taxonID	type	description	language	source
03C3AC5D0D1A4847ECF4FC7D29E8940F.taxon	distribution	Distribution: European part: Kola Peninsula (Sokolow, 1940; Shubina & Tsember, 2017); Komi Republic (Shubina & Tsember, 2017).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1A4847ECF4FBE92ACC94F7.taxon	distribution	Distribution: Far East: Sakhalin (Uchida, 1936 b).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1A4847ECF4FB21294496CF.taxon	distribution	Distribution: European part: Leningrad region (Sokolow, 1936; Sokolow & Jankovskaja, 1962); Kola Peninsula (Sokolow, 1940; Shubina & Tsember, 2017), Moscow region (Jankovskaja, 1964); Komi Republic (Shubina & Tsember, 2017); Caucasus: North Ossetia (Sokolow, 1927), Krasnodar region (Sokolow, 1940); Western Siberia: Altai region (Zhukova & Bezmaternykh, 2008; Stolbov, 2008); Khanty-Mansi Autonomous Okrug (Stolbov, 2024); Yamalo-Nenets Autonomous Okrug (Stolbov et al., 2025); Eastern Siberia: Irkutsk region, Republic of Buryatia, Zabaykalsky region (Semenchenko & Matafonov, 2014); Sakha Republic (Semenchenko et al., 2010); Far East: Khabarovsk Krai (Sokolow, 1950); Primorsky Krai, Sakhalin (Sokolow, 1940); Chukotka Peninsula (Wainstein, 1976 b); Magadan Region, Kamchatka (Wainstein, 1981 a); Jewish Autonomous Oblast (Semenchenko, 2005); Amur Region (Semenchenko, 2010).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1A4847ECF4FB21294496CF.taxon	discussion	Notes. 1) A number of old references may probably refer to Sperchon thienemanni Koenike, 1907, which was previously considered a variety of this species. 2) Sperchon glandulosus cubanicus Sokolow, 1940 occurs in the same samples as the nominative subspecies and requires re-examination, since its taxonomic status is not sufficiently defined.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1A4847ECF4F9292B6896B7.taxon	distribution	Distribution: Far East: Magadan region (Tuzovskij, 2008).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1A4847ECF4F9602A6D978B.taxon	distribution	Distribution: Far East: Kuril Islands (Uchida, 1936 a).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1A4847ECF4F9602A6D978B.taxon	discussion	Note: The species was originally assigned to the subgenus Hispidosperchon. However, the description of this species does not indicate the presence of a reticulated structure of the integument (the main feature of Hispidosperchon). Accodring to the description of Uchida, the integument has fine ridges. So very likely, this species belongs to Sperchon s. s. Also, it is impossible to determine the sex of the material from the description, which makes its identification difficult.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FF1D2BD890AF.taxon	distribution	Distribution: Western Siberia: Yamalo-Nenets Autonomous Okrug (Stepanov, 2006; Stolbov e t al., 2025); Far East: Chukotka (Wainstein, 1981 a, b); Amur Region (Semenchenko, 2010).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FE892AAD9133.taxon	distribution	Distribution: European part: Komi Republic (Shubina & Tsember, 2017); Far East: Primorsky Krai, Khabarovsk Krai (Sokolow, 1940; Semenchenko, 2008 a).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FE512B39924F.taxon	distribution	Distribution: Far East: Chukotka (Tuzovskij, 1981)	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FEE52A289187.taxon	distribution	Distribution: Eastern Siberia: Sakha Republic (Semenchenko et al., 2010); Far East: Magadan Region, Kamchatka (Wainstein, 1981 a, b).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FDA92BF092D3.taxon	distribution	Distribution: Eastern Siberia: Sakha Republic (Semenchenko et al., 2010); Far East: Magadan Region, Chukotka, Kamchatka (Tuzovskij, 2008), lower Amur basin (Semenchenko, 2008 a).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FD052C63929B.taxon	distribution	Distribution: Far East: Chukotka, Magadan region, Kamchatka (Tuzovskij, 2021).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FD5D2CD29343.taxon	distribution	Distribution: Far East: Magadan region (Tuzovskij, 2008); Amur region (Semenchenko, 2010).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FC952D31930B.taxon	distribution	Distribution: Far East: Chukotka (Wainstein, 1981 a, b); Khabarovsk Krai (Semenchenko, 2008 c).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FCED2D6293F3.taxon	distribution	Distribution: Far East: Magadan region (Wainstein, 1981 a, b); lower Amur basin (Semenchenko, 2008 a).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FC2428DD9447.taxon	distribution	Distribution: European part: Moscow (Sokolow, 1940); Kola Peninsula, Komi Republic (Shubina & Tsember, 2017).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FB4529B29567.taxon	distribution	Distribution: Western Siberia: Khanty-Mansi Autonomous Okrug (Stolbov, 2024); Far East: Chukotka (Wainstein, 1981 a, b).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FB9128DD9493.taxon	distribution	Distribution: European part: Leningrad region (Sokolow, 1925; 1936), Karelian Isthmus (Sokolow & Jankovskaja, 1962); Komi Republic (Shubina & Tsember, 2017); Sverdlovsk region (Stolbov et al., 2024); Western Siberia: Khanty-Mansi Autonomous Okrug (Stolbov, 2024); Yamalo-Nenets Autonomous Okrug (Stolbov et al., 2025).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FAB12B43952F.taxon	distribution	Distribution: Far East: Magadan region (Wainstein, 1981 a, b).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FA092D5D965F.taxon	distribution	Distribution: European part: Leningrad region (Sokolow, 1936); Komi Republic (Shubina & Tsember, 2017).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4FA092D5D965F.taxon	discussion	Note: Previously considered a form of S. glandulosus, so some older references may refer to this species.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4F9992B7996EB.taxon	distribution	Distribution: European part: Komi Republic (Tuzovskij, 2006; Shubina & Tsember, 2017); Western Siberia: Khanty-Mansi Autonomous Okrug (Stolbov, 2024); Altai Republic (Sokolow, 1940); Eastern Siberia: Sakha Republic (Semenchenko et al., 2010); Far East: Chukotka, Kamchatka, Wrangel Island (Wainstein, 1981 a); Khabarovsk Krai (Semenchenko, 2008 a); Amur Region (Semenchenko, 2010).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4F94D2DB09753.taxon	distribution	Distribution: European part: Leningrad region (Sokolow, 1936); Komi Republic (Shubina & Tsember, 2017).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4F8852A569727.taxon	distribution	Distribution: Western Siberia: Khanty-Mansi Autonomous Okrug.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4F8852A569727.taxon	discussion	Note: The description is given below.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1B4846ECF4F8F12B7197EF.taxon	distribution	Distribution: Far East: Primorsky Krai (Tuzovskij, 2002).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4FE022A569554.taxon	distribution	Distribution: European part: Republic of Karelia (Sokolow, 1930), Kola Peninsula (Sokolow, 1940; Shubina & Tsember, 2017), Leningrad Region (Sokolow, 1936); Tver Region (Thor, 1923; Sokolow, 1940); Kaluga Region (Jankovskaja, 1964); Yaroslavl Region (Zhavoronkova, 2000; Tuzovskij, 1977; 1996; Zhavoronkova et al., 2023); Samara Region (Tuzovskij, 2007); Voronezh Region (Silina et al., 2014); Komi Republic, Arkhangelsk Region (Shubina & Tsember, 2017); Sverdlovsk Region (Klimov et al., 2022; Stolbov et al., 2024): Caucasus: Krasnodar Krai (Jankovskaja, 1959); Western Siberia: Tyumen Region (Stolbov, 2010)	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4FE022A569554.taxon	discussion	Note: Wainstein (1981) suggested that S. clupeifer may have three subspecies: the nominate one, S. clupeifer plumifer Thor, 1902 and S. clupeifer anulatus Wainstein, 1981. He saw differences between these subspecies only in the presence of a chitinous plate around the excretory pore and the degree of plumoseness of the dorsal setae of the legs and their frequency. Since the presence or absence of a chitinous plate around the excretory pore is a species characteristic (Di Sabatino et al. 2010), it is impossible to agree with the subspecific status of S. plumifer (now recognized as a junior synonym of Sperchon hispidus Koenike, 1895). Sperchon clupeifer and S. hispidus are quite clearly distinguished by this feature, as well as by the length / height ratio of P- 4 and by the shape of the dorsal shields in the male (Di Sabatino et al. 2010). The degree of plumoseness of the dorsal setae of the legs also differs, but not always obviously (Di Sabatino et al. 2010). The third subspecies S. clupeifer anulatus also should it be considered a probably full species to the difference from S. clupeifer in the sclerotization of the excretory pore. In the density of the plumose dorsal setae of the legs and sclerotization of the excretory pore, it is closer to S. hispidus. But it differs from it in the degree of plumoseness of these setae. Moreover, it differs from the two species in the proportions of P- 4 - it is narrower in S. clupeifer anulatus (~ 8.0) vs. S. hispidus (4.5 – 5.4) vs. S. clupeifer (5.4 – 6.5). Thus, S. clupeifer anulatus can probably be recognized as a separate species, but the lack of information on males does not provide complete certainty. A re-examination of the type material of S. clupeifer anulatus is required and more material is needed, including males.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4FA862BC6951C.taxon	distribution	Distribution: European part: Komi Republic (Shubina & Tsember, 2017).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4FADE29BE95A8.taxon	distribution	Distribution: European part: Komi Republic, Kola Peninsula (Shubina & Tsember, 2017).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4FADE29BE95A8.taxon	discussion	Note: According to current data, it belongs to a complex of closely related species, reliably distinguishable only by males (Di Sabatino et al. 2010). Therefore, we are not sure that the identification of Shubina & Tsember (2017) refers to this species.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4F9822B2F96E4.taxon	distribution	Distribution: European part: Kola Peninsula (Shadin, 1940); Arkhangelsk region, Komi Republic (Shubina & Tsember, 2017); Sverdlovsk region (Klimov et al., 2022; Stolbov et al., 2024); Caucasus: North Ossetia (Sokolow, 1927); Far East: Primorsky Krai (Semenchenko, 2008 b).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4F9362C4796AC.taxon	distribution	Distribution: Far East: Chukotka, Kamchatka, Wrangel Island (Wainstein, 1981 a, b).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4F88E2C1D9714.taxon	distribution	Distribution: Far East: Khabarovsk Krai (Tuzovskij & Semenchenko, 2009).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4F8C62B5097A0.taxon	distribution	Distribution: European part: Leningrad region (Sokolow, 1936); Kola Peninsula (Sokolow, 1940; Shubina & Tsember, 2017); Tver region (Thor, 1923); Kaluga region (Jankovskaja, 1964); Samara region (Tuzovskij, 2007); Arkhangelsk region, Komi Republic (Shubina & Tsember, 2017)	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4FF1D2A5690AF.taxon	distribution	Distribution: Western Siberia: Khanty-Mansi Autonomous Okrug.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1C4841ECF4FF1D2A5690AF.taxon	discussion	Note: The description is given below.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4840ECF4FF1D28DD914B.taxon	distribution	Distribution: European part: Leningrad region (Sokolow, 1936); Lipetsk region (Silina, 2015); Yaroslavl region (this study); Komi Republic (Shubina & Tsember, 2017); Western Siberia: Sverdlovsk region (Klimov et al., 2022).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4840ECF4FE3D2A16926B.taxon	distribution	Distribution: European part: Komi Republic (Tuzovskij, 2013); Eastern Siberia: Irkutsk region (Tuzovskij, 2013); Far East: Kamchatka (Wainstein, 1976 a, 1981 a), Magadan region, Primorsky Krai (Wainstein, 1981 a); lower Amur basin (Semenchenko, 2008 a).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4840ECF4FDCD2C8C929B.taxon	distribution	Distribution: European part: Komi Republic (Shubina & Tsember, 2017); Sverdlovsk region (Stolbov et al., 2024); Western Siberia: Khanty-Mansi Autonomous Okrug (Stolbov, 2024); Eastern Siberia: Irkutsk region (Scheffler, 1972); Far East: Chukotka (Wainstein, 1981 a); Amur region (Semenchenko, 2010)	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4840ECF4FD5C29D1936F.taxon	distribution	Distribution: Far East: Magadan Region (Tuzovskij, 1982); Khabarovsk Territory (Semenchenko, 2008 c), Amur Region (Semenchenko, 2010).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4842ECF4FCF52C809186.taxon	description	(Figs 2 – 4, 11 A, B)	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4842ECF4FCF52C809186.taxon	materials_examined	Material examined. Holotype: female, Russia, Nether-polar Urals, Khanty-Mansi Autonomous Okrug, Beryozovsky district, Shaitanka River, 64 ° 34 ' 05.9 " N 59 ° 40 ' 50.6 " E, 407 m a. s. l., among moss on stones, 25.07.2023, leg. V. Stolbov (Fig. 1 A).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4842ECF4FCF52C809186.taxon	diagnosis	Diagnosis. Integument dorsally with flat papillae separated by short folds, ventrally with undulating folds. Glandularia platelets of dorsum not enlarged, without muscle attachment platelets (except Dl- 2). Cx-I + II not fused, Cx-III with a small medial glandular opening. Venter without muscle attachment platelets. Acetabula 1 and 2 rounded, the small, distance between these acetabula is 1 – 2 times greater than their length. Excretory pore surrounded bу а sclerotized ring, located distinctly closer to Vgl- 3. P- 2 not shortened or widened, with a pointed distoventral projection and two distal short setae and one long spiniform basal seta. P- 3 without ventral setae. Ventral peg-like seta on P- 4 small, far distanced rom each other. P- 5 twice as long as it is wide. IV-L- 3 – 5 with few slightly serrated dorsal setae.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4842ECF4FCF52C809186.taxon	description	Description Female. Integument dorsally with flat, papillae separated by short folds, which are located predominantly in the transverse direction of the idiosoma; ventrally with undulating fine folds, which are located predominantly in the transverse direction of the idiosoma. Dorsoglandularia and lateroglandularia are located on medium-sized (diameter 80 – 100) chitinous platelets with irregular shaped edges. Seta Dgl- 1 slightly spiniform, short, smooth. Preocularia setae are located on small chitinous platelets (diameter 35), postocularia — larger (diameter 90, approximately the same diameter as the ocular plate). Frontale pore (Fr) present. Dorsocentralia muscle attachment plates (Dc) absent, only one pair small (25) dorsolateralia plates Dl- 2 present. Five pairs of lyriform organs (i 1 - 5) present. Cx-I + II medially close to each other and in close contact, but not fused. Cx-III with a small medial glandular opening (Cxgl- 4). Coxoglandulare plates 2 (Cxgl- 2) and lateroglandularia 3 (Lgl- 3) medium-sized (70), ventroglandularia 1 (Vgl- 1) small (35), other ventroglandularia platelets (Vgl- 2 - 4) larger (80 - 90). Excretory pore surrounded bу а sclerotized ring (103), located distinctly closer to Vgl- 3 than to Vgl- 1. Genital plates not enlarged (225) with 17 – 18 medial and 6 – 7 lateral setae. Acetabula 1 and 2 (Ac- 1 - 2) rounded, small (35 – 40), distance between Ac- 1 and Ac- 2 is 1 – 2 times greater than their length. Acetabulum 3 round, larger (47). Postgenital and ventral sclerites (Vc) absent. Gnathosoma with rostrum shorter than gnathosomal base. Palp: P- 1 with one small spiniform dorsal setae; P- 2 not shortened or widened, with nine spiniform dorsal setae, a pointed distoventral projection with two distal short setae and one long (longer than distoventral projection) spiniform basal seta; P- 3 with five spiniform and two short filiform dorsal setae, ventral margin without setae and denticulation; P- 4 longer than P- 3, narrow (length / height 6.2), with two slightly developed ventral tubercles, bearing peg-like setae (proximal seta slightly larger that distal), proximal tubercle larger than distal one, proximal peg-like seta is located almost in the middle of P- 4, the distal one is closer to the distal margin of P- 4 (length of P- 5 approximately equal to distance from distal ventral peg-like seta on P- 4 to distal edge of P- 4), the distance between peg-like setae is more than twice as long as the height of P- 4, between peg-like setae two short filiform setae, distal spine seta pointed, slightly widened towards the top; P- 5 twice as long as wide, with large distal spines, two approximately the same size as ventral spines, one small dorsal spines before solenidion, one bifurcated seta (branches are equivalent) and four filiform setae (distance between bases of distal and ventral filiform setae very short than length of ventral setae). Leg segments slender, IV-L- 3 - 5 with few slightly serrated dorsal setae, other setae spiniform or short filiform. Ambulacrum with claw blade well protruding bearing a long dorsal and a shorter ventral clawlet. Male unknown. Measurements. Idiosoma L 1430, W 1100; distance between anterior end of Cx-I and posterior end of Cx-IV, 627; Cx-III W, 790. Palp dL / H ratio: P- 1 52 / 58, 0.89; P- 2 153 / 111, 1.37; P- 3 176 / 100, 1.76; P- 4 270 / 44, 6.1; P- 5 44 / 22, 2.0. Gnathosoma L 298; chelicera L 285. Leg segments dL: I-L- 2 – 6: 136, 175, 267, 283, 231; IV-L: 154, 201, 191, 405, 382, 317.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4842ECF4FCF52C809186.taxon	etymology	Etymology. The species is named after of the late Dr. Olga D. Zhavoronkova for her contributions to the study of water mites.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1D4842ECF4FCF52C809186.taxon	discussion	Remarks. The new species is most similar to Sperchon fluviatilis in the following characters: integument with undulating fine ridges; ventral setae P- 3 absent; ventral spiniform seta P- 2 present; distance between bases of distal and ventral filiform setae very short than length of ventral setae. However, the new species is noticeably different from S. fluviatilis by the following characters: length of P- 5 approximately equal to distance from distal ventral peg-like seta on P- 4 to distal edge of P- 4 (vs. length of P- 5 shorter than distance from distal ventral peg-like seta on P- 4 to distal edge of P- 4 in S. fluviatilis); acetabula not large, distance between Ac- 1 and Ac- 2 is 1 - 2 times greater than their length (vs. acetabula large, distance between Ac- 1 and Ac- 2 is 1 / 2 times less than their length in S. fluviatilis); folds of skin covers on back and behind genital opening are arranged in transverse rows (vs. arranged in irregular rows in S. fluviatilis); anus with sclerotized plate (vs. without sclerotized plate in S. fluviatilis).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1F4849ECF4F9EF2A0791FB.taxon	description	(Figs 5 – 7, 11 C, D)	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1F4849ECF4F9EF2A0791FB.taxon	materials_examined	Material examined. Holotype: male, Russia, Nether-polar Urals, Khanty-Mansi Autonomous Okrug, Beryozovsky district, Kobyla-Yu River, 64 ° 30 ' 24.1 " N 59 ° 33 ' 22.8 " E, 571 m a. s. l., 23.07.2023, leg. V. Stolbov (Fig. 1 B).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1F4849ECF4F9EF2A0791FB.taxon	diagnosis	Diagnosis. Integument dorsally with numerous long folds, between which are located few rounded or elongated papillae, ventrally with undulating fine folds. Glandularia platelets of dorsum not enlarged, with small muscle attachment platelets (Dc- 2 - 4 and Dl- 1 - 4), Dl- 1 consosting of two small platelets, Dl- 2 consisting of three small platelets. Cx-I + II not fused, Cx-III with a small medial glandular opening. Ventral with muscle attachment platelets present. Acetabula 1 and 2 elongated, distance between these acetabula is one times greater than their length. Excretory pore surrounded bу а sclerotized ring, located distinctly closer to Vgl- 3 than to Vgl- 1. P- 2 not shortened or widened, with a pointed distoventral projection with three distal short setae and one short spiniform basal seta. P- 3 without ventral setae. Ventral peg-like seta on P- 4 very small, far apart. P- 5 more than twice as long as wide. Few dorsal setae of IV-L- 3 - 5 not plumose.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1F4849ECF4F9EF2A0791FB.taxon	description	Description Male. Integument dorsally with numerous long folds, between which are located rare rounded or elongated papillae; ventrally with undulating fine folds. Dorsoglandularia and lateroglandularia are located on small (diameter 55 - 70) chitinous platelets with irregular shaped edges. Seta Dgl- 1 slightly spiniform, very short, smooth. Preocularia are located on small chitinous platelets (diameter 30), postocularia — larger (diameter 82 - 90, approximately the same diameter as the ocular plate). Frontale pore (Fr) present. Dorsocentralia and dorsolateralia muscle attachment plates (Dc- 2 - 4 and Dl- 1 - 4) present, Dc- 3 larger (110 - 125 / 60 - 70) than others (20 - 50), elongated with a constriction in the middle, Dl- 1 consisting of two small platelts, Dl- 2 consisting of three small platelets. Five pairs lyriform organs (i 1 - 5) present. Cx-I + II medially close to each other and in close contact, but not fused. Cx-III with a small medial glandular opening (Cxgl- 4). Coxoglandularia platelets 2 (Cxgl- 2), lateroglandularia 3 (Lgl- 3) and ventroglandularia 2 and 3 (Vgl- 2 - 3) platelets small (40 - 50), ventroglandularia 1 (Vgl- 1) a little smaller (30). Ventralia muscle attachment plates (V- 1 - 3) present, small (20 - 30). Excretory pore surrounded bу а sclerotized ring (60), located distinctly closer to Vgl- 3 than to Vgl- 1. Genital plates not enlarged (251) with 14 medial and 5 lateral setae. Acetabula 1 and 2 (Ac- 1 - 2) slightly elongated, small (55 - 65 / 30), distance between Ac- 1 and Ac- 2 is one times greater than their length. Acetabulum 3 round, larger (50). Postgenital platelet present (50). Pregenetal platelets small (25). Gnathosoma with rostrum shorter than gnathosomal base. Palp: P- 1 with one small spiniform dorsal setae; P- 2 not shortened or widened, with 12 spiniform dorsal setae, a pointed distoventral projection with three distal short setae and one short (not longer than distoventral projection) spiniform basal seta; P- 3 with three spiniform and two short filiform dorsal setae, ventral margin without setae and denticulation; P- 4 longer than P- 3, narrow (length / height 6.3), with two slightly developed ventral tubercles, bearing minute peg-like setae, proximal tubercle slightly larger than distal one, proximal peg-like seta is located slightly distal to middle of P- 4, distal one is closer to the distal margin of P- 4 (length of P- 5 noticeably longer than distance from distal ventral peg-like seta on P- 4 to distal edge of P- 4), the distance between peg-like setae is twice as long as the height of P- 4, between peg-like setae two minute filiform setae, distal spine seta rounded at top; P- 5 more than twice as long as wide, with large distal spines, two ventral spines of approximately the same size, one small dorsal spines before solenidion, one bifurcated seta (branches are equivalent) and four filiform setae (distance between bases of distal and ventral filiform setae slightly larger than length of ventral setae). Leg segments slender, IV-L- 3 - 5 with few thin dorsal setae, not plumose, other setae spiniform or short filiform. Ambulacrum with claw blade well protruding bearing a long dorsal and a shorter ventral clawlet. Female unknown. Measurements. Idiosoma L 1270, W 1010; distance between anterior end of Cx-I and posterior end of Cx-IV, 590; Cx-III W, 832. Palp dL / H ratio: P- 1 40 / 64, 0.62; P- 2 153 / 117, 1.3; P- 3 189 / 99, 1.9; P- 4 313 / 50, 6.3; P- 5 77 / 26, 2.9. Gnathosoma L 276; chelicera L 290. Leg segments dL: I-L- 2 – 6: 128, 191, 325, 351, 299; IV-L: 160, 177, 208, 390, 385, 320.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1F4849ECF4F9EF2A0791FB.taxon	etymology	Etymology. The species is named after the Ural Mountains where it was found.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D1F4849ECF4F9EF2A0791FB.taxon	discussion	Remarks. The new species is most similar to Sperchon questor in the following characters: integument with papillae and undulating fine folds; acetabula Ac- 1 and Ac- 2 slightly elongated; ventral setae P- 3 absent; ventral spiniform seta P- 2 present; length of P- 5 is noticeably less than distance from distal ventral peg-like seta on P- 4 to distal edge of P- 4; distance between bases of distal and ventral filiform setae longer than length of ventral setae. However, the new species is noticeably different from S. questor by the following characters: P- 4 not longer than P- 2 + 3 (vs. P- 4 longer than P- 2 + 3 in S. questor); ventrodistal process on P- 2 with 3 setae, not bifurcated (vs. ventrodistal process on P- 2 with two setae, clearly bifurcated in S. questor); dorsal integument with few papillae (vs. papillae numerous in S. questor).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D144849ECF4FE792DB39327.taxon	description	(Figs 8 – 10, 11 E, F)	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D144849ECF4FE792DB39327.taxon	materials_examined	Material examined. Three females, Russia, Yaroslavl region, Kutma River, 24.11.2024, leg. V. Kolesnikov; one female, Russia, Sverdlovsk Region, Balda River, 04.10.2020, leg. V. Stolbov (COX 1 sequence: OL 343041).	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D144849ECF4FE792DB39327.taxon	diagnosis	Diagnosis. Integument reticulated, with numerous folds. Glandularia platelets of dorsum not enlarged, dorsum with small muscle attachment plates (Dc- 2 - 4 and Dl- 1 - 4). Cx-I + II separated, Cx-III without glandular opening, Cxgl- 4 is the only seta between Cx-IV and genital plates. Venter with muscle attachment platelets. Acetabula 1 and 2 elongated, close to each other. Excretory pore without sclerotized ring, located close to Vgl- 3. P- 2 not shortened or widened, with a pointed distoventral projection with three distal short setae, ventral spiniform seta absent. P- 3 with four spiniform ventral setae. Ventral peg-like seta on P- 4 small. P- 5 short. IV-L- 3 - 5 with few, simple dorsal setae.	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
03C3AC5D0D084854ECF4FE3D2BB197B8.taxon	discussion	Notes * Due to limited data from the description, characters in the key based on the available information. The species requires re-examination. ** Since it is impossible to reliably determine to which species of the denticulatus species group the specimens from Russia belong (specimens and their images are missing), we use in the key only the definition of this group. *** The male of Sperchon clupeifer anulatus Wainstein, 1981 is unknown, so it is impossible to compare the dorsal plates. It is similar to S. hispidus in the presence of a sclerotized ring around the anus and dense dorsal setae on tibia IV, but in the less plumose dorsal setae on tibia IV it is more reminiscent of S. clupeifer. This subspecies has the narrowest P- 4 (~ 8.0). It is possible that this is a separate species	en	Kolesnikov, Vasiliy B., Stolbov, Vitaly A. (2025): Review of the genus Sperchon Kramer, 1877 (Acari, Hydrachnidia: Sperchontidae) from Russia. Zootaxa 5666 (1): 53-73, DOI: 10.11646/zootaxa.5666.1.2, URL: http://dx.doi.org/10.11646/zootaxa.5360.3.8
