taxonID	type	description	language	source
03C087AF2F37FFAEC495FD36FC306F53.taxon	materials_examined	Material examined. IRAQ: 1 ♀: S Rawandoz, Akoian Valley, 36 ° 30´N 44 ° 36´E. ~ 1400 m a. s. l., pitfall trap. 04.2018. leg. Ch. Reuter (cF); AFGHANISTAN: 1 ♂: ‘ ♂ ’, ‘ J, Klapperich Walang, 2750 m Salangtal, 15.11.52 ’, ‘ Hindukusch O. Afghanistan’, ‘ ex. coll. Scheerpeltz’, ‘ TYPUS Lathrimaeum afghanicum O. Scheerpeltz’, ‘ Lathrimaeum afghanicum n. sp. ’ (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F37FFAEC495FD36FC306F53.taxon	discussion	Remarks. Anthobium fusculum is widely distributed in the Palaearctic region and known from Europe to Caucasus, Middle East and Central Asia (Shavrin & Smetana 2018). Shavrin & Smetana (2018) and Shavrin (2023, 2024 a) listed “ Lathrimaeum kashmiricum Coiffait, 1982: 150 ” as a synonym of A. fusculum. In fact, the name “ kashmiricum ” was misspelled with “ cachemiricum ”. Lathrimaeum cachemiricum Coiffait, 1982 was originally described from “ Inde, Cachemire, Zagipal, près Pahalgam … ” (Coiffait 1982: 150, see also Shavrin 2013). Other species, Anthobium kashmiricum (Cameron, 1941), originally described as Lathrimaeum (Prionothorax) kashmiricum from “ Kashmir: Gulmarg … ” (Cameron 1941: 58) is a valid species of the reflexum group (Shavrin & Smetana 2019). It is here recorded from Iraq and Afghanistan for the first time.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F37FFACC495FA7BFAA269F4.taxon	description	(Figs 1, 4 – 5)	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F37FFACC495FA7BFAA269F4.taxon	materials_examined	Type material examined. Holotype (dissected) ♂ (Fig. 1): ‘ ♂ ’ <printed>, ’ 45. ’ <handwritten in black>, ‘ Oesterreichische | Karakorum Expedition | leg. Dr. E. PIFFL’ <printed>, ‘ Haramosch – Massiv [printed] | Sabil [?], Quelmoos | Gesiebe’ <handwritten>, ‘ Lathrimaeum | karakorumshane- | nsis, spec. | (prope | rugosum | Champ.) ’ <handwritten>, ‘ HOLOTYPE | Anthobium | scheerpeltzi sp. nov. | Shavrin A. V. 2025 ’ <red, printed> (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F37FFACC495FA7BFAA269F4.taxon	description	Description. Measurements: HW: 0.67; HL: 0.42; OL: 0.17; LT: 0.05; AL: 1.20; PL: 0.55; PWmax: 0.97; PWmin: 0.85; ESL: 1.30; EW: 1.22; MTbL: 0.65; MTrL (holotype): 0.30 (MTrL 1 – 4: 0.17; MTrL 5: 0.13); AW: 1.21; AedL: 0.43; BL: 3.20. Forebody moderately wide and convex (Fig. 1). Head and abdomen reddish brown; pronotum, and elytra yellow-brown (lateral parts of pronotum slightly paler); antennomeres 5 – 11 brown; mouthparts, antennomeres 1 – 4, legs and paratergites yellowish. Head with distinct microsculpture: fine and transverse in apical part of clypeus, coarser in middle portion between apical margins of eyes, finer and irregular in middle between level of basal third of eyes, infraorbital portions with dense and coarse isodiametric sculpture; neck with dense and fine isodiametric microreticulation; medioapical and mediobasal parts of pronotum with fine transverse meshes; scutellum and elytra without microsculpture; abdominal tergites with dense sculpture: isodiametric on abdominal tergites IV – V and transverse on tergites VI – VIII. Middle portion of head with dense and fine punctation, with interspaces between punctures between eyes about as long as one-two nearest punctures, punctation on infraorbital portion denser and coarser; visible part of neck without punctures; pronotum with moderately fine and dense punctation, distinctly sparser than that in middle part of head, finer in medioapical part, larger and sparser in lateral portion, and sparser in mediobasal third; scutellum with several fine punctures; punctation of elytra about as that on pronotum, denser and coarser around scutellum, finer and sparser along suture, larger and deeper in medioapical portion; abdominal tergites without visible punctation. Head 1.5 times as broad as long, with slightly elevated middle part and infraorbital portions, with wide semicircular impression in front of ocelli, and deep and wide anteriomedian depressions; anteocellar foveae missing; each anterior portion between antennal insertion and anterior margin of eye with moderately deep semicircular notch; postocular ridge acute, with distance between basal margin of eye and ridge about as long as three nearest ommatidia. Median surface of head with distinct elevations: irregular, transverse or diagonal in middle, and coarser and diagonal on infraorbital portions. Eyes large, strongly convex. Ocelli large, located slightly below level of posterior margin of eyes, distance between ocelli 1.4 times as long as distance between ocellus and posterior margin of eye. Apical maxillary palpomere about two and a half times as long as preceding segment, from basal part gradually narrowed toward subacute apex. Antennae reaching basal part of elytra when reclined, with distinctly elongate antennomeres 1 – 8 and slightly elongate 9 – 10; basal antennomere wide, about two and a half times as long as broad, 2 distinctly shorter and narrower than basal antennomere, 3 slightly longer and narrower than 2, 4 – 5 shorter than 3, 6 – 7 shorter than 5, 8 shorter than 7, 9 – 10 distinctly shorter than 8, apical antennomere 1.3 times as long as 10, from middle gradually narrowed toward subacute apex. Pronotum 1.7 times as broad as long, 1.4 times as broad as head, widest in middle, more narrowed posteriad than anteriad; apical portions widely rounded, distinctly protruded apicad; laterobasal margins widely concave in front of obtuse hind angles; middle portion evenly elevated, with fine and wide semicircular depression in mediobasal third; lateral edges with irregular acute crenulation; lateral portions widely impressed and slightly explanate, with deep moderately large oval pits slightly above middle. Elytra slightly longer than broad, gradually broadened posteriad, distinctly more than twice as long as pronotum, reaching apical margin of abdominal tergite IV; middle surface of each elytron with longitudinal elevations between punctures; lateral portions moderately narrow and indistinctly explanate; lateral edges with fine crenulation, indistinct in middle. Hind wings fully developed. Legs moderately long and slender, with dense and long setae on outer and inner margins of tibiae; metatarsi twice shorter than metatibia. Abdomen convex, slightly narrower than elytra, with a pair of small oval tomentose spots in middle of abdominal tergite V, with wide intersegmental membranes between abdominal tergites IV – V and narrow palisade fringe on margin of tergite VII. Male. Apical margin of abdominal tergite VIII straight. Apical margin of abdominal sternite VIII widely sinuate. Aedeagus moderately narrow, with wide median lobe slightly narrowed toward truncate apex; parameres wide, exceeding apex of median lobe, each broadened in preapical part and from apical third gradually narrowed apically, with three short apical setae; internal sac short and weakly sclerotized (Fig. 4). Lateral aspect of the aedeagus as in Fig. 5. Female unknown. Comparative notes. Regarding the general shape of the body and the presence of distinct crenulation on lateral edges of the pronotum, A. scheerpeltzi sp. nov. belongs to the fusculum group (Shavrin & Smetana 2018). Based on the general shape of the body and the morphology of the aedeagus, it is similar to A. wittmeri Shavrin, 2023, recently described from Kashmir (Shavrin 2023), from which it can be distinguished by the longer antennomeres 9 – 10 and elytra, coarser sculpture of the infraorbital portions, the lack of anteocellar foveae, and narrower apical parts of the median lobe and parameres.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F37FFACC495FA7BFAA269F4.taxon	biology_ecology	Natural history. The holotype was collected by sifting moss near a spring.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F37FFACC495FA7BFAA269F4.taxon	distribution	Distribution. Anthobium scheerpeltzi sp. nov. is known only from the type locality in Haramosh Mts, Pakistan.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F37FFACC495FA7BFAA269F4.taxon	etymology	Etymology. Patronymic; the species is dedicated to the memory of Otto Scheerpeltz (1888 – 1975).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F35FFACC495FBD6FD526F27.taxon	materials_examined	Material examined. 3 ♀♀: TAJIKISTAN: ‘ Hissar, Boch. Coll Hauser’, ‘ Collect. Hauser’ (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F35FFACC495FBD6FD526F27.taxon	discussion	Remarks. Geodromicus convexicollis is widely distributed in the Middle Asia and known from Kazakhstan, Uzbekistan, Kyrgyzstan, Tajikistan and Afghanistan.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F35FFABC495FA67FADE6AA8.taxon	materials_examined	Material examined. AFGHANISTAN: 8 ♂♂, 2 ♀♀ (one male and one female with additional red label: ‘ TYPUS Geodromicus afghanicus O. Scheerpeltz’): ‘ J. Klapperich Pagmangebirge 2300 m, 30.5.52 O. Afghanistan’, ‘ ex coll. Scheerpeltz’, ‘ COTYPUS Geodromicus afghanicus O. Scheerpeltz’ (NMW); 2 ♂♂: ‘ J. Klapperich Pagmangebirge 2400 m, 6.7.52 O. Afghanistan’, ‘ ex, coll. Scheerpeltz’, ‘ COTYPUS Geodromicus afghanicus O. Scheerpeltz’ (NMW); 1 ♂: ‘ J. Klapperich Sarekanda, 3600 m 23.7.53, Gebirge’, ‘ Badakschan NO - Afghanistan’, ‘ ex coll. Scheerpeltz’, ‘ COTYPUS Geodromicus afghanicus O. Scheerpeltz’ (NMW); 1 ♂, 1 ♀: ‘ J. Klapperich Qulatak, 1950 m Salangtal, 9.10.52 ’, ‘ Hindukush O. Afghanistan’, ‘ ex. coll. Scheerpeltz’, ‘ TYPUS Geodromicus cordicollis O. Scheerpeltz’ (NMW); 1 ♂: same data as the previous except one red label: ‘ COTYPUS Geodromicus cordicollis O. Scheerpeltz’ (cSh); 1 ♀: ‘ J. Klapperich Qulatak, 1950 m Salangtal, 9.10.52 ’, ‘ Hindukush O. Afghanistan’, ‘ ex coll. Scheerpeltz’, ‘ TYPUS Geodromicus salangensis O. Scheerpeltz’, ‘ Geodromicus salangensis n. sp. ’ (NMW); 2 ♀♀: ‘ Oesterreichische Karakorum Expedition leg. Dr. E. PIFFL’, ‘ Haramosch-Massiv Gumari’ (cSh, NMW); INDIA: KASHMIR: 1 ♂: Tangmarg, Pir Panjal Mt. 2600 m a. s. l. 21 – 25.05.1976. J. Martens & W. Schawaller leg. (SMNS).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F35FFABC495FA67FADE6AA8.taxon	discussion	Remarks. Geodromicus kashmirensis is known from Afghanistan, Pakistan and India (Ladakh, Kashmir).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFABC495FEE2FB3D682A.taxon	materials_examined	Material examined. KYRGYZSTAN: NARYN: 1 ♂: N Kalmak-Ashuu (pass). 3400 m a. s. l. 31.07.2009. W. Schawaller leg. (SMNS); OSH: 1 ♀: Lailak District, Sarken-Gush rivers. 2100 m a. s. l. 09.06.1996. A. Puchkov leg. (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFABC495FEE2FB3D682A.taxon	discussion	Remarks. Geodromicus lebedevianus is known from several localities in Kyrgyzstan.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFABC495FD63FE0969A0.taxon	materials_examined	Material examined. CHINA: GANSU: 1 ♂: Lianhuashan. 2800 – 2900 m a. s. l. 23.07. – 01.08.2007. J. Martens leg. (SMNS); HUBEI: 3 ♀♀: Dashenongjia Mts., 31 ° 5´N 110 ° 3´E. 2100 – 2900 m a. s. l. 10 – 14.08.2002. J. Turna leg. (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFABC495FD63FE0969A0.taxon	discussion	Remarks. Geodromicus paramerus is widely distributed in the eastern Palaearctic region and known from eastern Siberia and China.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFABC495FBEAFD606F22.taxon	materials_examined	Material examined. CHINA: HEBEI: 1 ♂: Mongol pass Chengde-Chifeng, 41 ° 6´N 118 ° 2´E. 30 – 31.05.2002. J. Turna leg. (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFABC495FBEAFD606F22.taxon	discussion	Remarks. Geodromicus sibiricus is widely distributed in the eastern Palaearctic region and known from Siberia, Far Eastern Russia, north-eastern China and Korea.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFABC495FA6BFDD66C84.taxon	materials_examined	Material examined. CHINA: SHAANXI: 1 ♂: 15 km S Shou-Man vill. Daba Shan, 32 ° 08´N 108 ° 37´E. 1800 m a. s. l. 25.05 – 14.06.2000 (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFABC495FA6BFDD66C84.taxon	discussion	Remarks. Geodromicus szechwanensis is known from several provinces of China (Henan, Hubei, Ningxia Autonomous Region, Sichuan, Shaanxi).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFAAC495F90EFDD56BEC.taxon	materials_examined	Material examined. PAKISTAN: 1 ♂, 2 ♀♀: ‘ Oesterreichische Karakorum Expedition leg. Dr. E. Piffl’, ‘ Haramosh-Massiv Iskere, Bachufer’ (NMW); INDIA: UTTARAKHAND: 1 ♀: ca. 30 km N Bageshwar, SE Dakhuri vill. 2600 – 2800 m a. s. l. 25 – 26.06.2003. Z. Kejval & M. Trýzna leg. (NMW); NEPAL: KARNALI: 1 ♂: Humla District, Ghatte, Yanchu Khola, 29 ° 54´24 ´´ N 81 ° 55´08 ´´ E. 2920 m a. s. l. 16 – 18.06.2022. A. Hetzel leg. (cF).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F32FFAAC495F90EFDD56BEC.taxon	discussion	Remarks. Hygrodromicus coriaceus is widely distributed in the Himalayan Region, and known from Kashmir to Nepal. Shavrin (2024 b) erroneosly provided Geodromicus palpalis Coiffait, 1977 as a member of the genus Asiageodromicus Shavrin, 2024, and provided a new combination. This is an unfortunate mistake! In fact, G. palpalis belongs to the genus Hygrodromicus Tronquet, 1981, which was established by me earlier (Shavrin 2024 c), and it was synonymized with H. coriaceus based on the study of the type and an additional material. It is here recorded from Pakistan for the first time.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F33FFAAC495FDA6FD2A6962.taxon	discussion	Remarks. Hygrodromicus ledouxi was originally described from “ Cachemire, Himachal Pradesh, Mari … ”. Recently, Shavrin (2024 b) synonymized Hygrogeus ledouxi Coiffait, 1983, originally described from “ Cachemire, Pahalgam, Sheshnag … ” (page 282), with the Himalayan Hygrodromicus coriaceus (Cameron, 1924). Thus, the former name (junior synonym) treated as a junior secondary homonym by the first revisor action, and H. ledouxi Coiffait, 1983: 285 should be selected here as a valid species.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F33FFAAC495FC2DFF496FB2.taxon	materials_examined	Material examined. CHINA: SICHUAN: 3 ♂♂, 4 ♀♀: 30 km W Nanping, Jiuzhaigou. 3100 m a. s. l. 13 – 15.06.1992. J. Turna leg. (NMW); 1 ♂: Gongga Shan, Hailuogou, above Camp 3, 29 ° 35´N 102 ° 00´E. 2800 – 3300 m a. s. l. 06 – 08.07.1998. J. Farkač leg. (NHMB); 6 ♂♂, 7 ♀♀: Jiuzhaigou. 3100 m a. s. l. 13 – 15.06.1992. M. Bok leg. (SMNS); YUNNAN: 1 ♀: Yulong Mts., 27.01 N 100.12 E. 3200 m a. s. l. 24 – 26.09.1993. Bolm leg. (NHMB); 1 ♂, 1 ♀: above Dali. 2000 – 2200 m a. s. l. 04 – 17.04.1999. W. Schawaller leg. (SMNS); 7 ♂♂, 5 ♀♀: same data. 2500 – 2700 m a. s. l. 08 – 18.04.1999. (SMNS).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F33FFAAC495FC2DFF496FB2.taxon	discussion	Remarks. Hygrodromicus maderi is widely distributed in China (Chongqing, Qinghai, Hubei, Sichuan, Yunnan).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F33FFAAC495FA16FD2B6D50.taxon	materials_examined	Material examined. INDIA: KASHMIR: 1 ♀: Tangmarg, Pir Panjal Gebirge. 2600 m a. s. l. 21 – 25.05.1976. J. Martens & W. Schawaller leg. (SMNS); NEPAL: BAGMATI: 3 ♀♀: Dolakha District, SW Kalinchok Mt. 3100 m. 19 – 23.04.1995. J. Martens & W. Schawaller leg. (SMNS).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F33FFAAC495FA16FD2B6D50.taxon	discussion	Remarks. Lesteva championi is widely distributed in the Himalayan Region, and known from India (Kashmir, Uttaranchal Pradesh, Uttarakhand) and Nepal.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F33FFA9C495F87BFB896B04.taxon	materials_examined	Material examined. NEPAL: BAGMATI: 1 ♀: Kavre [Kavrepalanchok] District, 25 km E Kathmandu, 7 km NNW Dhulikel, 27 ° 41´13.48 ´´ N 85 ° 32´14.98 ´´ E. 1900 m a. s. l. 19.11.2005. M. Jäch leg. (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F33FFA9C495F87BFB896B04.taxon	discussion	Remarks. Lesteva jaegeri was originally described from several localities in Bhojpur and Udayapur districts of eastern Nepal. It is here recorded from Bagmati and central Nepal for the first time.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA9C495FE8EFAA8694E.taxon	materials_examined	Material examined. AFGHANISTAN: 1 ♂, 1 ♀: ‘ J. Klapperich Kamdesch 2000 m Nuristan 27.4.53 Afghanistan’, ‘ ex coll. Scheerpeltz’, ‘ TYPUS Mannerheimia afghanica O. Scheerpeltz’ (NMW); 2 ♂♂, 2 ♀♀: same data as the previous except red label: ‘ COTYPUS Mannerheimia afghanica O. Scheerpeltz’ (NMW); 12 ♂♂, 6 ♀♀: ‘ J. Klapperich Kamdesch 2200 m Nuristan 28.4.53 Afghanistan’, ‘ ex coll. Scheerpeltz’, ‘ COTYPUS Mannerheimia afghanica O. Scheerpeltz’ (cSh, NMW); 1 ♂: ‘ J. Klapperich Achmede Dewane’, ‘ 2800 m, Bashgultal, Nuristan, 24.7.52, O. - Afghanistan’, ‘ ex coll. Scheerpeltz’, ‘ COTYPUS Mannerheimia afghanica O. Scheerpeltz’ (NMW).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA9C495FE8EFAA8694E.taxon	discussion	Remarks. Mannerheimia afghanica was originally described from “ Afghanistan, Nouristan, Kamdesch, 2 000 m ”. The type series is large (the holotype and 210 paratypes) and deposited in Zoologisches Forschungsmuseum Alexander Koenig, Bonn, Germany (ZFMK) and Muséum d’Histoire Naturelle de Paris, France (MHNH). Specimens of the studied material above are originated from the type localities but do not belong to the type series.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA9C495FC49FF496EC0.taxon	materials_examined	Material examined. INDIA: KASHMIR: 1 ♂: ‘ Pahlgam 30. V. ’, ‘ Pahlgam, 1200 ft. Kashmir A. Kincel. lgt. ’ (NMPC).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA9C495FC49FF496EC0.taxon	discussion	Remarks. Mannerheimia obscura is widely distributed in the Himalayan region, from Pakistan and Kashmir to Nepal.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA7C495FACAFB286C10.taxon	description	(Figs 2, 6 – 7)	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA7C495FACAFB286C10.taxon	materials_examined	Type material examined. Holotype (Fig. 2; dissected; right antennomeres 7 – 11 missing): ‘ NEPAL P: Karnali, D: Humla, | Simikot, ~ 14 km SE, 3450 m | 29 ° 53´37 ´´ N, 81 ° 55´36 ´´ E, | 18. VI. 2022, disturbed mixed | forest, leg. A. Hetzel, 22 - # 05 ’ <printed>, ‘ HOLOTYPE | Mannerheimia | feldmanni sp. nov. | Shavrin A. V. 2025 ’ <red, printed> (cF, to be eventually deposited in Naturkundemuseum Erfurt, Germany). Paratype ♂ (right antennomeres 6 – 11 and elytra missing): same data as the holotype, with additional red printed label: ‘ PARATYPE | Mannerheimia | feldmanni sp. nov. | Shavrin A. V. 2025 ’ <red, printed> (cF).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA7C495FACAFB286C10.taxon	description	Description. Measurements (n = 2): HW: 0.56 – 0.57; HL: 0.35; OL: 0.17; AL (holotype): 1.02; PL: 0.45; PWmax: 0.75; PWmin: 0.67; ESL: 0.92; EW: 1.10; MTbL (holotype): 0.55; MTrL (holotype): 0.32 (MTrL 1 – 4: 0.17; MTrL 5: 0.15); AW: 1.11; AedL: 0.55; BL: 2.85 – 3.00 (holotype). Body moderately small and wide, shiny (Fig. 2). Body reddish brown, with slightly paler pronotum and elytra; antennomeres 7 – 11 yellow-brown; mouthparts, antennomeres 1 – 6 and legs yellowish. Latero-apical parts of clypeus with fine diagonal microreticulation; scutellum with fine transverse meshes; abdominal tergites with fine transverse microsculpture, indistinct on tergites VII – VIII. Head with variable and moderately large punctation, sparser in middle; neck with sparse punctation similar to that in middle of head; pronotum with moderately dense punctation, larger and deeper than that in head, finer and sparser on lateral portions and sparser in median part, mediobasal portion without punctures; scutellum without visible punctures; punctation of elytra slightly denser than that on pronotum, denser around scutellum, finer and sparser in middle, each elytron with indistinct five tangled longitudinal rows of punctures, confused in basal and apical portions; abdominal tergites with moderately dense and fine punctation, sparser and finer on tergites VII – VIII. Apical portion of clypeus with several moderately long erect setae; latero-apical parts of pronotum with two erect setae; elytra without visible setation; abdomen with indistinct sparse and fine setation. Head 1.6 times as broad as long, with indistinctly elevated middle portion; anteocellar foveae short, about as long as diameter of each ocellus, relatively shallow, reaching level of basal third of eyes; anterior portion between antennal insertion and anterior margins of eyes slightly concave. Eyes large and convex. Postocular portions relatively short, strongly narrowed toward neck. Ocelli large, slightly convex, situated distinctly behind level of posterior margins of eyes; distance between ocelli slightly longer than distance between ocellus and posterior margin of eye. Apical maxillary palpomere about two and a half times as long as preceding segment, from about middle strongly narrowed toward subacute apex. Antennae reaching apical basal margin of pronotum when reclined; basal antennomere wide, about two and a half times as long as broad, antennomere 2 slightly narrower and distinctly shorter than basal antennomere, 3 about as long as and slightly narrower than 2, 4 small, about as long as broad, 5 distinctly longer than 4, 6 slightly broader than 5, 7 slightly broader than 6, 8 slightly shorter than 7, 9 – 10 slightly broader than 8, apical antennomere distinctly more than twice as long as 10, from middle gradually narrowed toward subacute apex. Pronotum 1.6 times as broad as long, 1.3 times as broad as head, widest in middle, gradually narrowed both anteriad and posteriad; apical margin rounded, slightly narrower than rounded posterior margin; anterior angles widely rounded, not protruded anteriad; laterobasal margins not sinuate in front of widely rounded posterior angles; laterobasal portions narrowly impressed. Elytra 1.1 times as broad as long, strongly broadened apicad, twice as long as pronotum; apical margins straight; surface of each elytron in middle longitudinally elevated between punctures. Hind wings fully developed. Metatarsus 1.7 times as long as metatibia. Abdomen slightly broader than elytra, with two small oval tomentose spots in middle of abdominal tergite V; abdominal tergite VII with indistinct narrow palisade fringe. Male. Apical margin of abdominal tergite VIII truncate or indistinctly sinuate. Apical margin of abdominal sternite VIII widely concave. Aedeagus with moderately wide basal portion, gradually narrowed apically; median lobe narrow, gradually narrowed toward preapical part and broadened in apical part, with small rounded apex; parameres distinctly longer than apex of median lobe, broadened in preapical parts, each gradually narrowed apically; internal sac long and moderately narrow, with two fields of spines becoming larger in apical part (Fig. 6). Lateral aspect of aedeagus as in Fig. 7. Female unknown. Comparative notes. Mannerheimia feldmanni sp. nov. can be distinguished from all known species of the genus by the shape of the apical part of the median lobe. Based on the shapes of antennomeres 4 – 10, the shape of the pronotum widest in the middle and with rounded hind angles, and the length of elytra, M. feldmanni sp. nov. is similar to M. afghanica Coiffait, 1982, known from Afghanistan (Shavrin 2021 b) and M. loebli Shavrin, 2022, recently described from Pakistan (Shavrin 2022 c), from which it differs by the slightly longer body, the sparser punctation of the narrower pronotum and elytra, significantly narrower median lobe and different morphology of the internal sac.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA7C495FACAFB286C10.taxon	distribution	Distribution. Mannerheimia feldmanni sp. nov. is known only from the type locality in western Nepal.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA7C495FACAFB286C10.taxon	biology_ecology	Bionomics. The specimens were collected at an elevation 3450 m a. s. l. by sifting litter in disturbed mixed forest.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F30FFA7C495FACAFB286C10.taxon	etymology	Etymology. Patronymic; the species is named to honour Benedikt Feldmann (Münster).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F3EFFA7C495F9B6FD0B6DEE.taxon	materials_examined	Material examined. NEPAL: KARNALI: 1 ♀: Humla District, SE Simikot, Rodikot, 29 ° 52´24 ´´ N, 81 ° 55´57 ´´ E. 3750 m a. s. l., montane meadow ab. treeline. 19.06.2022. A. Hetzel leg. (cF).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F3EFFA7C495F9B6FD0B6DEE.taxon	discussion	Remarks. Trichodromeus telnovi was recently described from Jumla (Karnali, western Nepal) based on a single male. The studied female originates from the same province and morphologically is similar to the holotype, but it is longer (the length of the body of the holotype is 5.45 mm), protarsomeres 1 – 4 are distinctly narrower, and the pronotum and the elytra are slightly broader. Measurements of the female: HW: 0.90; HL: 0.60; OL: 0.21; LT: 0.12; AL: 3.02; PL: 0.70; PWmax: 1.02; PWmin: 0.81; ESL: 1.45; EW: 1.72; AW: 1.60; BL: 6.10. Posterior margins of abdominal tergite VIII and sternite VIII rounded.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F3FFFA6C495FF3FFDF068D1.taxon	description	(Figs 3, 8 – 9)	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F3FFFA6C495FF3FFDF068D1.taxon	materials_examined	Material examined. NEPAL: 1 ♂: Manaslu, Barapokhari Lekh, 23 km NE Besisahar, 28 ° 21´N 84 ° 33´E. 3800 – 4100 m a. s. l., sieved from moss and Rhododendron leaf litter. 14.09.2000. A. Hetzel leg. (cF).	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
03C087AF2F3FFFA6C495FF3FFDF068D1.taxon	discussion	Remarks. Coryphium loebli was originally described from Gorha District (Chuling Khola), eastern Nepal and was attributed to the sikkimense group established by Zerche (1990). This species is clearly distinguished from other related species of the group by the shape of dorsal lobe and the morphology of the internal sac with the presence of large sclerotized structures. The aedeagus of C. loebli illustrated by Zerche (1993; Fig. 18) is deformed, thus I decide to provide figures of it of the studied specimen (Figs 8 – 9). Habitus as in Fig. 3. Measurements of the studied specimen: HW: 0.41; HL: 0.22; OL: 0.12; AL: 0.82; PL: 0.37; PWmax: 0.47; PWmin: 0.36; ESL: 0.60; EW: 0.64; AW: 0.67; AedL: 0.47; BL: 2.19.	en	Shavrin, Alexey V. (2025): New species and records of Omaliinae MacLeay, 1925 (Coleoptera: Staphylinidae) from the eastern Palaearctic region. Zootaxa 5601 (1): 157-168, DOI: 10.11646/zootaxa.5601.1.8, URL: https://doi.org/10.11646/zootaxa.5601.1.8
