Benthoctopus johnsonianus Allcock, Strugnell, Ruggeiro & Collins, 2006

Strugnell, Jan, Voight, Janet R., Collins, Patrick C. & Allcock, A. Louise, 2009, Molecular phylogenetic analysis of a known and a new hydrothermal vent octopod: their relationships with the genus Benthoctopus (Cephalopoda: Octopodidae), Zootaxa 2096 (1), pp. 442-459 : 448-449

Benthoctopus johnsonianus Allcock, Strugnell, Ruggeiro & Collins, 2006

Synonymy: Benthoctopus johnsoniana Allcock, Strugnell, Ruggeiro & Collins, 2006: 379–386 , figs 6–9.

Type material examined: Benthoctopus johnsoniana Allcock, Strugnell, Ruggeiro & Collins, 2006 holotype: BMNH 20060178 , RV Discovery, Stn 50518, 49°27'N 13°21'W, 7 June 1979, 2045–2110 m, male.

Additional material examined: Benthoctopus johnsonianus : * RV Discovery , 49°44'N 13°10'W (NE Atlantic), Stn 14309, 16 March 2002, 2011–2218 m, female, leg. & det. MA Collins. 8 additional specimens detailed in Allcock et al., 2006 : BMNH 20060179-82 , NMSZ 1999158.300 , NMSZ 1994128.46501 /.46503/.46506.

Diagnosis. Suckers closely set. Arms approximately 3.5 times ML. Arm formula 2>1>3>4. Funnel organ W-shaped. Gills with 8–11 lamellae per demibranch. LLI 10–11. Maximum spermatophore length 104, maximum SpLI 92, maximum number of spermatophores 19. Mature males without enlarged suckers. Integument smooth.

Remarks. The ending of the specific name has been corrected herein to agree with the male gender of Benthoctopus .