Dissomphalus, Ashmead, 1893
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publication ID |
https://doi.org/ 10.11646/zootaxa.5536.3.2 |
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publication LSID |
lsid:zoobank.org:pub:DADA0F2F-C442-4B2F-8C08-D918CFFB8709 |
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DOI |
https://doi.org/10.5281/zenodo.14248891 |
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persistent identifier |
https://treatment.plazi.org/id/038CCD37-FFB1-BA7D-3F8B-70FE79F2FDC5 |
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treatment provided by |
Plazi |
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scientific name |
Dissomphalus |
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Key to males of Dissomphalus from the punctatus species-group
1. Tergal process with tubercle ( Figs 1D, E View FIGURE 1 ).................................................................. 2
– Tergal process with tuft of setae ( Figs 1A, C, F View FIGURE 1 )............................................................. 3
2 (1). Abaxial margin of harpe projected anterad; basivolsella with spines ( Fig. 4H View FIGURE 4 )..................... D. johannae sp. nov.
– Abaxial margin of harpe projected anterad; basivolsella without spines ( Fig. 3G View FIGURE 3 ).................... D. deaneae sp. nov.
3 (1). Aedeagal dorsal lobe short ( Figs 2A View FIGURE 2 , 3A View FIGURE 3 , 5C View FIGURE 5 ).............................................................. 4
– Aedeagal dorsal lobe long or very long ( Figs 3C View FIGURE 3 , 4C View FIGURE 4 , 5E View FIGURE 5 ).................................................... 13
4 (3). Ventral margin of harpe bifurcated (see Colombo & Azevedo 2018).................. D. acutipupu Colombo & Azevedo
– Ventral margin of harpe not bifurcated ( Figs 5A, E View FIGURE 5 ).......................................................... 5
5 (4). Basivolsella with spines ( Figs 4H View FIGURE 4 , 5G View FIGURE 5 ).................................................................... 6
– Basivolsella without spines............................................................................. 7
6 (5). Harpe entirely wide; inner median filament almost same size of outer lobe directed dorsad; basal cover plate in diamond shape ( Fig. 5D View FIGURE 5 )......................................................................... D. margarethae sp. nov.
– Harpe wide basally; inner median filament anterior to apex of aedeagal outer lobe posterad; basal cover plate in inverted triangle shape ( Fig. 3F View FIGURE 3 )................................................................ D. dandarae sp. nov.
7 (5). Aedeagal dorsal outer lobe directed mesad ( Fig. 4A View FIGURE 4 )......................................................... 8
– Aedeagal dorsal outer lobe not directed mesad............................................................. 11
8 (7). Inner median filament aligned with apex of aedeagal dorsal lobes; basal cover plate very wide; aedeagal ventral valve higher than aedeagal dorsal valve ( Fig. 4A View FIGURE 4 )......................................................... D. elzae sp. nov.
– Inner median filament anterior to apex of aedeagal dorsal lobes ( Fig. 3F View FIGURE 3 ); basal cover plate narrow ( Figs 2F View FIGURE 2 , 3A View FIGURE 3 ); aedeagal ventral valve not higher than aedeagal dorsal valve ( Figs 3D View FIGURE 3 , 5A, B View FIGURE 5 )............................................ 9
9 (8). Harpe entirely wide with dorsal margin straight ( Figs 5D View FIGURE 5 , 6F View FIGURE 6 ); apex of aedeagal ventral valve parallel ( Fig. 5C View FIGURE 5 )......... 10
– Harpe wide basally with posterior margin very incurved; apex of aedeagal ventral valve slightly laterad ( Fig. 3A View FIGURE 3 )........................................................................................... D. ashauerae sp. nov.
10 (9). Digitus very long; posterior margin of basal cover plate broadly incurved; apex of harpe wide ( Fig. 1B View FIGURE 1 )...... D. napo Evans
– Digitus short; posterior margin of basal cover plate completely incurved; apex of harpe narrow ( Fig. 5C View FIGURE 5 ) .................................................................................................. D. lispectorae sp. nov.
11 (7). Basal cover plate very narrow with posterior margin completely incurved ( Fig. 2F View FIGURE 2 ); harpe entirely wide; inner median filament directed posterad ( Figs 3G View FIGURE 3 , 5B View FIGURE 5 , 6E View FIGURE 6 )...................................................................... 12
– Basal cover plate wide with posterior margin broadly incurved; harpe wide basally; inner median filament directed ventrad ( Fig. 7A View FIGURE 7 )............................................................................. D. sueliae sp. nov.
12 (11). Apex of harpe wide; abaxial margin of harpe not projected anterad; basivolsella and digitus sclerotized ( Fig. 3H View FIGURE 3 )................................................................................................ D. elisae sp. nov.
– Apex of harpe narrow; abaxial margin of harpe projected anterad; basivolsella and digitus not sclerotized ( Fig. 2F View FIGURE 2 )............................................................................................. D. ameliae sp. nov.
13 (3). Aedeagal dorsal lobe very long ( Figs 1C View FIGURE 1 , 5H View FIGURE 5 )............................................................. 14
– Aedeagal dorsal lobe long............................................................................. 15
14 (13). Basivolsella elongated with well delimited projection ( Fig. 1C View FIGURE 1 )............................... D. punctatus (Kieffer)
– Basivolsella not elongated without such projection ( Fig. 5H View FIGURE 5 )..................................... D. martae sp. nov.
15 (13). Basal part of basivolsella united to aedeagal ventral valve.................................................... 16 Basal part of basivolsella not united to aedeagal ventral valve................................................. 17
16 (14). Margin of aedeagal outer lobe sinuous; ventral margin of outer lobe with pointed projection upward ( Fig. 6G View FIGURE 6 )................................................................................................ D. silveirae sp. nov.
– Margin of aedeagal outer lobe rounded; ventral margin of outer lobe without pointed projection upward (see Colombo & Azevedo 2018)................................................................ D. cuca Colombo & Azevedo
17 (15). Harpe with dorsal margin slightly narrower in anterior part (see Redighieri & Azevedo 2006).................................................................................................. D. bahiensis Redighieri & Azevedo
– Harpe with dorsal margin not slightly narrower in posterior part............................................... 18
18 (17). Harpe color darker than other parts of genitalia ( Fig. 2E View FIGURE 2 )....................................... D. alzirae sp. nov.
– Harpe color not darker than other parts of genitalia.......................................................... 19
19 (18). Harpe very wide with dorsal margin entirely straight; basal cover plate in inverted triangle shaped ( Figs 5D View FIGURE 5 , 6F View FIGURE 6 )........ 20
– Harpe not very wide with dorsal margin strongly or slightly incurved dorsally; basal cover plate not inverted triangle shaped. .................................................................................................. 22
20 (19). Inner median filament anterior to aedeagal dorsal lobes ( Figs 5E View FIGURE 5 , 6E View FIGURE 6 , 7B View FIGURE 7 ); digitus narrow ( Figs 2F View FIGURE 2 , 4H View FIGURE 4 , 6H View FIGURE 6 ); posterior margin of hypopygium straight or nearly so ...................................................................... 21
– Inner median filament aligned with aedeagal outer lobes; digitus wide; posterior margin of hypopygium weakly outcurved ( Fig. 5B View FIGURE 5 )................................................................................... D. limae sp. nov.
21 (20). Digitus very long and narrow; basivolsella with narrow protuberance ( Fig. 6F View FIGURE 6 )........................ D. ritae sp. nov.
– Digitus short and wide; basivolsella with wide protuberance ( Fig. 5D View FIGURE 5 ).............................. D. luisae sp. nov.
22 (19). Aedeagal dorsal lobes wide ( Figs 3D View FIGURE 3 , 5F View FIGURE 5 , 6D View FIGURE 6 ) .............................................................. 23
– Aedeagal outer lobes narrow........................................................................... 25
23 (22). Harpe with dorsal margin deeply incurved posteriorly with apex narrow........................................ 24
– Harpe with dorsal margin not deeply incurved posteriorly with apex wide ( Fig. 3D View FIGURE 3 )................ D. carmenae sp. nov.
24 (23). Margin of aedeagal outer lobes rounded; basal cover plate with ventrolateral corner very narrow; apex of aedeagal ventral valve parallel ( Fig. 6D View FIGURE 6 )........................................................................ D. nisiae sp. nov.
– Margin of aedeagal outer lobes almost straight with pointed protuberance posterad; basal cover plate entirely wide; apex of aedeagal ventral valve curved laterad ( Fig. 5F View FIGURE 5 )................................................ D. lutzae sp. nov.
25 (22). Inner median filament curved laterad.................................................................... 26
– Inner median filament curved posterad, dorsad or ventrad.................................................... 27
26 (25). Posterior margin of basal cover plate incurved medially; basivolsellar protuberance poorly delimited; posterior margin of harpe outcurved ( Fig. 7B View FIGURE 7 )................................................................... D. vivianeae sp. nov.
– Posterior margin of basal cover plate not incurved dorsally; basivolsellar protuberance well delimited; posterior margin of harpe slightly incurved ( Fig. 2H View FIGURE 2 )............................................................. D. antonietae sp. nov.
27 (25). Gonostipe is about two times shorter than harpe ( Fig. 5E View FIGURE 5 )....................................... D. lumae sp. nov.
– Gonostipe as long as harpe or nearly so................................................................... 28
28 (27). Inner median filament posterior to aedeagal dorsal lobes; apex of harpe very narrow with abaxial margin projected anterad (see Colombo & Azevedo 2016).................................................. D. tupinikum Colombo & Azevedo
– Inner median filament aligned or anterior to apex of aedeagal outer lobes; apex of harpe wide or very wide............. 29
29 (28). Inner median filament aligned with apex of aedeagal lobes................................................... 30
– Inner median filament anterior to apex of aedeagal lobes..................................................... 31
30 (29). Aedeagal dorsal valve in bottle shaped; basal cover plate narrow with posterior margin straight; basivolsellar protuberance well delimited ( Fig. 2A View FIGURE 2 ).................................................................. D. montanus (Kieffer)
– Aedeagal dorsal margin almost in bottle or nearly so; basal cover plate wide with posterior margin slightly incurved; basivolsellar protuberance poorly delimited ( Fig. 3C View FIGURE 3 )...................................................... D. boriae sp. nov.
31 (29). Basal cover plate very wide medially in inverted triangle shape............................................... 32
– Basal cover plate is completely narrow, not inverted triangle shape............................................. 33
32 (31). Inner median filament directed posterad; basal cover plate with sclerotized oblique structure; apodeme far beyond genital ring ( Fig. 6H View FIGURE 6 )............................................................................. D. soniae sp. nov.
– Inner median filament directed dorsad; basal cover plate without such structure; apodeme not far beyond genital ring ( Fig. 2G View FIGURE 2 )....................................................................................... D. annae sp. nov.
33 (31). Apodeme far beyond genital ring; dorsal margin of harpe entirely narrow........................................ 34
– Apodeme not far beyond genital ring; dorsal margin of harpe not entirely narrow.................................. 37
34 (33). Median region of aedeagal dorsal valve slightly incurved medially; posterior margin of basal cover plate incurved medially ( Fig. 5A View FIGURE 5 ).............................................................................. D. leliae sp. nov.
– Median region of aedeagal dorsal valve outcurved; posterior margin of basal cover plate not incurved medially.......... 35
35 (34).Apex of aedeagal dorsal valve not abruptly narrow; basal cover plate entirely narrow with posterior margin straight; basivolsellar protuberance well demarcated and wide; ventral margin of harpe with small projection ( Fig. 6C View FIGURE 6 )...... D. mirandae sp. nov.
– Apex of aedeagal dorsal valve abruptly narrow; basal cover plate entirely narrow with posterior margin broadly rounded and narrow in dorsolateral margin; basivolsella without protuberance ( Fig. 4F View FIGURE 4 )....................... D. guidonae sp. nov.
37 (33). Basivolsella without protuberance....................................................................... 38
– Basivolsella with protuberance......................................................................... 40
38 (37). Apex of aedeagal outer lobes directed laterad; dorsal margin of harpe slightly incurved posteriorly; aedeagal ventral valve parallel ( Fig. 3B View FIGURE 3 )..................................................................... D. beatrizae sp. nov.
– Apex of aedeagal outer lobes directed posterad; dorsal margin of harpe strongly incurved posteriorly; aedeagal ventral valve angled laterad....................................................................................... 39
39 (38). Tergal process with dense tuft of setae; dorsal margin of harpe strongly incurved posteriorly; basal cover plate entirely narrow ( Fig. 2D View FIGURE 2 )......................................................................... D. alexandrinae sp. nov.
– Tergal process without dense tuft of setae; dorsal margin of harpe not strongly incurved posteriorly; basal cover plate very narrow in dorsolateral corner ( Fig. 4D View FIGURE 4 )................................................... D. evaristoae sp. nov.
40 (37). Basivolsellar protuberance narrow...................................................................... 41
– Basivolsellar protuberance wide........................................................................ 42
41 (40). Aedeagal dorsal valve robust; aedeagal ventral valve posterior to aedeagal dorsal valve; dorsal margin of harpe strongly incurved posteriorly ( Fig. 6A View FIGURE 6 )............................................................. D. maryae sp. nov.
– Aedeagal dorsal valve narrow; aedeagal ventral valve aligned to aedeagal dorsal valve; dorsal margin of harpe not strongly posteriorly ( Fig. 4C View FIGURE 4 ).................................................................... D. eulaliae sp. nov.
42 (40). Basal cover plate narrow; apex of harpe truncate and very wide................................................ 43 – Basal cover plate wide; apex of harpe sinuous and not very wide ............................................... 44 43 (42). Basidorsal corner of digitus strongly projected; posterior hypopygeal margin straight or nearly so; aedeagal dorsal valve narrow
( Fig. 4G View FIGURE 4 )......................................................................... D. jaquelineae sp. nov. – Basidorsal corner of digitus poorly projected; posterior hypopygeal margin sinuous; aedeagal dorsal valve in bottle shape or nearly so ( Fig. 6E View FIGURE 6 )...................................................................... D. penhae sp. nov. 44 (42). Posterior margin of basal cover plate incurved medially; basivolsellar protuberance poorly delimited ( Fig. 4E View FIGURE 4 )............
................................................................................... D. grazielae sp. nov. – Posterior margin of basal cover plate broadly incurved; basivolsellar protuberance well projected..................... 45 45 (44). Dorsal margin of harpe wide; basidorsal corner of digitus poorly projected ( Fig. 3E View FIGURE 3 )............... D. carolinae sp. nov.
Dorsal margin of harpe narrow; basidorsal corner of digitus strongly projected.................................... 46 46 (45). Tergal process with dense tuft of setae; inner median filament directed ventrad; ventral margin of harpe without projection ( Fig. View FIGURE 6
6B)............................................................................... D. meirelesae sp. nov. – Tergal process with tuft of little setae; inner median filament directed posterad; ventral margin of harpe with small projection
( Fig. 4B View FIGURE 4 )............................................................................ D. enedinae sp. nov.
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