Clito jonkersi

Grishin, Nick V., 2014, Two new species of Clito from South America and a revision of the Clito littera group (Lepidoptera: Hesperiidae: Pyrginae), Zootaxa 3861 (3), pp. 231-248 : 238

C. jonkersi de Jong, 1983

( Figs. 19–20 , 26 i, 27 part, 28 part)

C. jonkersi from Suriname (TL) and French Guiana (B. Hermier, pers. comm.) differs most prominently from the rest ( Figs. 19–20 , 26 i, 27 part), and is characterized by having a short and stout process off the costa in the valva. Hindwing of C. jonkersi is with a narrower pale discal band, edges of the band are better defined, spot in Sc+R1-Rs cell is rhomboid and is indented basad relatively to the spot at the base of Rs-M1 cell. Ventrally, C. jonkersi hindwing is with a row of well-defined submarginal spots; these spots are absent or indistinct in other species ( Figs. 19-20 , 27 part).

Figures

FIGURE 26. Male genitalia of Clito littera group species. a, g. C. littera lectotype, designated here, " Brasil ", P. Mabille collection 1923, R. Oberthür Coll., Brit. Mus. 1927 - 3, specimen No. BMNH (E) # 982948 [BMNH], in situ on a specimen, scales brushed off: a. various views rotating from right posterolateral through posterior to left posterolateral; g. ventrolateral (specimen Figs. 9 – 10); b, c. C. litteroides holotype, Venezuela: Aragua State, Rancho Grande [Biological Station, approximate GPS coordinates 10 ° 21 ' N 67 ° 41 ' W], elevation 1100 m, 4 - Jun- 1985, leg. S. S. Nicolay, genitalia vial NVG 130614 - 42 [USNM]: b. section of complete genitalia in posterolateral view; c. complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 7 – 8); d. C. nebulosa holotype, ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Mielke & Schröder (1994) (specimen Figs. 15 – 16); e, f. C. trinidadensis holotype, Trinidad, Morne Catherine, 24 - Mar- 1982, leg. M. J. W. Cock, specimen No. BMNH (E) # 982949 [BMNH], dorsal and lateral views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis, illustration from Cock (1998) (specimen Figs. 17 – 18); h. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall & M. A. Solis [USNM], complete genitalia in lateral view and uncus with distal part of tegumen in dorsal view (specimen Figs. 5 – 6); i. C. jonkersi holotype, Surinam, Domburg, 8 - Mar- 1975, leg. E. H. Jonkers [RMNH], lateral internal view of left valva and lateral view of uncus and gnathos, illustration from de Jong (1983) (specimen Figs. 19 – 20); j. C. palotschka paratype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, {25, 27} - Aug- 2002, leg. J. P. W. Hall & M. A. Solis [USNM], right valva and uncus in situ on a specimen, scales brushed off (specimen Figs. 3 – 4); k. C. palotschka holotype, Ecuador: Esmeraldas Prov., El Durango, km 39 Lita – San Lorenzo road, 1 ° 02.45 ' N, 78 ° 38.06 ' W, 230 - 300 m, 4 - Mar- 2001, leg. D. H. Ahrenholz [USNM], right valva in situ on a specimen, scales brushed off (specimen Figs. 1 – 2); l. C. anda, Brazil: Rondônia, Cacaulândia municipality, Fazenda Rancho Grande, genitalia No. GTA 4377, illustration from Austin (2000), ventral, lateral and dorsal views of tegumen, uncus, gnathos and associated structures, lateral internal view of right valva, lateral view of penis; m – o. C. anda [holo] type, French Guiana, Collect. C. Bar, R. Oberthür Coll., Brit. Mus. 1931 – 136, specimen No. BMNH (E) # 982947 [BMNH] (specimen Figs. 11 – 12), left valva on m. and Evans's sketches: n. pinned in the collection [BMNH]; o. published, and more stylized, version (Evans 1953), lateral internal view of left valva, lateral and ventral views of uncus, gnathos and distal half of penis. Species name is given below each image, holotypes and paratypes are labeled as HT and PT. Gray " F " indicates mirror image (left-right inverted). Images a, f, g, m – o. are copyright (©) Trustees of the Natural History Museum, London (used with permission).

FIGURE 27. Visual keys to species in the Clito littera group. Dorsal and ventral aspects for each species are shown on the left and right of the species name, respectively. Images shown are illustrations, sometimes composed of photographic segments from left and right sides and digitally edited to highlight the wing patterns instead of imperfections in specimens. Images are set to be approximately the same size. Unedited and to scale photographs of specimens are shown in Figs. 1 – 20. Since these species are defined on the basis of male genitalia, which are likely to be diagnostic, wing pattern characters are preliminary due to exceedingly small samples of available specimens, and exceptions are expected. Wing pattern characters should be treated as suggestions for further research. Photographs of C. jonkersi and C. nebulosa are by Bernard Hermier and Ernst Brockmann, respectively. Images of C. littera, C. trinidadensis and C. anda are copyright (©) Trustees of the Natural History Museum, London (used with permission).

FIGURES 17 – 23. Type specimens and illustrations of Clito from the C. littera group. 17 – 18. C. trinidadensis holotype, Trinidad, Morne Catherine, 24 - Mar- 1982, leg. M. J. W. Cock, specimen No. BMNH (E) # 982949 [BMNH] (genitalia shown in Figs. 26 e, f); 19 – 20. C. jonkersi holotype, Surinam, Domburg, 8 - Mar- 1975, leg. E. H. Jonkers [RMNH] (genitalia Fig. 26 i); 21. C. littera, unpublished illustration, likely of the lectotype, dorsal—left, ventral—right, pinned in the collection next to the specimen [BMNH]; 22. C. nebulosa original illustration of the holotype from Draudt (1924) dorsal side, half-specimen image is shown for easy comparison on 23. (and Figs. 15 – 16). Dorsal and ventral surfaces are shown on odd- and even-numbered figures, respectively, except illustrations. Labels are shown between the specimen images. Labels are reduced 2.5 times compared to specimens: small scale bar below C. trinidadensis locality labels refers to labels, and larger scale bars refer to specimens. Photographs 19 – 20 are by Bernard Hermier, images 17 – 18 and 21 are copyright (©) Trustees of the Natural History Museum, London (used with permission).