taxonID	type	description	language	source
03C39202C100750D48ABFD39D75EFEAF.taxon	type_taxon	Type species. Septsigilla tongbiguan Zhang & Zhang sp. nov.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C100750D48ABFD39D75EFEAF.taxon	etymology	Etymology. The generic name is a combination derived from the Latin “ septem ” (seven) and “ sigilla ”, referring to the seven sigilla on the sternum of the species described herein. The gender is neutral.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C100750D48ABFD39D75EFEAF.taxon	diagnosis	Diagnosis. This new genus shares similarities with other genera of Systariinae in possessing a compact eye group, separated ALS, relatively longer leg III (not less than 90 % of the length of leg II), notched trochanters, the presence of a cymbial groove and the absence of a cymbial spur on the male cymbium, as well as posteriorly positioned spermathecae and anteriorly positioned bursae on the female epigyne, but can be distinguished from the other genera in this subfamily (except for Tamin) by having the smallest anterior median eyes. The male can be distinguished from Tamin by the posteriorly elongated tegulum which extends to the median part of the tibia, and the whiplike embolus (Figs 3 A – C, 6 A – C; vs. the non-elongated tegulum and triangular embolus in Tamin, see Deeleman-Reinhold 2001: 213, fig. 250). The female differs from Tamin in that the bursae are almost as large as the primary spermathecae, and the connecting tubes are sac-like (Figs 3 E, 6 E, vs. the bursae significantly larger than the primary spermathecae and the connecting tubes being narrow and tubular in Tamin, see Deeleman-Reinhold 2001: figs 243, 256).	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C100750D48ABFD39D75EFEAF.taxon	description	Description. Medium-sized spiders (5.38 – 6.56 mm), prosoma yellowish brown, with black radial patterns; radial and cervical grooves indistinct; fovea reddish-brown, longitudinal, presenting at posterior part of thoracic region (Figs 1 A, C, 4 A, C). Ocular area narrow and dark, ca. 0.6 times width of CRW; AME smallest, median eyes circular, lateral eyes oval-shaped, all eyes with black rings, anterior eye row slightly recurved, posterior eye row almost straight in dorsal view (Figs 2 A – B, 5 A – B). Chelicera yellowish brown, with three promarginal teeth and two or three retromarginal teeth. Endite longer than wide, without fold or impression (Figs 1 B, D, 4 B, D). Sternum shield-shaped, longer than wide, with seven sigilla, paired sigilla with nearby slit sensilla (Figs 1 B, 2 C, 4 B, 5 C). Legs pale yellow, leg formula 4123. Leg III about 0.9 times the length of Leg II. Coxa Ⅰ with retrocoxal window (RCW, Fig. 7), metatarsi III and IV with conspicuous preening brushes, tibiae III and IV, metatarsi III and IV each with three pairs of robust ventral spines. Leg tarsi with relatively degenerated claw tufts (Fig. 7). Opisthosoma oval, dorsally brownish yellow to gray, with two pairs of muscular impressions (Figs 1 A, C, 4 A, C). ALS spacing about half their diameter, tapering distally; PMS slender and short; PLS with two segments, basal segment cylindrical and similar in length to ALS, distal segment conical, shorter than half length of basal segment (Figs 2 D – E, 5 D – E). Male Palp. Tibia length and width nearly equal, ventral side with one small basal apophysis. RTA slender, with rectangular extension at base in ventral view, curved to ventral side in retrolateral view. Cymbium with basal concavity on retrolateral side. The posterior elongation of tegulum extending to median part of tibia. Sperm duct running along distal and retrolateral margins of tegulum. Embolic base spherical, half of base membranous. Conductor membranous, originating from distal part of tegulum, edges rolling upward (Figs 3 A – C, 6 A – C, 8). Epigyne. Epigynal plate slightly sclerotized. Atrium triangular, posteriorly positioned, carrying sclerotized anterior margin. Copulatory opening locating below sclerotized anterior margin of atrium, not visible in ventral view (Figs 3 D, 6 D). Primary spermatheca and secondary spermatheca heavily sclerotized, connected to each other. Copulatory duct short and thick. Bursa slightly sclerotized, approximately oval-shaped. Connecting tube flat and sac-like (Figs 3 E, 6 E). Composition. Septsigilla tongbiguan Zhang & Zhang sp. nov.; S. zhongi Zhang & Zhang sp. nov.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C100750D48ABFD39D75EFEAF.taxon	distribution	Distribution. China (Xizang, Yunnan).	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C103750A48ABFE19D74EFD47.taxon	description	Figs 1 – 3, 7 A, 8 A, 9	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C103750A48ABFE19D74EFD47.taxon	materials_examined	Type material. Holotype ♂ (MHBUARA- 2024 - 887 - 1), Yunnan, Yingjiang County, Tongbiguan Town, 24.5915 ° N, 97.6776 ° E, 1336.2 m elev., 9 October 2024, leg. K. Yu, Y. Ni and Y. Xu. Paratypes. 6 ♂ 4 ♀ (MHBUARA- 2024 - 887 - 2 – 11), same data as holotype. Other material examined. 1 ♂ 2 ♀ (MHBUARA- 2024 - 881 - 1 – 3), Yunnan, Yingjiang County, Tongbiguan Town, 24.6268 ° N; 97.6395 ° E, 1371.6 m elev., 5 October 2024, leg. K. Yu, Y. Ni and Y. Xu; 2 ♀ (MHBUARA- 2024 - 64 - 1 – 2), Yunnan, Longling County, Longjiang Town, 98.7670 ° E, 24.8259 ° N, 2183 m elev., 5 April 2024, leg. J. Zhao and X. Qi.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C103750A48ABFE19D74EFD47.taxon	etymology	Etymology. The specific name is a noun in apposition derived from the type locality.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C103750A48ABFE19D74EFD47.taxon	diagnosis	Diagnosis. The new species resembles Septsigilla zhongi sp. nov., but can be distinguished by: 1) chelicera with three retromarginal teeth (vs. chelicera with two retromarginal teeth); 2) the patella without apophysis (DPA, Figs 3 A, C; vs. the patella presents an apophysis on its dorsal part, Figs 6 A, C); 3) the smooth embolus without any apophysis (Figs 3 A ‒ C, 8 A; vs. the embolus with three small embolic teeth on its distal two thirds, Figs 6 A ‒ C, 8 B); 4) the top margin of the connecting tube obviously higher than the bursa (Fig. 3 E; vs. the top margin of the connecting tube close to the bursa, Fig. 6 E).	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C103750A48ABFE19D74EFD47.taxon	description	Description. Male holotype (Figs 1 A ‒ B, 2 A ‒ D) total length 6.52; prosoma 3.33 long, 2.63 wide; opisthosoma 3.20 long, 1.80 wide. Eye sizes and interdistances: AME 0.09, ALE 0.13, PME 0.12, PLE 0.13; AME ‒ AME 0.09, AME ‒ ALE 0.05, ALE ‒ ALE 0.34, PME ‒ PME 0.11, PME ‒ PLE 0.12, PLE ‒ PLE 0.60, ALE ‒ PLE 0.06. OAW 0.77, CRW 1.30, OAW / CRW 0.59, MOA 0.34 long, anterior width 0.25, posterior width 0.37, posterior eye row slightly wider than anterior eye row. CH 0.13, CH / AME 1.44. Endites and labium yellowish-brown. Sternum yellow, 1.60 long, 1.29 wide. Leg measurements: Ⅰ 9.10 (2.43, 0.99, 2.28, 1.91, 1.49), II 7.60 (2.24, 0.93, 1.72, 1.45, 1.26), III 6.82 (2.02, 0.89, 1.25, 1.69, 0.97), IV 9.25 (2.51, 0.84, 2.10, 2.76, 1.04); leg formula 4123. Leg spination: femora: Ⅰ d 1 - 1 - 1, p 0 - 0 - 2; II d 1 - 1 - 1, p 0 - 0 - 1; III d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 1 - 1; IV d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 0 - 1. Tibiae: Ⅰ p 0 - 0 - 1, v 1 - 2 - 2; II p 0 - 1 - 1, v 1 - 2 - 1; III d 1 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 2; IV d 1 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 2. Metatarsi: Ⅰ v 2 - 2 - 0; II p 0 - 1 - 0, v 2 - 2 - 0; III d 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2; IV d 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2. Opisthosoma dorsally grey, with a pair of white, irregular stripes and five pairs of white transverse stripes. Palp (Figs 3 A – C, 8 A). RTA hook-shaped in retrolateral view, with rectangular extension at base. Cymbial concavity developed, about half as long as cymbium. Posterior elongation of tegulum extending to median part of tibia. Embolus originating at 7 - to 8 - o’clock-position from tegulum, extending clockwise to 2 - o’clock-position, then curving towards the distal end of cymbium. Conductor fan-shaped. Female paratype (Figs 1 C – D, 2 E, MHBUARA- 2024 - 887 - 8) total length 6.56, prosoma 3.11 long, 2.19 wide; Opisthosoma 3.45 long, 1.92 wide. Eye sizes and interdistances: AME 0.09, ALE 0.13, PME 0.12, PLE 0.12; AME ‒ AME 0.08, AME ‒ ALE 0.05, ALE ‒ ALE 0.35, PME ‒ PME 0.12, PME ‒ PLE 0.13, PLE ‒ PLE 0.59, ALE ‒ PLE 0.07. OAW 0.77, CRW 1.44, OAW / CRW 0.53, MOA 0.35 long, anterior width 0.27, posterior width 0.35. CH 0.12, CH / AME 1.33. Leg measurements: I 7.11 (2.09, 0.94, 1.77, 1.32, 1.00), II 6.52 (1.93, 0.98, 1.45, 1.28, 0.88), III 6.05 (1.78, 0.87, 1.18, 1.44, 0.78), IV 8.44 (2.39, 1.02, 1.78, 2.30, 0.94); leg formula 4123. Labium 0.49 long, 0.41 wide. Sternum 1.52 long, 1.31 wide. Leg spination: femora: Ⅰ d 1 - 1 - 1, p 0 - 0 - 1; II d 1 - 1 - 1, p 0 - 0 - 1; III d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 0 - 1; IV d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 0 - 1. Tibiae: Ⅰ v 1 - 1 - 1; II v 0 - 1 - 1; III: d 1 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 2; IV d 1 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 2. Metatarsi: Ⅰ v 2 - 2 - 0; II v 2 - 1 - 0; III d 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2; IV d 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2. Coloration like male. Epigyne (Fig. 3 D ‒ E). Epigynal plate as long as wide. Posterior triangular atrium width about three times its length in ventral view. Primary spermatheca spherical, separated by about its diameter, secondary spermatheca similar in size to primary spermatheca. Copulatory duct rectangular, bending at median part. Anterior bursae oval. Connecting tubes flattened sac-like, inner margins converging in median section.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C103750A48ABFE19D74EFD47.taxon	distribution	Distribution. China: Yunnan (Fig. 9).	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C104750648ABFD71D74DFD47.taxon	description	Figs 4 – 6, 7 B, 8 B, 9	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C104750648ABFD71D74DFD47.taxon	materials_examined	Type material. Holotype ♂ (MHBUARA- 2024 - 464 - 1), Xizang, Medog County, Drepung Town, 29.1805 ° N, 95.1422 ° E, 1681.4 m elev., 23 June 2024, leg. K. Yu, H. Zhang and Y. Zhang. Paratypes. 4 ♂ 2 ♀ (MHBUARA- 2024 - 464 - 2 ‒ 7), same data as holotype.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C104750648ABFD71D74DFD47.taxon	etymology	Etymology. The specific name is in honor to Prof. Yang Zhong (1964 ‒ 2017), a botanist who made remarkable contributions to the conservation of biodiversity in Xizang.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C104750648ABFD71D74DFD47.taxon	diagnosis	Diagnosis. See S. tongbiguan Zhang & Zhang sp. nov.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C104750648ABFD71D74DFD47.taxon	description	Description. Male holotype (Figs 4 A ‒ B, 5 A ‒ D) total length 6.32; prosoma 3.38 long, 2.43 wide; opisthosoma 2.94 long, 1.58 wide. Eye sizes and interdistances: AME 0.11, ALE 0.13, PME 0.14, PLE 0.13; AME ‒ AME 0.10, AME ‒ ALE 0.04, ALE ‒ ALE 0.34, PME ‒ PME 0.11, PME ‒ PLE 0.11, PLE ‒ PLE 0.60, ALE ‒ PLE 0.07. OAW 0.75, CRW 1.30, OAW / CRW 0.58, MOA 0.35 long, anterior width 0.30, posterior width 0.37. CH 0.12, CH / AME 1.09. Chelicera promargin with three teeth, retromargin with two teeth. Labium 0.45 long, 0.35 wide. Sternum 1.64 long, 1.33 wide. Leg measurements: Ⅰ 9.60 (2.64, 1.26, 2.23, 2.04, 1.43), II 8.42 (2.31, 1.21, 1.86, 1.85, 1.19), III 7.63 (2.24, 1.03, 1.48, 1.97, 0.91), IV 10.61 (3.02, 1.37, 2.21, 2.95, 1.06); leg formula 4123. Leg spination: femora: Ⅰ d 1 - 1 - 1, p 0 - 0 - 2; II d 1 - 1 - 1; III d 1 - 1 - 1, p 0 - 1 - 1, r 1 - 1 - 1; IV d 1 - 1 - 1, p 0 - 1 - 1, r 1 - 1 - 1. Tibiae: Ⅰ p 0 - 1 - 1, v 2 - 2 - 2; II p 0 - 1 - 1, v 2 - 2 - 2; III d 1 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 2; IV d 1 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 2. Metatarsi: Ⅰ p 0 - 1 - 0, v 2 - 2 - 0; II p 0 - 1 - 0, v 2 - 2 - 0; III d 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2; IV d 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2. Opisthosoma dorsally grey, with a pair of white vertical stripes at anterior center, five white stripes medially, and five white horizontal stripes laterally on each side. Palp (Figs 6 A ‒ C, 8 B). Cymbial concavity shallow, about 1 / 3 times as long as cymbium. The posterior elongation of tegulum extending backward to median part of tibia. Embolus whiplike, originating at 8 - o’clock-position, extending clockwise to 1 - o’clock-position, then curving anteriad in ventral view and possessing three small embolic teeth on its distal 2 / 3 part. Conductor fan-shaped. Female paratype (Figs 4 C ‒ D, 5 E, MHBUARA- 2024 - 464 - 6) total length 6.56; prosoma 3.60 long, 2.36 wide, cephalic region elevated. Opisthosoma 4.55 long, 2.66 wide. Eye sizes and interdistances: AME 0.12, ALE 0.15, PME 0.13, PLE 0.13; AME ‒ AME 0.10, AME ‒ ALE 0.04, ALE ‒ ALE 0.38, PME ‒ PME 0.14, PME ‒ PLE 0.14, PLE ‒ PLE 0.66, ALE ‒ PLE 0.06. OAW 0.82, CRW 1.52, OAW / CRW 0.54, MOA 0.37 long, anterior width 0.32, posterior width 0.39. CH 0.09, CH / AME 0.75. Labium 0.52 long, 0.45 wide. Sternum 1.69 long, 1.40 wide. Leg measurements: I 8.01 (2.30, 1.28, 1.86, 1.57, 1.00), II 7.48 (2.19, 1.15, 1.68, 1.50, 0.96), III 6.98 (1.99, 1.03, 1.47, 1.72, 0.77), IV 9.91 (2.86, 1.32, 2.20, 2.58, 0.95); leg formula 4123. Leg spination: femora: Ⅰ d 1 - 1 - 1, p 0 - 0 - 3; II d 1 - 1 - 1, p 0 - 0 - 1; III d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 0 - 1; IV d 1 - 1 - 1, p 0 - 1 - 1, r 0 - 0 - 1. Tibiae: Ⅰ v 0 - 1 - 1; II v 0 - 1 - 1; III d 1 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 2; IV d 1 - 0 - 0, p 1 - 1 - 0, r 1 - 1 - 0, v 2 - 2 - 2. Metatarsi: Ⅰ v 2 - 2 - 0; II v 2 - 2 - 1; III d 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2; IV d 2 - 2 - 2, p 1 - 1 - 1, r 1 - 1 - 1, v 2 - 2 - 2. Prosoma color as in male, opisthosoma dorsally brownish-yellow, without distinct patterns. Epigyne (Fig. 6 D ‒ E). Epigynal plate length about 1.5 times width. Atrium width about four times its length in ventral view. Primary spermatheca spherical, secondary spermatheca ovoid-shaped, smaller than primary spermatheca. Copulatory duct nearly straight. Bursa drop-shaped.	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C39202C104750648ABFD71D74DFD47.taxon	distribution	Distribution. China: Xizang (Fig. 9).	en	Zhang, Haibin, Yu, Kun, Zuo, Anru, Zhang, Feng (2025): Description of a new genus, Septsigilla gen. nov., with two new species from China (Araneae: Miturgidae: Systariinae). Zootaxa 5666 (1): 125-135, DOI: 10.11646/zootaxa.5666.1.7, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
