taxonID	type	description	language	source
03C2D37AFFEBFFDFBDF4DBC3FD6BFAF4.taxon	description	urn: lsid: zoobank. org: act: 3 DA 55 FF 8 - 7 A 75 - 4 FD 7 - 83 F 0 - 650 AB 759 F 868 (Figs 1, 2 and 3)	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFEBFFDFBDF4DBC3FD6BFAF4.taxon	materials_examined	Material examined Holotype. INDIA: West Bengal, / North 24 - Parganas (N), / Mondouri, / ex Mangifera indica L. / 12. vi. 2023, coll. A. Das, 1 adult ♀ mounted on slide (ZSI, Registration No. 19676 / H 15) Paratypes. Collection data same as for holotype, 5 adult ♀♀ mounted singly on 5 slides (ZSI, Registration Nos. 19677 / H 15 to 19681 / H 15); and 10 adult ♀♀ mounted singly on 10 slides (BCKV). Total number of paratype specimens = 15. Additional, non-type specimens. Collection data same as for holotype, 20 slide-mounted adult ♀♀ (BCKV).	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFEBFFDFBDF4DBC3FD6BFAF4.taxon	description	Adult female description Appearance in life (Fig. 1): Body oval, moderately convex above, with the whole dorsal surface symmetrically reticulated on each side with dark brown lines forming various rectangular to polygonal cell-like patterns (Figs 1 A, B). In young specimens this patterning is slight or absent, but is obvious in comparatively mature females; later, the cell-like pale areas become sclerotized and the entire dorsum becomes dark brown except for the marginal areas. Found on stems and leaves of Mangifera indica L., associated with the ant, Oecophylla smaragdina Fabricius (Formicidae: Hymenoptera) (Fig. 1 D); a large number of ants were found attending the exposed scale insect colony (Fig. 1 C). Slide-mounted adult female (n = 36, but measurements are based on the holotype and 9 paratypes) (Figs 2 and 3). Body broadly oval (Fig. 2 A), 1.7 – 2.8 mm long, 1.4 – 1.9 mm wide; length to width ratio 1.3 – 1.5: 1; anal cleft 380 – 520 μm long, roughly one-fifth of body length. Dorsum. Derm membranous, marked by small, clear, circular-to-oval cell-like areolations (Figs 2 K, 3 L), widely spaced, maximum areolation width 7.5 – 12.5 μm, distributed over the whole dorsum, each areolation containing 1 microduct (Fig. 3 L 1). Dorsal setae spinose, neither thick nor thin, straight (Figs 2 H 1, 3 I 1) or very slightly curved (Figs 2 H 2, 3 I 2), with apices pointed and basal sockets well developed, each 7.5 – 12 μm long; sparsely distributed over entire dorsum, slightly more frequent posteriorly. Dorsal submarginal tubercles (Figs 2 G, 3 J) numbering 2 on each side, each 15 μm wide in dorsal view. Preopercular pores and dorsal tubular ducts absent. Anal plates each triangular, each plate 115 – 150 μm long, 75 – 100 μm wide, with anterolateral margin (100 – 125 μm long), sub-equal in length to posterolateral margin (90 – 120 μm long); plates together roughly quadrate (Figs 2 L, 3 H); each plate with 4 apical and subapical setae. Anogenital fold with a total of 2 pairs of long anterior margin setae (Fig. 2 M), each 70 – 85 μm long, and each side with 2 lateral margin setae (Fig. 2 N), each 65 – 85 μm long. Anal ring elongate oval, maximum length 40 ‒ 55 μm, with 2 irregular rows of translucent pores, bearing 8 (rarely 7) long setae (Fig. 2 J), each 170 – 220 μm long. Eyes (Figs 2 E, 3 M) present dorsally and submarginally, each situated within a clear circular area about 10 μm in diameter. Margin. Stigmatic clefts deep, obvious, lacking sclerotization, each containing 3 spiracular setae (Figs 2 R 1, 3 K 1), each with a well-developed basal socket; median seta longest, slender and about 55 – 75 μm long, lateral setae smaller, generally sub-equal in length, each 20 – 35 μm long; a few spiracular clefts with 2 lateral spines uneven in length (Figs 2 R 2, 3 K 2), 1 about twice as long as other; all spiracular setae straight or slightly curved apically, and with pointed or bluntly pointed apices. Marginal setae easily distinguishable from stigmatic spines, each seta 21 – 45 μm long, slender, with well-developed basal socket (Figs 2 I 1 – I 6, 3 G 1 – G 5); setae highly variable in structure, with various degrees of curvature between apically slightly wavy to sickle-shaped or abruptly bent; about 50 % of setae simple with bluntly pointed apices, the rest weakly fimbriate or a few bifid apically. Distribution of marginal setae as follows: with 60 – 63 anteriorly between anterior spiracular furrows and, on each side, with 13 – 15 between anterior and posterior spiracular furrows, and 46 – 50 between posterior spiracular furrow and anal cleft. Venter. Derm membranous; ventral tubular ducts absent; microducts not seen. Submarginal setae (Figs 2 D, 3 E) mostly forming a row parallel to margin, each 8 – 10 μm long, setose, very slender and straight with a pointed apex. Other ventral setae short, simple, straight and sharply-pointed apically, similar to submarginal setae, each 7 – 8 μm long, scattered sparsely throughout. Interantennal setae (Fig. 2 T) numbering 2 or 3 pairs of variable lengths, each seta 40 – 95 μm long. Pregenital long setae (Fig. 2 O) numbering 3 segmental pairs, each seta 70 – 125 μm long. Antennae well developed, each with 7 or 8 segments (Figs 2 B, 3 B), total length 240 – 375 μm; lengths of segments in an 8 - segmented antenna (in μm): segment I, 30 – 40; II, 35 – 45; III, 45 – 65; IV, 35 – 55; V, 25 – 40; VI, 20 – 35; VII, 20 – 34, and VIII, 45 – 60. Clypeolabral shield (Fig. 2 P) 85 – 110 μm long, 100 – 110 μm wide, without setae. Labium 40 – 50 μm long and maximum 45 – 55 μm wide. Legs well developed (Figs 2 C, 3 C); all legs with a sclerotized tibio-tarsal articulation; tarsal digitules slender, knobbed, extending beyond claw digitules claw lacking denticle; claw digitules broad, equal, expanded at apex; (Fig. 3 C). Lengths of prothoracic leg segments (in μm): coxa 65 – 75, trochanter 45 – 57, femur 80 – 88, tibia 80 – 94, tarsus 48 – 58, claw 16 – 17, tarsal digitule 35 – 40, and claw digitule 25 – 30. Lengths of mesothoracic leg segments (in μm): coxa 72 – 85, trochanter 50 – 55, femur 88 – 95, tibia 85 – 95, tarsus 55 – 65, claw 17 – 18, tarsal digitule 35 – 40, and claw digitule 30 – 35. Lengths of metathoracic leg segments (in μm): coxa 85 – 100, trochanter 50 – 60, femur 90 – 105, tibia 85 – 98, tarsus 60 – 68, claw 20 – 24, tarsal digitule 35 – 50, and claw digitule 25 – 30. Spiracles all similar and sub-equal in size (Fig. 2 S), each 18 – 25 μm wide across atrium. Spiracular disc-pores forming band 1 or 2 pores wide (Figs 2 Q, 3 D) between each spiracle and margin; each spiracular disc-pore 5 – 6 μm in diameter, with 5 or 6 loculi (mostly 5); each anterior spiracular pore band containing 16 – 20 pores, each posterior band containing 16 – 28 pores. Pregenital disc-pores often difficult to locate, present in a group of about 25 – 35 pores near or around anogenital fold (Fig. 3 F), each pore 8 – 10 μm in diameter and with 7 – 9 (rarely 10) loculi, occasionally with 1 or 2 pores present medially on preceding 1 or 2 segments.	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFEBFFDFBDF4DBC3FD6BFAF4.taxon	etymology	Etymology. The species epithet mondouriensis is formed from the name of the type locality, Mondouri, combined with the Latin suffix “ - ensis ”, meaning “ from Mondouri ”.	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFEBFFDFBDF4DBC3FD6BFAF4.taxon	discussion	Comments. Of the 13 Coccus species previously known from India, the adult female of C. mondouriensis sp. nov. is most similar to those of C. ophiorrhizae (Green) and C. longulus (Douglas), both of which also lack dorsal and ventral tubular ducts. In addition to lacking tubular ducts, C. mondouriensis shares with C. ophiorrhizae the following character states: (i) absence of preopercular pores; (ii) anogenital fold with a total of 4 anterior margin setae, and 2 lateral margin setae on each side; and (iii) interantennal setae usually numbering 2 pairs. However, C. mondouriensis can easily be distinguished from C. ophiorrhizae by the presence of the following features (character states of C. ophiorrhizae in parentheses): (i) marginal setae variable in form, with apices either bluntly pointed or truncate, or weakly fimbriate to bifurcate (marginal setae only simple, without any with apices fimbriate, branched or bifurcate), and (ii) dorsal setae spinose, straight or very slightly curved (dorsal setae cylindrical). According to the literature (Hamon & Williams 1984; Gill 1988; Tang 1990), Coccus longulus appears to be morphologically variable; in case it is a cryptic species complex, the following comparison is based only on the data for C. longulus given by Ben-Dov (1977), who studied the type specimens. Coccus mondouriensis is similar to C. longulus in lacking tubular ducts and discal setae on anal plate, and in having dorsal submarginal tubercles. However, it can be distinguished from C. longulus by the following features (character states of C. longulus in parentheses): (i) preopercular pores absent (with 4 – 14 preopercular pores); (ii) body broadly oval (body distinctly elongate oval, often with lateral margins almost parallel); (iii) dorsal setae spinose, straight or very slightly curved (simple and conspicuously curved); (iv) each anal plate with 4 apical and subapical setae (with 5 apical and subapical setae); (v) anogenital fold with a total of 4 anterior margin setae and 2 lateral margin setae on each side (with 6 – 9 anterior margin setae and 3 lateral margin setae on each side); and (vi) interantennal setae numbering 2 or 3 pairs (interantennal setae absent [based on Ben-Dov 1977: 90, fig. 1]). Cao et al. (2022) described C. nanningensis Cao & Feng from China, feeding on Ficus carica (Moraceae). This species also resembles C. mondouriensis in having (i) similar types of marginal setae, and in lacking (ii) dorsal and ventral tubular ducts and (iii) preopercular pores. Coccus mondouriensis may be distinguished from C. nanningensis by the presence of the following features (data for C. nanningensis in parentheses): (i) dorsal setae spinose, straight or very slightly curved (dorsal setae setose, extremely fine and pointed), (ii) each anal plate with 4 apical and subapical setae, but lacking discal setae (each plate with 3 apical setae and a robust discal seta); (iii) anogenital fold with a total of 4 anterior margin setae and 2 lateral margin setae on each side (with 6 anterior margin setae and 3 lateral margin setae on each side); (iv) marginal setae numbering 60 – 63 between the two anterior spiracular furrows (39 – 45) and, on each side, 13 – 15 between anterior and posterior spiracular furrows (10 – 12), and 46 – 50 between posterior spiracular furrow and anal cleft (22 – 25); and (v) pregenital setae numbering 3 pairs (mainly 2 pairs). It is therefore considered that C. mondouriensis is a species new to science that is morphologically close to both C. ophiorrhizae and C. nanningensis. It is associated with the ant, Oecophylla smaragdina; a large number of which were found attending the colony outside the ant’s nest.	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFECFFD2BDF4D8B8FEAEFB64.taxon	description	urn: lsid: zoobank. org: act: A 4 E 327 BD-F 77 D- 4 CF 9 - B 49 D-AF 45 FA 8 D 119 C (Figs 4, 5 and 6)	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFECFFD2BDF4D8B8FEAEFB64.taxon	materials_examined	Material examined Holotype. INDIA: West Bengal, / 24 - Parganas (N), Mondouri, / ex. Citrus aurantifolia / (Christm.) / Swingle, 7. v. 2024, coll. A. Das, inside the nest of Oecophylla smaragdina Fab., 1 adult ♀ mounted on a slide (ZSI, Registration No. 19682 / H 15). Paratypes. Collection data same as for holotype, 5 adult ♀♀ mounted singly on 5 slides (ZSI, Registration Nos. 19683 / H 15 to 19687 / H 15); and 5 adult ♀♀ mounted singly on 5 slides (BCKV). Total number of paratype specimens = 10. Additional, non-type specimens. Collection data same as for holotype, 8 slide-mounted adult ♀♀ (BCKV).	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFECFFD2BDF4D8B8FEAEFB64.taxon	description	Adult female description Appearance in life (Fig. 4): Body broadly oval to elongated oval; median area slightly convex, longitudinal median ridge apparent in lateral view; body light yellowish, marginal area transparent; the whitish ventral stigmatic bands and colour of the host plant to which it is attached visible through dorsum (Figs 4 C, D); median area of body turning brown with age. Eyes black, prominent, situated very close to margin. Coccus oecophyllae sp. nov. was found inside the nest of the weaver ant, Oecophylla smaragdina F. (Fig. 4 A), which the ants had built by binding the leaves together with white silk. The scale insects were found feeding on leaves (Fig. 4 B) and a stem (Fig. 4 C) within the nest. Slide-mounted adult female (n = 19, but measurements are based on the holotype and 7 paratypes only) (Figs 5 and 6). Body broadly oval (Figs 5 A, 6 A), 2.7 – 3.1 mm long, 1.6 – 1.8 mm wide; anal cleft 400 – 480 μm long, onesixth to one-seventh of length of body. Dorsum. Derm membranous with small, very pale dermal round-to-oval areolations, not numerous. Dorsal setae (Figs 5 Q 1 – Q 3, 6 J 1 – J 4) moderately numerous, scattered over entire dorsum, each 8 – 13 μm long; setae of two types present: (i) most frequent type spinose, straight or very slightly curved with pointed or slightly bluntly pointed apex (Figs 5 Q 2 – Q 3, 6 J 1 – J 2); and (ii) cylindrical setae each with a very slightly enlarged and rounded apex, present less frequently (Figs 5 Q 1, 6 J 3, J 4). Dorsal submarginal tubercles (Figs 5 E, 6 N) conspicuously raised and convex, sparse, numbering 6 – 9 in total, present adjacent to margins, each about 10 μm wide and 15 – 23 μm high. Preopercular pores absent. Microducts present throughout, each situated within an areolation. Dorsal tubular ducts (Figs 5 O, 6 E) few, numbering 4 – 6 in total, present on the submargin except between two anterior clefts, each with a broad outer ductule, 8 – 10 μm long and 2 μm wide, and a thin inner ductule about 15 μm long with a small terminal gland. Each anal plate triangular (Figs 5 K, 6 I), 137 – 160 μm long and 85 – 100 μm wide, plates together roughly quadrate with anterolateral margin (105 – 125 μm) and posterolateral margin (110 – 120 μm); each plate with 4 apical and subapical setae, each 20 ‒ 23 μm long. Anogenital fold with a total of 2 pairs of anterior margin setae (Fig. 5 L), each 65 – 85 μm long, and each lateral margin with 2 setae (Fig. 5 M), each 75 – 92 μm long. Anal ring subcircular, 35 – 52 μm wide, bearing 6 long setae, each 160 – 180 μm long; pores in anal ring forming 2 irregular rows. Anal tube 150 – 200 μm long. Eyes (Figs 5 F, 6 M) situated near margin, each 20 – 25 μm wide. Margin. Stigmatic clefts well developed, weakly sclerotized; each cleft containing 3 spiracular setae (Figs 5 S 1, 6 L 1), usually with 1 long median seta, 60 – 65 μm long, straight to slightly curved apically and bluntly pointed, and 2 smaller, straight lateral setae with pointed apices, subequal in length, each 22 – 32 μm long, about half to one-third as long as median seta. Occasionally spiracular setae are grossly subequal in length (each 40 – 60 μm long) (Figs 5 S 2, 6 L 2), and rarely with longest seta situated laterally rather than in medial position (Figs 5 S 3, 6 L 3). Marginal setae (Figs 5 J 1 – J 7, 6 H 1 – H 6) mostly small, slightly curved to sickle-shaped or even abruptly curved, each 12 – 35 μm long, with apices usually bifid or weakly to moderately fimbriate, but occasionally some simple; anal lobe margin with a few longer stout setae, each about 30 ‒ 40 μm long and straight to slightly curved with a well-developed basal socket. Distribution of marginal setae as follows: 27 – 31 anteriorly between two anterior spiracular furrows and, on each side, with 10 – 12 between anterior and posterior spiracular furrows, and 22 – 24 between posterior spiracular furrow and anal cleft. Venter. Derm membranous. Submarginal setae (Figs 5 D and 6 F 1, F 2) slender, straight and spinose, with sharply pointed apices, each 7 – 15 μm long. Interantennal setae (Fig. 5 V) numbering 2 pairs, each seta 50 ‒ 65 μm long. Pregenital long setae (Fig. 5 N) numbering 3 pairs, each seta 60 – 85 μm long, occasionally with 1 seta absent from one side, making the total number 5. Other ventral setae similar to submarginal setae, straight or curved, each 10 – 30 μm long, sparsely distributed. Antennae (Figs 5 B, 6 B) well developed, mostly each with 7 (occasionally 8) segments, each 250 – 285 μm long; lengths of segments in a 7 segmented antenna (in μm): segment I, 42; II, 30; III, 57; IV, 55; V, 18; VI, 28; and VII, 50. Clypeolabral shield (Fig. 5 U) 110 – 125 μm long, 90 – 100 μm wide, without setae; labium 50 – 65 μm long and 60 – 75 μm wide. Legs (Figs 5 C, 6 C) well developed, each with a tibio-tarsal articulatory sclerotization; tarsal digitules thin, setose, knobbed, extending beyond claw digitules claw digitules equal, broad, expanded at apex; claw lacking denticle. Lengths of prothoracic leg segments (in μm): coxa 63 – 75, trochanter 38 – 78, femur 92 – 100, tibia 80 – 88, tarsus 60 – 65, claw 13 – 20, tarsal digitule 33 – 40, claw digitule 20 – 24; mesothoracic leg segments (in μm): coxa 75 – 85, trochanter 45 – 50, femur 100 – 108, tibia 92 – 100, tarsus 65 – 74, claw 15 – 18, tarsal digitule 32 – 42, claw digitule 17 – 20; metathoracic leg segments (in μm): coxa 95 – 100, trochanter 50 – 65, femur 107 – 115, tibia 85 – 95, tarsus 65 – 75, claw 14 – 20, tarsal digitule 37 – 50, claw digitule 22 – 30. Spiracles all similar, each 50 – 65 μm long and 27 – 30 μm wide across peritreme. Sometimes with very shallow spiracular furrow between each spiracle and margin, lacking sclerotization. Spiracular disc-pore band 1 or 2 pores wide between each spiracle and stigmatic cleft (Figs 5 R, 6 D), and 2 or 3 pores may extend beyond spiracle; each anterior pore band with 16 – 26 pores, and posterior band with 19 – 32 pores; each spiracular disc-pore 3 – 5 μm in diameter, containing 5 loculi. Pregenital disc-pores (Figs 5 I, 6 G) numbering about 30 (27 ‒ 37), each 8 – 10 μm in diameter, with 9 or 10 peripheral loculi and 1 central loculus, present around anogenital fold; sometimes also a few present on medial area of preceding abdominal segment. Ventral tubular ducts (Figs 5 O, 6 E) all of one type, each with an outer ductule 12 – 18 μm long and 2 – 3 μm wide and a narrow inner ductule about 15 – 18 μm long and 1.5 μm wide with a flowershaped terminal gland, scattered mostly medially near mouthparts, on thorax and a few laterally on some abdominal segments: with a group of 3 or 4 ducts located medially just posterior to labium, and, on each side, with about 4 ducts between clypeolabral shield and procoxa; a comparatively broad transverse band of about 9 ducts present mesad to mesocoxae; with 3 between each meso- and metacoxa, and 4 medially at same level; 5 or 6 sparsely scattered on inner submarginal area laterad to hind coxa; and 1 on each submarginal area of abdominal segments V and VII. Microspines (Fig. 5 P) present on derm of anal lobe, distributed in small linear groups.	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFECFFD2BDF4D8B8FEAEFB64.taxon	etymology	Etymology. The species epithet oecophyllae, meaning “ of Oecophylla ”, refers to the intimate association of the scale insect with the ant species Oecophylla smaragdina (oecophyllae: genitive case, feminine singular).	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFECFFD2BDF4D8B8FEAEFB64.taxon	discussion	Comments. Choi & Lee (2018) gave the diagnostic morphological characters of adult females of 11 Asian species of Coccus possessing ventral tubular ducts, including the description of a new species, Coccus ficicola Choi & Lee from China, feeding on Ficus benghalensis L. (Moraceae). Comparing C. oecophyllae sp. nov. with these 11 species and other Coccus species present in India, the new species is found to have a novel set of morphological traits, including an unusual distribution pattern of ventral tubular ducts. Morphologically, C. oecophyllae sp. nov. is closest to C. ficicola in sharing common character states like the presence of: (i) dorsal submarginal tubercles and tubular ducts on both dorsum and venter; (ii) two pairs of interantennal setae; (iii) each anal plate with four apical and subapical setae, but lacking discal setae; (iv) anogenital fold with a total of four anterior margin setae; and (vii) tibio-tarsal scleroses. However, C. oecophyllae may be easily distinguished from C. ficicola by the presence of the following features (contrasting character states for C. ficicola are given in parentheses): (i) dorsal setae of two types: spinose with pointed or slightly bluntly pointed apices, and cylindrical with very slightly enlarged and rounded apices (dorsal setae of one type, cylindrical with blunt apices); (ii) marginal setae with apices usually strongly bifid or weakly to moderately fimbriate, but occasionally a few with simple tips; (mostly with simple pointed apices, but sometimes with slightly bifid tips); (iii) marginal setae numbering 27 ‒ 31 between the two anterior spiracular furrows (52 ‒ 59), and, on each side, 10 ‒ 12 between anterior and posterior spiracular furrows (12 – 17), and 22 ‒ 24 between posterior spiracular furrow and anal cleft (45 ‒ 53); (iv) preopercular pores absent (preopercular pores present); (v) antennae each generally seven segmented (eight segmented); (vi) ano-genital fold with two lateral margin setae on each side (three); and (vii) pregenital disc pores each with 10 ‒ 12 loculi (nine or 10 loculi). Coccus oecophyllae sp. nov. is also similar to C. discrepans (Green), as both species have: (i) dorsal submarginal tubercles and ventral tubular ducts; (ii) preopercular pores absent; (iii) apices of marginal setae frayed, bifurcated or pointed; (iv) antennae each with seven segments (occasionally eight); (v) pregenital setae numbering three pairs; (vi) tibio-tarsal scleroses present; (vii) anogenital fold with two pairs of anterior margin setae and each lateral margin setae with two setae; and (ix) two pairs of interantennal setae. However, C. oecophyllae can be distinguished from C. discrepans by the following (character states of C. discrepans are given in parentheses): (i) dorsal setae of two types, spinose and cylindrical (one type, large and spinose as found in India by Avasthi & Shafee (1991) or short and rod-like with apices blunt or slightly pointed, as found in Taiwan by Tao et al. (1983 )); (ii) dorsal tubular ducts present (dorsal tubular ducts absent); (iii) ventral tubular ducts present on medial areas of pro-, meso- and metathorax and submarginal areas of metathorax and abdomen (present on medial area of mesothorax, and with a few laterad to mid-coxae, not present on abdomen); and (iv) claw digitules similar in size and shape (different in size and shape). It is therefore considered that C. oecophyllae is a species new to science that is morphologically close to C. ficicola and C. discrepans. The new species is associated with the ant, Oecophylla smaragdina, and was found inside the nest.	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
03C2D37AFFE1FFD5BDF4D9C9FA0CFD1F.taxon	discussion	Note: As the available description of Coccus ramakrishnai (Ramakrishna Ayyar) is inadequate, it is not included in the key.	en	Das, Anirban, Pramanik, Aniruddha, Das, Bijan Kumar (2025): Two new ant-attended species of the soft scale insect genus Coccus L. (Hemiptera: Coccomorpha: Coccidae) from India, with a key to Indian Coccus species. Zootaxa 5647 (2): 131-144, DOI: 10.11646/zootaxa.5647.2.3, URL: https://doi.org/10.11646/zootaxa.5647.2.3
