Panopides anticus Pascoe, 1871

Setliff, Gregory P., 2008, Revision of the Genus Panopides Pascoe (Coleoptera: Curculionidae: Cryptorhynchinae), The Coleopterists Bulletin 62 (1), pp. 83-98 : 83-98

Panopides anticus Pascoe

( Figs. 1–5 ; 12 ; 15–25 )

Panopides anticus Pasoce 1871:199 [key], 200 [description], pl. VIII: fig. 4 [figured].

Panopides antica Pascoe , [unjustified emendation in Hustache (1936: 260)].

Diagnosis. This species is readily distinguished from its congeners by its ornate maculation on the elytra and pronotum. Other than the vestiture and slightly thinner legs, it is externally very similar to P. philippinicus from which it can be further separated by the poorly developed apical arms of the male spiculum gastrale and thicker basal endophallic sclerites. It differs from P. riedeli by its large size (8.5–8.9 mm compared to 6.2–6.4 mm), longer, more gracile legs that are curved and have a setal brush in males, more separated meso- and metacoxae, and comparatively weakly developed elytral prominence.

88 Redescription. Length 8.5–8.9 mm, width 3.5–4.0 mm (n 5 4). Vestiture of elytra, pronotum, tibia, and abdominal ventrites brown (except intercoxal process of ventrite 1). Scutellum, pleural sclerites, prosternal canal, meso- and metasternum, intercoxal process of ventrite 1, dorsal portions of femora proximal to body, ventral portions of femora, and dorsum of tarsi densely clothed in pale white to yellowish-white scales. Anterior region of pronotal disk in front of cervical constriction, vertex of head, and base of rostrum also clothed in pale yellowish-white scales; pronotal base with oblong macula medially and pair of indistinct dorsolateral patches of pale scales. Elytra with humeri to anterolateral pleural margin with sparse, pale scales and large, pale, sutural maculation at apex past declivity. Each elytron with oblique macula of pale scales from suture near scutellum reaching near midline at intervals 5 or 6, pair of oblique macula forming chevron with vertex at scutellum when elytra closed. Elongate setae originating from granules on dorsum and legs mostly dirty white to pale yellowish-brown. Head. Antennae slender, funicular articles 1 + 2 longer than

8) female lateral view; 9) same, dorsal view.

antennal club; club ovate. Thorax. Pronotum as broad as long; short medial carina visible at anterior margin. Scutellum elongate-elliptical, much broader than long. Elytral punctures very large and deep, interspace between punctures smaller than diameter of puncture. Intervals moderately convex, interval 3 with longitudinal prominence starting near base and terminating at elytral declivity; prominence elevated above other intervals, narrow, not obscuring adjacent striae, with rounded apex and two rows of closely appressed, shiny granules. Meso- and metacoxae separated by about the length of mesocoxa. Male protibia strongly to moderately curved with well developed setal brush on ventral margin comprised of paired rows of very long, brown, hair-like setae ( Fig. 12 ). Hind femur exceeding elytral apex by one-third its length, mesofemora easily exceeding elytral apices. Abdomen. Posterior margin of male tergite VII truncate, broadly emarginate medially, with 10–11 pairs of plectral tubercles arranged in 2 longitudinal rows on either side of midline ( Fig. 19 ). Female tergite VII with posterior margin rounded, plectral rows not reaching the anterior or posterior margins ( Fig. 24 ). Male terminalia. Tergite VIII subquadrate, longer than broad, 90 posterior margin rounded; sternite VIII obsolescent ( Fig. 15 ). Spiculum gastrale with apodeme shorter than aedeagal apodemes; apical arms weakly developed, more or less symmetrical ( Fig. 18 ). Tegminal apodeme longer than width of tegminal ring ( Fig. 20 ). Aedeagus with apodemes more than twice as long as aedeagal body, body weakly curved, apex rounded. Endophallus with inverted Yshaped apical sclerite and pair of elongate basal sclerites; basal sclerites subequal in length to aedeagal body, basal extremities exposed with apices straight. Paired middle sclerites near base of aedeagal body ( Figs. 16–17 ). Female terminalia. Hemisternites subcylindrical, 3.6 times longer than styli, narrow, slightly broadened at base; styli 4.3 times long as broad, cylindrical, with minute apical setae as in Fig. 21 . Tergite VIII twice as long as broad, apex slightly broader than subapical region ( Fig. 23 ). Sternite VIII with apical plate spatulate, tapering at base, plate as long as apodeme ( Fig. 25 ).

Remarks. Two female specimens labeled with British Museum syntype disk labels were examined. The label data for both specimens are identical except on one specimen the determination label has the word ‘‘type’’ written in Pascoe’s small handwriting. This specimen is selected here as the lectotype. A lectotype designation is required to ensure the stability of nomenclature in case additional type material is discovered. One male specimen from Manado, N. Sulawesi, was discovered in the Pascoe special collection and studied at the NHML but was not borrowed for critical examination.

Distribution. Northern Sulawesi

92 Material examined. Lectotype (designated here): ♀, ‘‘SYN-TYPE [white disk with blue margin]/ Tondano [blue disk]/ Panopides anticus Pasc Type / Pascoe Coll. B.M. 1893–60’’ ( NHML) ; Paralectotype: 1 ♀ ‘‘SYN-TYPE? [white disk with blue margin; question mark handwritten]/ Tondano [blue disk]/ Panopides anticus Pasc / Pascoe Coll. B.M. 1893-60’’ ( NHML) ; Other material examined: 1 Oi, ‘‘ Minardo [probably Manado, N. Sulawesi]/ Bowring 63 47*’’ [on underside] ( NHML) ; 1 Oi, ‘‘ Minado [5 Manado; blue disk]’’ ( NHML) ; 1 ♀, ‘‘ Totok N. Celebes /‘‘ E. Gowing Scopes Collection, NHML (E) 2005–4’’ [on underside] ( NHML) .

Figures

Fig. 1. Detail of female head and rostrum of Panopides anticus, frontal view.

Figs. 2–5. Habitus of Panopides anticus. 2) male, lateral view; 3) same, dorsal view; 4) female, lateral view; 5) same, dorsal view.

Figs. 12–14. Right male prothoracic leg of Panopides species, anterior view. 12) P. anticus; 13) P. philippinicus; 14) P. riedeli. Scale line 5 1.0 mm.

Figs. 15–20. Male terminalia of Panopides anticus. 15) tergite VIII and sternite VIII, ventral view; 16) aedeagus, lateral view, tegmen removed; 17) aedeagus, ventral view, tegmen removed; 18) spiculum gastrale, ventral view; 19) tergite 7, dorsal view; 20) tegmen, ventral view. Scale line 5 1.0 mm.

Figs. 21–25. Female terminalia of Panopides anticus. 21) hemisternites, dorsal view; 22) spermatheca, lateral view; 23) tergite VIII, dorsal view; 24) tergite VII, dorsal view; 25) sternite VIII, ventral view. Left scale line for Figs. 21–23 5 0.5 mm, and right scale line for Figs. 24–25 5 1.0 mm.