identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03CD1C633B14527CFF4160F2FE4FFBC2.text	03CD1C633B14527CFF4160F2FE4FFBC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	(Burgosia) Boeters 2003	<div><p>Subgenus Burgosia Boeters, 2003</p><p>Type species: Spiralix (Burgosia) burgensis Boeters 2003 by original designation.</p></div>	https://treatment.plazi.org/id/03CD1C633B14527CFF4160F2FE4FFBC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Álvarez, José Francisco Martín;Quiñonero-Salgado, Sergio;López-Soriano, Joaquín;Glöer, Peter	Álvarez, José Francisco Martín, Quiñonero-Salgado, Sergio, López-Soriano, Joaquín, Glöer, Peter (2024): Description of three new stygobiont snails of the families Hydrobiidae and Moitessieriidae from the southwestern Iberian Peninsula. Nemus 14: 88-101, DOI: 10.5281/zenodo.17026029
03CD1C633B145279FF2A6757FDDEFE86.text	03CD1C633B145279FF2A6757FDDEFE86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spiralix (Burgosia) aenigmatica Álvarez & Quiñonero-Salgado & López-Soriano & Glöer 2024	<div><p>Spiralix (Burgosia) aenigmatica sp. nov.</p><p>(Figs. 1-2)</p><p>Type material. Holotype RMNH.MOL.350846 (Naturalis) . Paratypes: 4 s. in RMNH.MOL.350847 (Naturalis). 5 s. in SQS coll. 12 s. in JFMA coll .</p><p>Type locality. Fuente de Sevilla, Zufre (Huelva); 29S X734085 Y4190034. 05⁄10⁄2022; 16⁄12⁄2022; 09⁄01⁄2023; 14⁄05⁄2023 , JFMA leg. 375m a.s.l. (Fig. 10A). The spring of the type locality flows out of limes- tone, and is collected in an artificial vaulted basin, which eventually feeds an irrigation pool “ alberca ” used for water supply to a neighbouring orchard .</p><p>Etymology. The specific name refers to the enigma represented by the large gap in the distribution range of the subgenus, until now only known from the Cantabrian Mountains and surrounding areas, in the north of Spain, very far from the locality where the new species was found.</p><p>Description. Shell small, translucent, fragile, subcylindrical and turriculated, with deep sutures and 3–3.5 convex whorls with orthocline growth lines. Size of about 1.0 mm high and 0.5 mm wide. Teleoconch microsculpture formed by irregularly arranged cuneiform marks. Protoconch microsculpture formed by angular depressions, irregularly arranged. Umbilicus deep. Aperture oval, aligned with the vertical axis, 0.4 mm high and 0.3 mm wide.</p><p>Dimensions. Holotype 0.90 mm high and 0.52 mm wide. For complete size ranges see Table 1 and Fig. 3. This is the smallest species in this genus.</p><p>Habitat. Stygobitic.</p><p>Differentiating characters. Spiralix (Burgosia) burgensis Boeters, 2003 has a similar shape, but has larger whorls, making it higher (1.24–1.44 mm compared to 0.86–1.01 mm) and wider (apex 0.73–0.85 mm compared to 0.47–0.64 mm; last whorl 0.88–0.96 mm vs. 0.58–0.71 mm). The ovoid aperture is slanted instead of vertical.</p><p>Spiralix (Burgosia) affinitatis Boeters, 2003 is quite different, having much more inflated whorls, resulting in an overall ovoid outline. Its length (1.07–1.24 mm) and width (apex 0.66–0.76; last whorl 0.78–0.93mm) fit in between Spiralix aenigmatica sp. nov. and S. burgensis . This species also has a slanted aperture.</p><p>Boeters (2003) did not elaborate on the microsculpture of the species he described, but Quiñonero-Salgado et al. (2017) presented SEM images of the six thenknown species, including the two species mentioned above. From this, it is evident that the microsculpture of the shell of S. aenigmatica sp. nov. is characteristic for the subgenus Burgosia: irregular cuneiform marks on the teleoconch, best seen on the first teleoconch whorls, and the protoconch with a pitted sculpture of angular depressions.</p><p>Some other species in the nominal genus Spiralix from the eastern Iberian Peninsula have to some degree similar marks at the initial whorl of the teleoconch (see Corbella et al., 2014), but they are much less apparent and developed than in Burgosia (detailed conchological redescription of this genus by Quiñonero-Salgado et al., 2017). So, the subgenus classification of the new species should be considered as tentative, until new evidence (anatomical or molecular) allows for more reliable criteria.</p><p>Remarks. The family Moitessieriidae includes only strict stygobiont species. Within this family, in Spain the genus Spiralix Boeters, 1972 is represented by nine species or subspecies in the Valencian Community, all included in the nominate subgenus Spiralix, and another seven species of the subgenus Burgosia Boeters, 2003 restricted to the north of the Iberian Peninsula. Of these seven species in the second subgenus, two are endemic to Asturias (Quiñonero-Salgado et al., 2018a), two are from Cantabria, another two are exclusively found in the province of Burgos (northern part of the Castilla y León Autonomous Community), and one more is found both in Burgos and the Basque Country. While many other Moitessieriidae species are known in Catalonia, Aragón and the Valencian Community, little is known about this family in the rest of Spain, to date with no known representatives in the whole centre and southern part.</p><p>The finding of a species of the subgenus Burgosia in Andalusia enlarges its distribution range about 600 km to the south from all previously known species. Such a big gap is quite surprising for a subgenus that presumably has very limited dispersion capability. Further exploratory research is needed to determine whether there is a biogeographic explanation for this distribution, or if it is caused by a lack of knowledge of the stygobiont malacofauna in a large part of the Iberian Peninsula. The huge increase in our knowledge of stygobiont molluscs in Iberia over the last two decades makes the second explanation more likely.</p></div>	https://treatment.plazi.org/id/03CD1C633B145279FF2A6757FDDEFE86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Álvarez, José Francisco Martín;Quiñonero-Salgado, Sergio;López-Soriano, Joaquín;Glöer, Peter	Álvarez, José Francisco Martín, Quiñonero-Salgado, Sergio, López-Soriano, Joaquín, Glöer, Peter (2024): Description of three new stygobiont snails of the families Hydrobiidae and Moitessieriidae from the southwestern Iberian Peninsula. Nemus 14: 88-101, DOI: 10.5281/zenodo.17026029
03CD1C633B1D5275FC976635FC6DFAAC.text	03CD1C633B1D5275FC976635FC6DFAAC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Guadiella algarvensis Álvarez & Quiñonero-Salgado & López-Soriano & Glöer 2024	<div><p>Guadiella algarvensis sp. nov.</p><p>(Figs. 7-8)</p></div>	https://treatment.plazi.org/id/03CD1C633B1D5275FC976635FC6DFAAC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Álvarez, José Francisco Martín;Quiñonero-Salgado, Sergio;López-Soriano, Joaquín;Glöer, Peter	Álvarez, José Francisco Martín, Quiñonero-Salgado, Sergio, López-Soriano, Joaquín, Glöer, Peter (2024): Description of three new stygobiont snails of the families Hydrobiidae and Moitessieriidae from the southwestern Iberian Peninsula. Nemus 14: 88-101, DOI: 10.5281/zenodo.17026029
03CD1C633B1D5275FC9967B1FC5DFB0C.text	03CD1C633B1D5275FC9967B1FC5DFB0C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Guadiella Boeters 2003	<div><p>Genus Guadiella Boeters, 2003</p><p>Type species: Belgrandiella andalucensis Boeters, 1983 by original designation</p></div>	https://treatment.plazi.org/id/03CD1C633B1D5275FC9967B1FC5DFB0C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Álvarez, José Francisco Martín;Quiñonero-Salgado, Sergio;López-Soriano, Joaquín;Glöer, Peter	Álvarez, José Francisco Martín, Quiñonero-Salgado, Sergio, López-Soriano, Joaquín, Glöer, Peter (2024): Description of three new stygobiont snails of the families Hydrobiidae and Moitessieriidae from the southwestern Iberian Peninsula. Nemus 14: 88-101, DOI: 10.5281/zenodo.17026029
03CD1C633B1D5275FF7467B6FB92FB82.text	03CD1C633B1D5275FF7467B6FB92FB82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Onubiella suberensis Álvarez & Quiñonero-Salgado & López-Soriano & Glöer 2024	<div><p>Onubiella suberensis sp. nov.</p><p>(Figs. 4 -6)</p><p>Type material. Holotype RMNH.MOL.350848 (Naturalis) (Fig.5A) . Paratypes:5 s. RMNH.MOL.350849 (Naturalis); 20 s. in JFMA collection; 20 s. in PG collection.</p><p>Type locality. Fuente de Sevilla, Zufre (Huelva province): 29S X734085 Y4190034; 05⁄10⁄2022; 16⁄12⁄2022; 09⁄01⁄2023; 14⁄05⁄2023 , JFMA leg. 375m a.s. l. (Fig. 10A). See description of the type locality in the species described above, Spiralix aenigmatica sp. nov. Description. Very small, valvatiform, planispiral or very low spire. Protoconch of about 1.5 whorls, teleoconch about 1.5 convex whorls with clear suture (especially dorsal side) and wide umbilicus. The whorls attach to the lower part of the previous whorl, without any overlap, remaining about flat on the dorsal side, whereas difference in whorl height is clearly reflected on the ventral side. Protoconch mottled with very fine, irre- gularly-arranged pits, teleoconch smooth with growth lines. Aperture round, peristome slightly reflected .</p><p>Anatomy. Penis simple without any outgrowth, broad at the basis and slim at the distal end. Female sex tract unknown.</p><p>Etymology. suber (Latin =cork), refers to the Fuente del Corcho (= Cork Spring) in which the species was discovered.</p><p>Dimensions. See Table 2 and Fig. 6.</p><p>Habitat. Stygobitic.</p><p>Other localities. Fuente del Corcho, La Corte-Cortegana (Huelva province); 29S X690054 Y4202569. 25⁄03⁄2018; 22⁄05⁄2021; 21⁄06⁄ 2021. 580 m a. s.l.</p><p>(Fig. 10B).</p><p>Distribution. Only known from the two localities mentioned above.</p><p>Differentiating characters. The shell of Corbellaria celtiberica Callot-Girardi &amp; Boeters, 2012, described from the province of Soria in northern Spain, is very similar but does not have the narrow neck (note the reflected peristome is only present in fully grown specimens). Also, the shell is slightly higher and height increases with width, whereas in Onubiella gen. nov. it is constant. In Corbellaria aperture height increases with width, in Onubiella gen. nov. also, but less. Anatomically, Onubiella suberensis sp. nov. has a simple penis without any outgrowth, while C. celtiberica has a simple lobe (Callot-Girardi &amp; Boeters, 2012, Fig. 10D).</p></div>	https://treatment.plazi.org/id/03CD1C633B1D5275FF7467B6FB92FB82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Álvarez, José Francisco Martín;Quiñonero-Salgado, Sergio;López-Soriano, Joaquín;Glöer, Peter	Álvarez, José Francisco Martín, Quiñonero-Salgado, Sergio, López-Soriano, Joaquín, Glöer, Peter (2024): Description of three new stygobiont snails of the families Hydrobiidae and Moitessieriidae from the southwestern Iberian Peninsula. Nemus 14: 88-101, DOI: 10.5281/zenodo.17026029
