Calathus (Certocalathus), Schmidt & Will, 2020

Schmidt, Joachim & Will, Kipling, 2020, A new subgenus for “ Acalathus ” advena (LeConte, 1846) and the challenge of defining Calathina based on morphological characters (Coleoptera, Carabidae Sphodrini), Zootaxa 4722 (4), pp. 326-338 : 332

Subgenus Certocalathus nov.

Third antennomere without pubescence. Mentum tooth bifid. Pronotal laterobasal seta situated at margin. Lateral bead of prosternal process absent. Parascutellar seta present. Elytra with very short, fine pubescence across all intervals for length of elytra, or pubescence restricted to the outer two intervals and apical third; with 2-3 discal setae in the third interval; discal setae absent in the fifth and seventh intervals. Elytral microsculpture nearly isodiametric. Metepisternum elongated. Male protarsomeres dilated. Lateral grooves present on metatarsomeres I-IV. Apical gonocoxite short, robust, with two ensiform setae at external margin; sensory pit of apical gonocoxite welldeveloped. Aedeagus left side up in repose; median lobe apical disc absent; apical hook of long paramere present; membranous lobe of short paramere present.