Ponderia remyi, Merle & Pacaud & Ledon & Goret, 2024

Merle, Didier, Pacaud, Jean-Michel, Ledon, Daniel & Goret, Bernard, 2024, New Cenozoic Muricidae (Mollusca: Gastropoda) from Europe, Geodiversitas 46 (15), pp. 495-551 : 528

Ponderia remyi n. sp.

( Figs 19 E-H; 27K, L)

urn:lsid:zoobank.org:act:13F57F4E-5263-4EC8-92FE-105E24BA99C9

TYPE MATERIAL. — Holotype. France • Normandy, Manche, Fresville; Calcaire de Fresville ; middle Eocene (uppermost Lutetian ); MNHN.F.A90548 (Remy coll.), H: 26 mm ( Figs 19E, F ; 27K ).

Paratypes. France • 5 spm; same as for the holotype; MNHN.F.A90549 (Remy coll.) • 1 spm; idem; MNHN.F.A90555 (Remy coll.) ( Fig. 19G, H ; 27L ) • 3 spm, MNHN.F.A90550 ( MNHN coll.) • 1 spm; Normandy, Manche , Fresville (coteau de Vauville); idem; MNHN.F.A27328 ( MNHN coll.) • 2 spm; idem; MNHN.F.B63590 (de Morgan coll.).

ETYMOLOGY. — Dedicated to Nicolas Remy who collected the material of this species.

TYPE HORIZON. — Calcaire de Fresville, middle Eocene (uppermost Lutetian), see Bignot et al. (1968).

TYPE LOCALITY. — France, Normandy, Manche, Fresville, see APGN (2024).

DISTRIBUTION. — Only known from the type locality.

DESCRIPTION

Bulbous, paucispiral protoconch of 1.5 whorls. Slender, fusiform teleoconch up to 21 mm in height, up to 9 mm in width. High spire, up to six subcarinate whorls. Last whorl (sixth whorl) up to 69% of total length. Apical angle 35° excluding P1 spine and up to 55° including P1 spine. Spiral sculpture with strongly marked primary cords. First and second whorl: no primary cord. Third whorl: appearance of weak P2. Fourth whorl: P2 strongly marked and appearance of weak P1. Fifth whorl: strongly marked P1 and P2. Sixth and last whorl: P1 to P3 well marked, P4 weak (convex part of whorl); secondary cords occasionally s1 and s2. Axial sculpture with 8-9 lamellose protovarices per whorl from first to fourth whorl to four lamellose protovarices. From fifth whorl to sixth whorl, three major varices and two or three nodulose intervarices intercalated. Varices with three major cord spines (P1 to P3). P1 spine appearing at beginning of first whorl and forming open tube at end of first whorl. From second to last whorl P1 spine well developed, open. Intervarical nodules spirally aligned and present on P1 and P2. Ovate aperture up 32% of diameter (including wings) and up to 76% of length of last whorl (including siphonal canal). Columellar lip smooth, poorly erect. Parietal lip adherent. Shoulder channel of P1 open. Outer lip lacking denticle. Outer lip not crenulate. Pseudoumbilicus narrow. Siphonal canal open, up to 55% of apertural length. COMPARISONS

Amongst European Eocene species, Ponderia remyi n. sp. can be compared to P. daguini ( Magne, 1941) from the Early Eocene of Gan (Aquitaine and Paris Basins) and to P. bispinosa (Sowerby, 1823) from the Middle Eocene of the Paris Basin and Hampshire ( England) and from the Late Eocene of Lattorf Basin ( Germany). Ponderia daguini is easily distinguished from P. remyi n. sp. in having a broader shape, more numerous cords (with P5, P6 and ADP) and nodulose cords. Ponderia bispinosa usually bears more foliaceus varices. The cord spines of P. bispinosa varies from two (P1 and P3) to five (P1 to P5). The morphotypes with five cords spines are common in the Lutetian and in the Bartonian of the Paris Basin, where those with two cord spines are common in the Bartonian of Hampshire and in the Priabonian of Lattorf ( von Koenen 1889; Merle et al. 2011). The varices of P. bispinosa are more winged than those of P. remyi n. sp. In addition, P. bispinosa clearly differs in having a multispiral protoconch and a microsculpture formed of small elongated granules (see Merle 1990).

Figures

FIG. 19. — Spiral cords and internal denticles of the outer lip in Textiliomurex chodmonensis n. sp. (A-D) and Ponderia remyi n. sp. (E, H): A, B, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), Oise, Paris Basin, France, middle Eocene (Lutetian); C, D, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-en-Vexin (Carrière Darcy); E, F, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Manche, Normandy, France, middle Eocene (Lutetian); G, H, MNHN.F.A90555, Fresville (Remy coll.). Scale bars: 10 mm. Credits: P. Loubry (MNHN/CNRS).

FIG. 27. — New species of Timbellus De Gregorio, 1885, Pterynotus Swainson, 1833 (Pterymarchia Houart, 1995), Textiliomurex Merle, 2011, Ponderia Houart, 1986 and Flexopteron Shuto, 1969: A-C, Timbellus submicropterus n. sp.: A, holotype, MNHN.F.A24111 (MNHN coll.), Fresville, France, Lutetian; B, paratype, MNHN.F.A24126 (Munier-Chalmas coll.), Gourbesville, France, Lutetian; C, paratype, UCBL-EM 33438 (Bourdot coll.), Fresville; D, T. cf. submicropterus n. sp., MNHN.F.A90557 (Brébion coll.), Saint-Aignan-Grand-Lieu (Pierre-Aiguë), France, Bartonian; E, F, Pterynotus (Pterymarchia) gaasensis n. sp.: E, holotype, MNHN.F.A32492 (Merle coll.), Gaas (Espibos), France, Rupelian; F, paratype, MNHN.F.A90554 (Merle coll.), Gaas (Espibos); G, H, P. (P.) pelouatensis n. sp.: G, holotype, MNHN.F.J09945, Saucats (Peloua), France, Burdigalian; H, paratype, MNHN.F.J17632, Saucats (Peloua); I, J, Textiliomurex chodmonensis n. sp.: I, holotype, MNHN.F.A81144 (Pacaud coll.), Chaumont-en-Vexin (Carrière Darcy), France, Lutetian; J, paratype, MNHN.F.A81145 (Pons coll.), Chaumont-en- Vexin (Carrière Darcy); K, L, Ponderia remyi n. sp.: K, holotype, MNHN.F.A90548 (Remy coll.), Fresville, Lutetian; L, paratype, MNHN.F.A90555, Fresville (Remy coll.); M, N, Flexopteron constantinense n. sp.: M, paratype, MNHN.F.A77761 (MNHN coll.), Carentan-les-Marais (Cauvin), France, Priabonian; N, holotype, MNHN.F.A77760 (MNHN coll.), Carentan-les-Marais (Cauvin). Scale bars: A-D, I-L, 10 mm; E-H, N, 5 mm; M, 3 mm. Credits: A, B, D-F, N, P. Loubry (MNHN/ CNRS); C, E. Robert (UCBL).