Trubatsa ganensis, Merle & Pacaud & Ledon & Goret, 2024

Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard, 2024, New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe, Geodiversitas 46 (15), pp. 495-551 : 508

publication ID	https://doi.org/10.5252/geodiversitas2024v46a15
publication LSID	urn:lsid:zoobank.org:pub:C32EAAFD-85D8-45CD-8DF7-B894E1537713
DOI	https://doi.org/10.5281/zenodo.13985812
persistent identifier	https://treatment.plazi.org/id/0EBD7F4D-8646-456B-BD5C-10FA4A8625DD
taxon LSID	lsid:zoobank.org:act:0EBD7F4D-8646-456B-BD5C-10FA4A8625DD
treatment provided by	Plazi (2024-10-24 08:29:49, last updated 2024-11-26 02:03:40)
scientific name	Trubatsa ganensis
status	sp. nov.

Treatment

Trubatsa ganensis View in CoL n. sp.

( Figs 5 View FIG A-D; 25I-K)

urn:lsid:zoobank.org:act:0EBD7F4D-8646-456B-BD5C-10FA4A8625DD

Trubasta [sic] parisiensis – Merle 1990: 169, pl. 3, figs 13-15, pl. 4, fig. 12 [non d’Orbigny, 1850].

TYPE MATERIAL. — Holotype. France • Aquitaine Basin, Pyrénées-Atlantiques, Gan ( Tuilerie ); Uppermost part of the Marnes de Gan only (top biozone N12/base biozone NP13); Early Eocene ( Ypresian ); MNHN.F.A90520 (Varone coll.), H: 17 mm ( Figs 5A, B View FIG ; 25I View FIG ).

Paratype. France • 1 spm; same as for the holotype; MNHN.F.R61441 ( Merle coll., figured specimen of Merle (1990: pl. 3, fig. 13) • 1 spm; idem; MNHN.F.A90518 ( Merle coll., with complete siphonal canal, Figs 5C View FIG ; 25J View FIG ) • 1 spm; idem; MNHN.F.A90517 ( Merle coll. with incomplete siphonal canal; Figs 5D View FIG ; 25K View FIG ) • 12 spm; idem; MNHN.F.A90519 ( Merle coll.) • 17 spm; idem; MNHN.F.A72383 (Varone coll.).

ADDITIONAL MATERIAL. — 4 spm; idem;MNHN.F.A90521 (Combault coll.).

ETYMOLOGY. — From the type locality, Gan (Pyrénées-Atlantiques).

TYPE HORIZON. — Uppermost part of the Marnes de Gan only (top biozone N12/base biozone NP13), see Nolf et al. (2002: 173, fig. 4), Early Eocene (Ypresian).

TYPE LOCALITY. — France, Aquitaine Basin, Pyrénées-Atlantiques, Gan (Tuilerie) ( Merle 1986: 8; Nolf et al. 2002: 185, 186).

DISTRIBUTION. — Only known from the type locality.

DESCRIPTION

Bulbous protoconch of 1.5-2 whorls (see Merle 1990: pl. 4, fig. 12). Teleoconch up to 17 mm (canal siphonal not complete) in height, up to 11 mm in width, biconic in profile, composed of five whorls. Moderately high spire. Last whorl representing 72% of total height. Spiral angle 56°. Suture weakly impressed. Axial sculpture consisting in four slightly foliate varices per whorl. On spire, anal tube (P1) and very short spine (P2). Anal tube forming angle of approximately 60-65° with axis of shell. Anal tube (P1) totally integrated within varices, eliminating shoulder. Small callus at base of anal tube. On convex part of last whorl and abapically to anal tube four spinelets (P2, s2, P3, P4 and P5). Intervarical spaces usually smooth, but with P3-P5 cords on holotype. Rounded aperture up to 25% of diameter, up to 74% of height of last whorl (including siphonal canal). Edge, erect, smooth, peristome complete. Siphonal canal closed, usually broken, occupying 60% of apertural lenght (complete siphonal canal longer, see Fig. 5C View FIG ).

COMPARISONS

Merle (1990: 169) attributed this material collected at Gan to Typhis parisiensis d’Orbigny, 1850 View in CoL and to the genus Trubatsa Dall, 1889 View in CoL . The generic attribution is correct and the distinction between Siphonochelus Jousseaume, 1880 View in CoL and Trubatsa View in CoL is now validated by molecular data ( Houart et al. 2021). Nevertheless, after further comparison, specific attribution of the Gan material to this species incorrect. Firstly, the protoconchs are different, that of T. ganensis n. sp. is paucispiral ( Merle 1990: pl. 4, fig. 12) and that of T. parisiensis View in CoL is multispiral (see juvenile specimen from Barton illustrated by A. Morton (2023; www.dmap.co.uk/fossils). In addition, the Bartonian specimens are more sculptured than the specimens from Gan ( Fig. 5E, F View FIG ). They display stronger spines (P2 to P5) and P6, spiral cords (P2 to P5) in the intervarical spaces and a partition recalling that of T. gaasensis (Tournouër in Benoist, 1880) from the Rupelian of Aquitaine Basin. Trubatsa parisiensis View in CoL occur in the Bartonian from England and in the Priabonian from Germany ( von Koenen 1889). Merle (1985) also suggested that Typhis sinuosus Cossmann, 1902 from the Bartonian of Bois-Gouët ( France, Loire-Atlantique) was a junior subjective synonym of T. parisiensis View in CoL . According to Merle (1985), the shape of the varices of the type specimen (Dumas coll.) of T. sinuosus recalls that of Trubatsa View in CoL . However, it is a juvenile (H: 8 mm) and the varices of the juveniles of Typhina rutoti Cossmann, 1882 View in CoL are also similar. Moreover, the type specimen of T. sinuosus is worn, lacks a protoconch and finally lacks diagnostic characters. Thus, we consider T. sinuosus as a nomen dubium.

References

COSSMANN M. 1902. - Mollusques eoceniques de la Loire-inferieure - Fascicule 1. Bulletin de la Societe des Sciences naturelles de l'Ouest de la France 2 (2): 55 - 209.

HOUART R., BUGE B. & ZUCCON D. 2021. - A taxonomic update of the Typhinae (Gastropoda: Muricidae) with a review of New Caledonia species and the description of new species from New Caledonia, the South China sea and Western Australia. Journal of Conchology 44 (2): 103 - 147.

JOUSSEAUME F. P. 1880. - Division methodique de la famille des Purpurides. Le Naturaliste 2 (42): 335 - 338. http: // biodiversitylibrary. org / page / 33928744

MERLE D. 1985. - Les communautes fossiles en tant qu'indicateurs paleobiologiques de milieux: exemples pris sur le Cuisien de Gan (Pyrenees-Atlantiques) et l'Egerien de Hongrie. Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser. 7, sect. C, 24: 257 - 262.

MERLE D. 1986. - Contribution a l'etude paleontologique du gisement cuisien de Gan (Pyrenees Atlantiques): systematique, evolution et paleoecologie. Unpublished Memoir of the Ecole pratique des Hautes Etudes (EPHE), 2 vols, Section Sciences de la Vie et de la Terre, Paris, 394 p.

MERLE D. 1990. - Revision des Muricidae du Cuisien de Gan et de Bos d'Arros (Bassin d'Aquitaine, France). Bulletin du Museum national d'Histoire naturelle, Paris 4 eme ser., 11 (1989), sect. C, 3: 145 - 185.

MORTON A. 2023. -- A Collection of Eocene and Oligocene Fossils. Available at: www. dmap. co. uk / fossils / (last access on november 2023).

NOLF D., STEURBAUT E. & CAHUZAC B. 2002. - Progres recents dans la connaissance des gisements cenozoiques en Aquitaine meridionale (Chalosse, Bearn et Bas-Adour: SW France. Revue de Micropaleontologie 45 (3): 164 - 194. https: // doi. org / 10.1016 / S 0035 - 1598 (02) 90017 - 6

VON KOENEN A. 1889. - Das Norddeutsche Unter-Oligocan und seine Mollusken-Fauna. Abhandlungen zur geologischen Specialkarte von Preussen und den Thuringischen Staaten 10 (1): 1 - 280.

Figures

FIG. 5. — Spiral cords in Trubatsa ganensis n. sp. (A-D), T. parisiensis (d’Orbigny, 1850) (E, F) and T. calviniacensis n. sp. (G, H): A, B, holotype, MNHN.F.A90520 (Varone coll.), Gan (Tuilerie), Pyrénées-Atlantiques, Aquitaine Basin, France, early Eocene (Ypresian); C, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); D, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); E, F, Alan Morton coll., Upper Barton Bed H, Barton-Highcliff, United Kingdom, middle Eocene (Bartonian); G, H, holotype, MNHN.F.A25590 (Pacaud coll.) Cauvigny (Châteaurouge), Oise, Paris Basin, France, middle Eocene (Lutetian). Scale bars: 5 mm. Credits: A-D, G, H, P. Loubry (MNHN/CNRS); E, F, A. Morton.

FIG. 25. — New species of Muricopsis Bucquoy & Dautzenberg, 1882, Favartia Jousseaume, 1880, Dermomurex Monterosato, 1890, Trubatsa Dall, 1889 and Timbellus De Gregorio, 1885: A, B, Muricopsis pontileviensis n. sp.: A, holotype, MNHN.F.A71120 (Dollfus coll.), Pontlevoy, France, Langhian; B, paratype, MNHN.F.A77743, (Dollfus coll.), Pontlevoy; C, D, M. neraudeaui n. sp.: C, holotype, MNHN.F.A77746 (Néraudeau sample), Saint-Jacques-de-la-Lande, France, Messinian;D, paratype,MNHN.F.A77747 (Néraudeau sample),Saint-Jacques-de-la-Lande;E, F, Favartia jansseni n. sp.:E, holotype,RGM.225979,Winterswijk,The Netherlands, Langhian; F, paratype,MNHN.F.A77748 (Ledon coll.),Winterswijk;G, H, Dermomurex (Gracilimurex) ligerianus n. sp.: G, holotype,MNHN.F.A33456 (Dollfus coll.), Pontlevoy; H, paratype, MNHN.F.A33457 (Dollfus coll.); I-K, Trubatsa ganensis n. sp.: I, holotype, MNHN.A90520 (Varone coll.), Gan (Tuilerie), France, Ypresian; J, paratype, MNHN.F.A90518 (Merle coll.), Gan (Tuilerie); K, paratype, MNHN.F.A90517 (Merle coll.), Gan (Tuilerie); L, T. calviniacensis n. sp., holotype,MNHN.F.A25590 (Pacaud coll.), Cauvigny (Châteaurouge),France, Lutetian; M-O, Timbellus magnificus n. sp.: M, holotype, MNHN.F.A90522 (Pacaud coll.), Baron (Sablière Heudebert), France, Bartonian; N, paratype, MNHN.F.A90525 (Schtrock coll.), Saint-Vaast-lès-Mello (Barisseuse), France, Bartonian; O, paratype, MNHN.F.A90523 (Pacaud coll.), Baron (Sablière Heudebert). Scale bars: A-L, 5 mm; M-O, 20 mm.