Gymnetis chalcipes Gory & Percheron, 1833

Iorio, Osvaldo Di, 2013, A review of the Cetoniinae (Coleoptera: Scarabaeidae) from Argentina and adjacent countries: systematics and geographic distributions, Zootaxa 3668 (1), pp. 1-87 : 24-26

Gymnetis chalcipes Gory & Percheron, 1833

Figs. 44–45 (aspects), 52 (parameres), 56–57 (anatomy), 62 (distribution)

[ Gymnetis] Chalcipes. Dej.: Dejean 1833: 169 (catalog; distribution; nomen nudum).

Gymnetis chalcipes Gory & Percheron, 1833: 354 (description; distribution), plate 71, Fig. 2 ); Blanchard 1850: 35 (catalog; distribution); Bruch 1911a: 214 (catalog; distribution); Schenkling 1921: 92 (catalog; distribution; references); Krajčík 1998: 76 (catalog; distribution).

Paragymnetis chalcipes G. & P. subsp. chalcipes G. & P. : Schürhoff 1937: 67 (new combination; distribution); Blackwelder 1944: 263 (catalog; distribution).

Gymnetina chalcipes (G. & P.) : Korytkowski & Ojeda 1970: 77 (catalog; distribution).

Gymnetina (Gymnetoides) chalcipes chalcipes (G. & P., 1833): Monné 1970: 61 (references).

Paragymnetis chalcipes ( Gory & Percheron, 1833) : Silva et al. 1968: 555 (distribution; natural history); Morelli 2000: 155–165 (distribution; larva); Navarrete-Heredia 2001: 427 (distribution; natural history; references).

Paragymnetis litigiosa , not Gory & Percheron, 1833: Di Iorio 2003: 179 (distribution; natural history): misidentification.

Distribution: PERU: Amazonas: Bagua, [MELP], identified by Pereyra 1970 ( Korytkowski & Ojeda 1970). BRAZIL ( Dejean 1833, Gory & Percheron 1833, Schenkling 1921): Río Grande do Sul: [locality not stated] ( Silva et al. 1968). ARGENTINA ( Schenkling 1921, Krajčík 1998): Corrientes ( Blanchard 1850); Buenos Aires: La Plata, in Bruch Collection ( Bruch 1911a); [Pdo. Berisso], Los Talas, near La Plata ( Bruch 1919); Punta Lara, 20-II-[1927], Maldonado Bruzzone leg., 4 exx., identified by Bruch ( Maldonado Bruzzone 1927); Buenos Aires ( Bruch 1929); ( Schenkling 1921, Blackwelder 1944); Florencio Varela ( Di Iorio 2003). URUGUAY: [locality not stated] ( Monné 1970); Lavalleja: Route 8, km 120, near “La Fuente Salus” ( Morelli 2000); Canelones: Sauce ( Morelli 2000).

Material examined: PERU: Loreto: Río Marañón, Montenegro , 6-XI-1961, Williner leg., 1 ex. [ ODIC], “ Gymnetina (Gym.) / chalcipes / (G. et P.) / A. Martínez det. 1965” . BRAZIL: Rio de Janeiro: [Rio de Janeiro?], G. Pellerano Collection, 1 ex. [ MACN] ; Paraná: Carambei, 15-II-2004, 4 exx. [ LABA] ; PARAGUAY: “Para- / guay” [printed on green paper], 1 ex. [ HBBA], without labels, 3 exx. [ HBBA] ; ARGENTINA: Misiones: Col. Berg, 1 ex. [ MLPA] ; El Soberbio, 30-XI-1969, J. Carreras leg., 1 ex. [ JCDT] ; Córdoba: Río Primero , 14-II-1953, A. Martínez leg., 2 exx. [ ODIC] ; Entre Ríos: 190_, C. Bruch, 1 ex. [ CBBA] ; Gualeguaychú, [M. Doello Jurado leg.], # 12322, 1 ex. [ MACN] ; Concordia, 13-II-2002, Zubarán leg., 1 ex. [ GZSM] ; Federación, G. Zubarán leg., 4- XII-2000, 1 ex. [ GZSM], 12-XII-2000, 1 ex. [ GZSM], 15-I-2003, 1 ex. [ GZSM] ; Dto. Colón, Ruinas del Viejo Molino , 20–25-XI-2009, 1 ex. [ ZIMP] ; Ibicuy, I-2012, C. Ufor leg., 4 exx. [ CUJCP], 2 exx. [ ODIC] ; La Pampa: Catriló, I-1980, 1 ex. [ ODIC] ; Buenos Aires: “Pcia Buenos Aires / 4 189 7 / C. Bruch ”, 1 ex. [ CBBA], “ Gymnetis / chalcipes / G.P.” [handwritten by Bruch] ; same label, except 1-II-1904, 2 exx. [ CBBA] ; Buenos Aires, 17-XII-1904, C. Bruch, 1 ex. [ MLPA], “ Gymnetis / chalcipes ” [handwritten by Bruch] ; Buenos Aires, 12-IX-1917, A. Zotta leg., 1 ex. [ MACN], “ Gymnetis / litigiosa / G. & P.” ; Buenos Aires, 12. 939, G. Pellerano Collection, 1 ex. [ MACN] ; Buenos Aires, Bosq Collection, 2 exx. [ MLPA], Richter Collection, 4 exx. [ MLPA] ; [Buenos Aires, 12- III-1926, E.A. Campos leg.], # 17526, 1 ex. [ MACN] ; San Nicolás, XII-1982, Bianchini leg., 1 ex. [ ARCU] ; Luján, XII-1985, [A. Oliva leg.], 1 ex. [ MACN] ; Delta, Isla de los Laureles, 21-XI-1939, Bosq Collection, 1 ex. [ MLPA] ; Delta, M.J. Viana, VII-1947, 1 ex. [ MLPA], VII-1951, 1 ex. [ MLPA] ; Isla Martín García, IV-1937, M.J. Viana leg., 1 ex. [ MLPA] ; Delta [Río] Paraná, in front of Campana, V-1968, E. Ogueta, 1 ex. [ ODIC] ; Delta del Paraná , Estación Inta, 12-I-1972, 1 ex. [ ODIC], 8-II-1972, 1 ex. [ ODIC] ; Río Luján , F.C.G.B.M., 30-XII-2011 (pupae), Di Iorio leg., 10 exx. [ ODIC] ; Escobar, II-2005, C. Ufor leg., 2 exx. [ CUJCP] ; Otamendi, 30-VII-2008, Di Iorio leg., 1 ex. [ ODIC] ; Merlo, Barrio San Martín, 11-IV-1995, Arias leg., 1 ex. [ JCDT] ; Castelar, VII-1980, 1 ex. [ ODIC] ; [Pdo. Moreno], [Pdo. Moreno], La Reja, 28-III-1971, A. del Valle leg., 1 ex. [ ARCU] ; Morón, F. Penco leg., without data, 3 exx. [ FPBA], 29-XI-2001, 1 ex. [ FPBA], 16-IX-2002, 1 ex. [ FPBA], 25-III-2004, 1 ex. [ FPBA], 15-XII-2006, 1 ex. [ JCDT] ; Morón, Barrio Alessandro , 21-I-2009, V. Osses leg., 1 ex. [ JCDT] ; Ituzaingó, Parque Leloir, XI-2001, 1 ex. [ FPBA] ; José C. Paz, II-1996, C. Servian leg., 1 ex. [ FPBA], XII-2005, Borquez leg., 3 exx. [ LABA] ; San Miguel, V-1971, Williner leg., 1 ex. [ MACN] ; Bella Vista, XII-1953, J.M. Gallardo leg., 1 ex. [ MACN], “ Paragymnetis / chalcipes / (G. & P.) / det. B.C. Ratcliffe 2003”, II-1969, Bachmann leg., 1 ex. [ ODIC], 12-IV-1970, V. Lasaignes leg., 1 ex. [ ARCU] ; Hurlingham, 22-I-1997, Di Iorio leg., 1 ex. [ ODIC] ; Grand Bourg, XII-1983, Di Iorio leg., 1 ex. [ ODIC] ; Los Polvorines, 12-I-2012 (larvae), Di Iorio leg., 4 exx. [ ODIC] ; Campo de Mayo , IV-1966, 1 ex. [ ODIC], Di Iorio leg., 24-XII-2011 (pupae), 47 exx., 15-II-2012 (larvae), 7 exx. [ ODIC] ; Don Torcuato, 15-IV-2012 (larvae), Di Iorio leg., 1 ex. [ ODIC] ; Tigre, 1-I-1903, [illegible letter], Carbouli leg., 1 ex. [ MACN], “ Gymnetis / litigosa / G. & P.”, II-1923, 1 ex. [ MACN], IV-1936, M.J. Viana leg., 1 ex. [ MLPA], VI-1944, M.J. Viana leg., 1 ex. [ MLPA] ; Las Conchas, Col. Frers, 1 ex. [ MACN], “ Gymnetis / litigiosa / Gory et Perch.” [handwritten by Frers] ; San Fernando, 16-X-2007, L. Aguado leg., 2 exx. [ LABA] ; San Isidro, Col. Frers, # 461, 1 ex. [ MACN], 13-XI-1969, 1 ex. [ ABJCP] ; San Isidro (Puerto), 5 exx. [ HAHC], I-1973, A. Martínez leg., 1 ex. [ ODIC] ; Martínez, 4-II [or XI?]-1926, Bridarolli leg., 1 ex. [ MACN], “ Gymnetis / inquinita / Thoms. / Bruch det.” [printed], 9-VIII-1981, de Lavallaz leg., 1 ex. [ ARCU] ; La Lucila, 23-XI-1969, Guarrera leg., 1 ex. [ MLPA] ; Vicente López, G. Pellerano Collection, II-1931, 1 ex. [ MACN], I-1936, 1 ex. [ MACN], XII-1937, 1 ex. [ MACN], I-1940, 1 ex. [ MACN] ; Reserva Ecológica Vicente López, 12-II-2012, L. Damer leg., 1 ex. [ LDBA], 3-I-2013, Di Iorio leg., 2 exx. [ ODIC] ; Olivos, 1-XII-1918, 1 ex. [ MLPA], “ Gymnetis / chalcipes / (G. et P.)”, II-1988, C. Fernández leg., 1 ex. [ ARCU] ; Munro, 20-III-2012 (larvae), Di Iorio leg., 42 exx. [ ODIC] ; Gonzalez Catán, 21-IX-1970, Castronuovo leg., 1 ex. [ ARCU] ; Quilmes, 17-XI-1983, R. Mauri leg., 1 ex. [ MLPA] ; Adrogué, 18-X-1991, A. Cicchino leg., 1 ex. [ MLPA] ; La Plata, 22-III-1991, M. Lopez Ruff leg., 1 ex. [ MLPA] ; Punta Lara, VI-1949, M.J. Viana leg., 1 ex. [ MLPA], 13-IV-1969, A. Torno leg., 1 ex. [ ARCU], I-1977, 1 ex. [ ODIC] ; Florencio Varela, 19-III-1996, Di Iorio leg., 2 exx. [ ODIC] ; Capital Federal: Devoto, IV-1981, 1 ex. [ ODIC] ; Capital Federal, [Villa Pueyrredón], A. Fortino leg., 8-I-1996, 1 ex., 3-I-1997, 1 ex., 22-I-1997, 1 ex., II-1999, 1 ex., II-2004, 1 ex. [all AFBA] ; Palermo, 25-XI-1978, Kerner leg., 1 ex. [ ARCU], Di Iorio leg., 18-XI-2011 (pupae), 23 exx. [ ODIC], 19- XII-2011 (P), 3 exx. [ ODIC], 26-IV-2012 (larvae), 2 exx. [ ODIC], 26-IX-12 (larvae), 2 exx. [ ODIC] ; Villa Crespo, Parque Centenario, 13-IX-2012 (larva), Di Iorio leg., 1 ex. [ ODIC] ; Ciudad Universitaria, 12-XII-1975, Cignoni leg., 1 ex. [ ARCU], I-1981, Di Iorio leg., 1 ex. [ ODIC], II-1986, M. Kun leg., 2 exx. [ ARCU], XI-2010, H. Iuri leg., 2 exx. [ ODIC], 17-II-2012 (L), Di Iorio leg., 2 exx. [ ODIC], 2011–2012, 175 exx. [ ODIC], XII-2012, 1 ex. [ ODIC] ; Villa Pueyrredon, Parque General Paz, 22-II-2012 (larvae), Di Iorio leg., 26 exx. [ ODIC] ; Capital Federal, III-2004, 1 ex. [ EAPW] ; Belgrano, 26-X-2012, P. Torres leg., 1 ex. [ ODIC] ; Agronomía, 6-I-2012, A. Fortino leg., 1 ex. [ AFBA] ; Reserva Ecológica Costanera Sur, 9-XI-2011, L. Damer leg., 1 ex. [ LDBA] ; URUGUAY: “Banda / Oriental”, Col. Berg, 1 ex. [ MLPA] ; Cerro Largo: Cañada de los Burros, Tremoleras Collection, 1 ex. [ MLPA] ; Fraile Muerto, 23-III-1932, Tremoleras Collection, 1 ex. [ MLPA] ; “ Po. Tía Lucía ”, Tremoleras Collection, 1 ex. [ MLPA] ; Cuchillo de Melo , 20-XII-1915, Tremoleras Collection , 1 ex. [ MLPA] ; Colonia: Col. A. Stevenin, 1 ex. [ MACN], “ Gymnetis / chalcipes / G & P” ; San José: “San José / B a de Cufré”, 1 ex. [ MLPA], “ Gymnetis / chalcipes / Gory Perch.” ; Montevideo: Montevideo , Felippone leg., 1 ex. [ MACN], “ Gymnetis / irregularis / G. et P.” ; Maldonado: A. Stevenin Collection, 2 exx. [ MACN].

Male genitalia: parameres ( Figs. 52 ); armature of the internal sac ( Figs. 56–57 ).

Note: Gymnetis chalcipes may be distinguished by the cupreous shine on the lateral and anterior margin of the clypeus, tarsi, tibiae, mesosternal process, and abdominal sternites; the same reflection is brown in G. undata , and black in G. litigiosa .

Figures

FIGURE 2. Geographic distributions of Gymnetini from Argentina and adjacent countries: Gymnetis bonplandi (triangles); Gymnetis hepatica (circles); Hologymnetis sp. (square).

FIGURES 44–51. General aspects of Gymnetini from Argentina: 44–45, Gymnetis chalcipes (44, common phenotype; 45, light phenotype); 46, Gymnetis litigiosa (La Rioja: Los Molinos); 47–48, Gymnetis hepatica (47, paratype, Santiago del Estero: Sachayoj; 48, paratype, Chaco: Charata); 49, Gymnetis undata (Misiones, Bruch Collection); 50, Gymnetis hebraica; 51, M. insculpta.

FIGURES 52–55. Male parameres: 52, Gymnetis chalcipes (Buenos Aires: Florencio Varela); 53, Gymnetis litigiosa (La Rioja: Los Molinos); 54, Gymnetis hepatica (paratype, Santiago del Estero: Sachayoj); 55, Gymnetis undata.

FIGURES 56–61. Armature of the internal sac and details of spines: 56–57, Gymnetis chalcipes; 58–59, Gymnetis litigiosa (La Rioja: Los Molinos); 60–61, Gymnetis pudibunda.