Chlaenius (Pseudochlaeniellus) sobrinus Dejean, 1826

Anichtchenko, Alexander & Kirschenhofer, Erich, 2017, To the knowledge of Oriental species of subgenus Pseudochlaeniellus Jeannel, 1949 (Coleoptera, Carabidae, Chlaenius), Zootaxa 4231 (2), pp. 187-202 : 190-192

Chlaenius (Pseudochlaeniellus) sobrinus Dejean, 1826

( Figs. 7–12 , 34–36 )

Chlaenius sobrinus Dejean, 1826: 316

Chlaenius callichloris Bates, 1873: 250 syn. n.

C. (Pseudochlaeniellus) ladon Kirschenhofer, 1998: 322 syn. n.

Material. CHINA: 1 ♀ —„China, Sechuan, Wudu, 2.6.1997, leg. E. Kučera “, “ Chlaeniellus caesitus Andr. , det. Kirschenhofer ” (cMH); 4 ex—“ CH: Yunnan, Hekou: Hekou T. Naxihe R. n. Shanyao St. 88 m, 22°31'05.3"N 104°00'19.7"E , 18.XII.2003 B.Kataev leg.” ( MPU). INDIA: 13 ♂, 11 ♀ — “N India, Uttarakhand, 23– 26.04.2012, 5 km N Ramnagar, valley of Kosi riv., N 29°25'56.50", E 79°8'21.40", Anichtchenko A. leg.” (cAA; DUBC); 8 ♂, 10 ♀ — “N India, Uttarakhand, Rajaji Nat. Park, 4 km NE Chilla , N 30°0'13.88", E 78°14'37.83", 8.04.2012, Anichtchenko A. leg.” (cAA; DUBC) . LAOS: 1 ♀ — “ Chlaenius (Pseudochlaeniellus) germanus Kirschenhofer, 2001 ”, “ Laos, Vientiane Prov., Vang Vieng, bank of Nam song river, 280 m, at light, No 6. 21.III.1998, leg. O. Merkl & G. Csorba ( NMW); 5 ♀ — “ Laos, Vientiane prov., Vang Vieng 6 km N, Pakpo W env., Nam Song river , 18°59'27,00"N 102°26'20.00"E, 27–28.II.2014, R. Cibulskis leg” (cAA, DUBC) . NEPAL: 1 ♂ —“Chitwan”, “ Chlaenius nigrosuturatus det. Kirschenhofer ” (cDW) . TAIWAN: 1 ♀ —“ Chlaenius callichloris , det. Kirschenhofer 1996”, “ Akau , 15–30.XI.1907, Formosa, Sauter ” ( NMW) . THAILAND: 1 ♂, Paratype—“ Chlaenius (Pseudochlaeniellus) ladon mihi Kirschenhofer ”, “ Thailand: Amphoe Chom Thong , Ban Huai Muang , 4.1. 1989, 250m, leg. Trautner & Geigenmüller ( NMW) . VIETNAM: 1 ♀ —“ Holotypus Chlaenius (Pseudochlaeniellus) ladon mihi Kirschenhofer”, "Cochinchine Mytho—Tanchou , V.48 "; " Chlaenius sp. vestitus Payk. det. Straneo"; "Coll. J. Ochs in Coll. Curti MHNG-1991" (MHNG); 1 ♀— “ Paratypus Chlaenius (Pseudochlaeniellus) ladon mihi Kirschenhofer ”, " Kam Samnar Mekong , 1.48"; " Chlaenius hemichlorus Fairm. ?, det. Curti, M. "; " Coll. J. Ochs in Coll. Curti MHNG-1991" ( MHNG); 2 ♂ — “ Tonkin ” ( NMW).

Diagnosis. This small sized species can be diagnosed easily by yellow translucent lateral margins of pronotum. This important character was also mentioned by Dejean (1826).

Description. Body length 7.3–8.6 mm. Dorsal surface with dense yellow pubescence. Coloration of head and pronotum metallic green with golden or bluish overtones on the disc. Narrow lateral margin of pronotum yellowish translucent. Mouthparts, antennomeres and legs yellow. Antennomeres IV–XI slightly darker. Elytra metallic green, with narrow yellow lateral band on two external intervals, apically enlarged to stair-step apical spot. Ventral surface black, except wide lateral band on all abdominal segments.

Head shiny, without microreticulation, sometimes in females with barely visible microsculpture; near eyes and on vertex moderately punctate. Pronotum strongly weakly and irregularly microreticulate. Punctation on disc deep and irregular, denser towards the base, punctures separated by 1–2 their diameters. Lateral margins narrow. Posterior angles right. Elytra with striae narrow and deep, moderately punctate at the base, punctures gradually disappearing to the apices. Intervals weakly convex at base.

Aedeagus ( Figs. 34–36 ). See comments.

Sexual dimorphism. The only Oriental species of subgenus in which sexual dimorphism is expressed. In females elytra markedly wider.

Comments. Species was described from “Indes orientales” and widely distributed in mid- and lowlands of Oriental region. The study of Paratype of C. (Pseudochlaeniellus) ladon Kirschenhofer, 1998 syn. n. showed that there is no characters outside the range of intraspecific variation. New synonymy is proposed here.

The species C. callichloris Bates, 1873 syn. n. was described after specimens from Japan and Kin-Kiang on the Yang-tsze ( China). Probably this species erroneously cited for Japan, because Habu (1987) don’t saw specimens from Japan, despite the fact that specimens of this species are usually very numerous and common. On the other hand, its presence there is quite possible. Studied specimens from China are conspecific with C. sobrinus . We noticed weak differenced in aedeagal structure between specimens from India ( Figs. 33–34 ) and from Vietnam ( Fig. 35 ), i.e. apical part of median shaft slightly longer in specimens from Vietnam. Perhaps further investigation revealed that the population in the east of the area deserve the status of subspecies.

Distribution. China (Jiangxi; Sichuan; Yunnan), India, Laos, Nepal, Taiwan, Thailand, Vietnam.

Figures

FIGURES 7 – 12. Habitus of C. (Pseudochlaeniellus) sobrinus: 7 — Type. Female (MNHN); 8 — Vietnam. Female (cAA); 9 — C. (P.) ladon syn. n., Paratype. Vietnam (NMW); 10 — China. Male (cAA); 11 — Tonkin, Male (NMW); 12 — Ramnagar, India. Male (cAA).

FIGURES 28 – 36. Aedeagus: 28 — Holotype of C. chloodes = C. neochloodes (BMNH); 29 — C. caesitius, Loc. Typ. (NMW); 30 — C. puncticollis. India (cAA); 31 — Paratype of C. mesopotamicus = lucasi (NMW); 32 — C. germanus, Karnataka, India (cAA); 33 — C. germanus, Sylhet, Bangladesh. (BMNH); 34 - 35 variability of aedeagus of C. sobrinus, Rajaji, India (cAA); 36 — C. sobrinus, Tonkin, Vietnam (NMW).