres III–X slightly curved, length ratios of antennomeres I–XI 1.0: 0.2: 0.9: 1.2: 1.2: 1.2: 1.2: 1.1: 1.1: 1.1: 1.2, length to width ratios of antennomeres I–XI 2.6: 1.0: 2.7: 3.6: 3.8: 3.6: 3.8: 3.7: 4.3: 4.3: 5.1; slender in females ( Fig. 19B ), antennomeres III–X straight, length ratios of antennomeres I–XI 1.0: 0.3: 0.7: 0.9: 1.0: 0.8: 0.8: 0.8: 0.8: 0.8: 0.8, length to width ratios of antennomeres I–XI 3.5: 1.9: 3.8: 5.1: 5.3: 4.7: 4.6: 4.4: 4.4: 4.3: 4.6. Pronotum transverse, 1.2–1.3x wider than long, disc with one pair of lateral depressions, reticulate microsculpture, and with dense, coarse punctures; lateral margin basally narrowed; apical and basal margins subtruncate. Elytra elongate and parallel-sided, 1.9x longer than wide; disc with reticulate microsculpture, dense, coarse punctures, arranged into longitudinal rows, and with distinctly longitudinal ridges between longitudinal rows of punctures. Apical margin of abdominal ventrite V in females ( Fig. 19H ) rounded, but slightly depressed near medial area. Tarsomeres I of front legs swollen in males ( Fig. 19I ), subparallel in females ( Fig. 19J ). Penis ( Figs 19 C–19E) extremely slender, 10.4x longer than wide; apex lanceolate, with medial, shallow, narrow notch, parallel-sided; tectum extremely elongate from apical 1/10 to basal 1/3, apically narrowed; straight in lateral view, slightly curved near base; triangular sclerites small; internal sac with elongate endophallic sclerite complex, 0.6x as long as penis, composed of two sclerites, basal piece much longer (1.4x) than apical piece, apical sclerite with acute apex, dorsal sclerite with apex acute and hook-like in lateral view; ventral sclerites absent. Gonocoxae ( Fig. 19F ) elongate, slightly broadened at apical 1/6, connected from basal 1/3 to apical 1/5; apices narrowly rounded, each gonocoxa with eight setae along outer margin from apex to apical 1/6; with one pair of short lateral processes at basal 2/5. Ventrite VIII ( Fig. 19G ) longitudinal and well sclerotized; disc with several long setae at sides and near apical margin, and with dense, short setae along apical margin; spiculum extremely slender. Receptacle of spermatheca ( Fig. 19K ) strongly swollen; pump slender and strongly curved; proximal spermathecal duct deeply inserted into receptacle, broad and short.

Variation. Some specimens from southern Taiwan have metallic green bodies ( Figs 4H, 4I ). Specimens at high altitudes (more than 2000 m) have deep metallic purple bodies and dark brown legs with black tibiae and tarsi ( Fig. 4G ).

Diagnosis. Males of T. sauteri Chûjô are similar to those of T. geiseri sp. nov. based on absence of ventral sclerites (present in others), longer basal piece (subequal or short than apical piece in others) but are characterized by the dorsal sclerite separate from the basal and apical pieces ( Figs 19 C–19E).

Food plants. Rosaceae : Prunus campanulata ; P. phaeosticta ; Moraceae : Ficus sarmentosa var. nipponica ; Fagaceae : Pasania hancei var. ternaticupula f. naticupula ; Lythraceae : Lagerstroemia subcostata ; Actinidiaceae : Actinidia callosa ; Myricaceae : Myrica rubra .

Distribution. Endemic to Taiwan. Theopea sauteri is one of most widespread species of the genus in Taiwan. Adults appear at various elevations ( Fig. 10C ) and almost year round except January and December. Most adults were collected in lowlands (0~ 1500 m) (88%) and from April to July (87%).

Figures

FIGURES 4. Habitus of Theopea cheni sp. nov. and T. sauteri Chûjô. 4A. T. cheni sp. nov., male, dorsal view; 4B. Same, ventral view; 4C. T. cheni sp. nov., female, dorsal view; 4D. T. sauteri, male, Mutanchihshan (ËṚḾƜ), dorsal view; 4E. Same, ventral view; 4F. T. sauteri, male, Liyuan (Mȁ), dorsal view; 4G. T. sauteri, female, Liyuan (Mȁ), dorsal view; 4H. T. sauteri, male, Lilungshan (BÊƜ), dorsal view; 4I. Same, ventral view.

FIGURES 10. Distribution maps of Theopea irregularis Takizawa, T. kanmiyai (Kimoto), and T. sauteri Chûjô. Solid line: 1000 m, broken line: 2000 m. 10A. T. irregularis; 10B. T. kanmiyai; 10C. T. sauteri.

FIGURES 19. Diagnostic characters of Theopea sauteri Chûjô. 19A. Antenna, male; 19B. Antenna, female; 19C. Penis, dorsal view; 19D. Penis, lateral view; 19E. Penis, ventral view; 19F. Gonocoxae; 19G. Abdominal ventrite VIII; 19H. Apical margin of abdominal ventrite V, female; 19I. Tarsi of front leg, male; 19J. Tarsi of front leg, female; 19K. Spermatheca.