identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03D60217CC48BA6517876987FD3D23BE.text	03D60217CC48BA6517876987FD3D23BE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yoyetta Moulds 2012	<div><p>List of  Yoyetta species</p><p>Yoyetta aaede (Walker, 1850)</p><p>Yoyetta abdominalis (Distant, 1892)</p><p>Yoyetta australicta (Popple &amp; Emery, 2022)</p><p>Yoyetta celis (Moulds, 1988)</p><p>Yoyetta corindi (Popple &amp; Emery, 2022)</p><p>Yoyetta cumberlandi (Emery, Emery &amp; Popple, 2015)</p><p>Yoyetta darug sp. nov.</p><p>Yoyetta delicata (Popple &amp; Emery, 2022)</p><p>Yoyetta denisoni (Distant, 1893)</p><p>Yoyetta douglasi (Popple &amp; Emery, 2020)</p><p>Yoyetta electrica (Emery, Emery &amp; Popple, 2019)</p><p>Yoyetta enigmatica (Popple &amp; Emery, 2020)</p><p>Yoyetta fluviatilis (Emery, Emery &amp; Popple, 2015)</p><p>Yoyetta fumea sp. nov.</p><p>Yoyetta grandis (Emery, Emery &amp; Popple, 2019)</p><p>Yoyetta humphreyae (Moulds &amp; Popple, 2018)</p><p>Yoyetta hunterorum (Moulds, 1988)</p><p>Yoyetta ignita (Popple &amp; Emery, 2022)</p><p>Yoyetta incepta (Walker, 1850)</p><p>Yoyetta kershawi (Goding &amp; Froggatt, 1904)</p><p>Yoyetta landsboroughi (Distant, 1882)</p><p>Yoyetta loftyensis (Popple &amp; Emery, 2020)</p><p>Yoyetta ngarabal (Popple &amp; Emery, 2020)</p><p>Yoyetta nigrimontana (Emery, Emery &amp; Popple, 2015)</p><p>Yoyetta psammitica sp. nov.</p><p>Yoyetta regalis (Emery, Emery &amp; Popple, 2019)</p><p>Yoyetta repetens (Emery, Emery &amp; Popple, 2015)</p><p>Yoyetta robertsonae (Moulds, Popple &amp; Emery, 2020)</p><p>Yoyetta robusta (Popple &amp; Emery, 2020)</p><p>Yoyetta serrata (Emery, Emery &amp; Popple, 2019)</p><p>Yoyetta spectabilis (Emery, Emery &amp; Popple, 2019)</p><p>Yoyetta subalpina (Emery, Emery &amp; Popple, 2019)</p><p>Yoyetta timothyi (Emery, Emery &amp; Popple, 2019)</p><p>Yoyetta tristrigata (Goding &amp; Froggatt, 1904)</p><p>Yoyetta verrens (Emery, Emery &amp; Popple, 2019)</p><p>Genus  Yoyetta Moulds, 2012</p><p>The diagnosis follows Moulds (2012) with modifications of Emery et al. (2015). Type species:  Cicadetta celis Moulds, 1988 .</p></div>	https://treatment.plazi.org/id/03D60217CC48BA6517876987FD3D23BE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Emery, David L.;Emery, Nathan J.;Popple, Lindsay W.	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC4BBA6F17876862FD66250E.text	03D60217CC4BBA6F17876862FD66250E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yoyetta darug Emery & Emery & Popple 2025	<div><p>Yoyetta darug sp. nov.</p><p>(Plate 1; Plate 2A–B; Plate 3A–B; Plate 4A; Figs. 1, 2C–D, 3–5).</p><p>Types.  Holotype ♂, Silverdale NSW, 33 o 52’ 08’’S 150 o 37’11’’E, 14.xi.2007, D. Emery (EME0489-002) (K.624928, AM).</p><p>Paratypes. NEW SOUTH WALES. 1♂ 1♀, Bourke St., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.0&amp;materialsCitation.latitude=-33.793056" title="Search Plazi for locations around (long 151.0/lat -33.793056)">Parramatta</a>, 33°47’35’’S 151°00’00’’E, 6.xii.2022, D. Emery (EME0489-017, -018) (K. 624929–930, AM);  1♂, same location as holotype, 7.xi.2007, D. Emery &amp; L. Popple (EME0489-001; genitalia prep 489-1);  1♂, Castlereagh <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.74417&amp;materialsCitation.latitude=-33.681114" title="Search Plazi for locations around (long 150.74417/lat -33.681114)">Nature Reserve</a>, 33 o 40’52’’S 150 o 44’39’’E, 18.x.2015, N. Emery (EME0489-003);  1♂, same location, 27.i.2019, N. Emery (EME0489-004); 1♂, same location, 22.xi.2017, N. Emery &amp; C. Foster (EME0489-005); 1♂, same location, 23.x.2020, D. Emery (EME0489-006);  1♂, Lake Parramatta Reserve,  Sth James Ruse Drive, 33 o 793’S 150 o 010’E, 31.x.2020, J. Poyitt (EME0489-007);   1♂, Longneck Lagoon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.56029&amp;materialsCitation.latitude=-33.57361" title="Search Plazi for locations around (long 150.56029/lat -33.57361)">Scheyville</a> NPk, 33°34’25’’S 150°33’37’’E, 27.xii.2018, J. Poyitt (EME0489-008, genitalia prep 489-2);   2♂, Carysfield Park, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.99806&amp;materialsCitation.latitude=-33.903614" title="Search Plazi for locations around (long 150.99806/lat -33.903614)">Bass Hill</a>, 33°54’13”S 150°59’53”E, 6.xii.2022, D. Emery (EME0489-019 – 020);   1♂, Bourke St., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.0&amp;materialsCitation.latitude=-33.793056" title="Search Plazi for locations around (long 151.0/lat -33.793056)">Parramatta</a>, 33°47’35’’S 151°00’00’’E, 11.xi.2022, D. Emery (EME0489-009);  2♂, same location, 24.xii.2022, D. Emery (EME0489-010 – 011); 2♂ 2♀, same location, 28.x1.2022, D. Emery (EME0489-012 – 014; 014 &amp; 015 mating); 1♂, same location, 4.xi.2022, D. Emery (EME0489-016); 2♂ same location, 6.xii.2022, D. Emery (EME489-019 – 020); 3♂ 2♀, same location 11.xii.2022, D. Emery (EME489-021 – 024; 021 &amp; 022 mating);  1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.44305&amp;materialsCitation.latitude=-33.07" title="Search Plazi for locations around (long 150.44305/lat -33.07)">Nattai</a>, 33°04’12’’S 150°26’35’’E, 20.xii.2020, D. Emery (EME0489-025);   1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.44305&amp;materialsCitation.latitude=-33.236668" title="Search Plazi for locations around (long 150.44305/lat -33.236668)">Burragorang Nature Res.</a>, 33°14’12’’S 150°26’35’’E, 29.xii.2010, D. Emery (EME0489-026);   1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.625&amp;materialsCitation.latitude=-34.349167" title="Search Plazi for locations around (long 150.625/lat -34.349167)">Avon Dam Rd.</a>, 34°20’57’’S 150°37’30’’E, 23.xii.1998, D. Emery (EME0489-027);   1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.76694&amp;materialsCitation.latitude=-34.066666" title="Search Plazi for locations around (long 150.76694/lat -34.066666)">Mt Annan Botanical Gardens</a>, 34°04’00’’S 150°46’01’’E 12.xi.2024, N. Emery (EME0489-028) (DE);   3♂, Silverdale,  Nepean Basin, 2.xii.2003, L. Popple &amp; D. Emery, 489-0001 to 489-0003 (1M genitalia prep 489-01);   1♀, Bourke St., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.0&amp;materialsCitation.latitude=-33.799168" title="Search Plazi for locations around (long 151.0/lat -33.799168)">Parramatta</a>, 33°47’57’’S 151°00’00’’E, 11.xi.2022, D. Emery (EME0489-015, mating with -014);   1♂,  Bass Hill, xii.2009, D. Emery (LWP);   1♂, Sth. of James Ruse Drive, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.01&amp;materialsCitation.latitude=-33.79361" title="Search Plazi for locations around (long 151.01/lat -33.79361)">Lake Parramatta Reserve</a>, NSW, 33°47’37”S 151°0’36”E, 17.ii.2018, J. Poyitt, (HEMC489.001);  2♂ 1♀, same location, 29.xii,2018, J. Poyitt (HEMC489.002, 011, 012); 1♂, same location, 31.x.2020, J. Poyitt (HEMC489.009); 1♂, same location, 14.i.2021, J. Poyitt (HEMC489.007); 4♂, same location, 22–30.x.2022, J. Poyitt (HEMC489.017 – 020); 1♂, same location, 19.xi.2022, J. Poyitt (HEMC489.044); 1♀, same location, 28.xi.2022, J. Poyitt (HEMC489.022);  1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.01&amp;materialsCitation.latitude=-33.79361" title="Search Plazi for locations around (long 151.01/lat -33.79361)">Lake Parramatta Reserve</a>, NSW, 33°47’37”S 151°0’36”E, 26.xii.2018, S. Jones (HEMC489.016);  1♂, same location, 24.xi.2019, J. Poyitt, (HEMC489.010); 1♂, same location, 8.xii.2019, J. Poyitt, (HEMC489.009); 2♂, same location, 10.i.2024, J. Poyitt (HEMC489.044, -045);  1♂, Castlereagh <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.76445&amp;materialsCitation.latitude=-33.681946" title="Search Plazi for locations around (long 150.76445/lat -33.681946)">Nature Reserve</a>, 33 o 40’ 55’’S 150 o 45’52’’E, 1.xii.2018, J. Poyitt, (HEMC489.006);   1♂, Longneck Lagoon, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.56029&amp;materialsCitation.latitude=-33.57361" title="Search Plazi for locations around (long 150.56029/lat -33.57361)">Scheyville</a> NPk, 33°34’25’’S 150°33’37’’E, 27.xii.2018, J. Poyitt (HEMC489.003);   1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.96138&amp;materialsCitation.latitude=-33.785557" title="Search Plazi for locations around (long 150.96138/lat -33.785557)">Norwest Blvd</a>, 33°47’08’’S 150°57’41’’E, 10.xii.2018, J. Poyitt (HEMC489.014);   1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.98834&amp;materialsCitation.latitude=-33.720554" title="Search Plazi for locations around (long 150.98834/lat -33.720554)">Fred Caterson Res.</a>, NSW, 33°43’14’’S 150°59’18’’E, 15.xi.2020, J. Poyitt (HEMC489.013);   1♂, Lookout Trail, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.89471&amp;materialsCitation.latitude=-33.54889" title="Search Plazi for locations around (long 150.89471/lat -33.54889)">Cattai</a> NPk., 33°32’56’’S 150°53’41’’E, 30.i.2021, J. Poyitt (HEMC489.005);   1♂, Box Hill fire <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.90054&amp;materialsCitation.latitude=-33.638615" title="Search Plazi for locations around (long 151.90054/lat -33.638615)">Stn.</a>, 33°38’19’’S 151°54’02’’E, 30.x.2022, J. Poyitt (HEMC489.021);   1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.01334&amp;materialsCitation.latitude=-33.793888" title="Search Plazi for locations around (long 151.01334/lat -33.793888)">Redeemer Baptist School</a>, 33°47’38’’S 151°00’48’’E, 30.xi.2022, J.Poyitt (HEMC489.045);   3♂ 2♀, Century Cct., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.01334&amp;materialsCitation.latitude=-33.793888" title="Search Plazi for locations around (long 151.01334/lat -33.793888)">Norwest</a>,NSW, 33°47’38’’S 151°00’48’’E, 5–10.xii.2022, J. Poyitt (emerging on  Eucalyptus crebra; HEMC489.025, 027, 029, 030, 032);   4♂ 1♀, Solent Cct., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.96056&amp;materialsCitation.latitude=-33.73611" title="Search Plazi for locations around (long 150.96056/lat -33.73611)">Norwest</a>, NSW, 33°44’10’’S 150°57’38’’E, 10–16.xii.2022, J. Poyitt (emerging on  Eucalyptus crebra; HEMC489.034 – 038);   2♂, Ted Horwood Res., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=151.005&amp;materialsCitation.latitude=-33.760555" title="Search Plazi for locations around (long 151.005/lat -33.760555)">Baulkham Hills</a>, 33°45’38’’S 151°00’18’’E, 2.i.2023, J. Poyitt (HEMC489.039, 040);   1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.98611&amp;materialsCitation.latitude=-33.72639" title="Search Plazi for locations around (long 150.98611/lat -33.72639)">Castle Hill Showground</a>, 33°43’35’’S 150°59’10’’E, 9.i.2023, J. Poyitt (HEMC489.041);   1♂, Douglas Park Drive, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.70694&amp;materialsCitation.latitude=-34.196114" title="Search Plazi for locations around (long 150.70694/lat -34.196114)">Douglas Park</a>, 34°11’46’’S 150°42’25’’E, 10.i.2023, J. Poyitt (HEMC489.042);   1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.80806&amp;materialsCitation.latitude=-34.26139" title="Search Plazi for locations around (long 150.80806/lat -34.26139)">Cataract Dam Picnic Area</a>, NSW, 34°15’41’’S 150°48’29’’E, 10.i.2023, J. Poyitt (HEMC489.043) (JP).</p><p>Distribution, habitat and seasonality. Restricted to central eastern New South Wales, where it is known subcoastally from around Illawarra and Burragorang regions, north to the Hunter River area (Fig. 1). It is most prevalent in Greater Western Sydney. Adults occur in open forest, mainly on rough-barked eucalypts, especially ironbarks ( Eucalyptus siderophyloia,  E. fibrosa in Western Sydney and  E. crebra around Parramatta and the Hills District) and stringybarks (e.g.  Eucalyptus obliqua) (Figs. 2A–B). Females actively fly to ironbarks and secrete themselves vertically along the grooves in the rough bark on the main trunk at around 2–5 m, from which they wingflick to attract males (two mating couples were captured in these grooves). In this position they are extremely well camouflaged and difficult to dislodge when attempts are made by human observers to capture them using a longhandled net. Males fly to the locations of the females and continuously move, tick and buzz, leaving if no further encouragement occurs. Males respond to simulated wing-flicks (e.g. finger snapping) and playback of their song recordings, but are quite wary.</p><p>This species has been encountered from late October to early February and co-occurs with several related and similar looking  Yoyetta species, including  Yoyetta humphreyae,  Yoyetta repetens and  Yoyetta cumberlandi .</p><p>Etymology. The specific epithetic honours a language of the traditional peoples of the Sydney Basin bioregion where this species is found (a noun in apposition).</p><p>Description of adult male (Plate 1A–B, E–G; Plate 2A–B; Plate 3A–B; Plate 4A; Fig. 3).</p><p>Head slightly narrower than the lateral margins of pronotum; mostly brown to orange-brown with extensive black colouration between and surrounding the ocelli; supra-antennal plates black, occasionally edged dull orange-brown; ocelli pale red; eyes dark brown to black; postclypeus mainly black, dorsal side dark orange-brown, ventral surface with narrow orange lateral margins and medial line; anteclypeus shiny black; rostrum reaching bases of hind coxae, brown basally becoming dark brown or black apically; antennae black.</p><p>Thorax mostly black. Pronotum with a narrow orange-brown fascia along midline surrounded by black; interior of pronotum brown to dark brown, with irregular black areas, especially along paramedian and lateral fissures; pronotal collar black on anterior margin, orange-brown to dark brown over remainder, black around margins of lateral angles. Mesonotum orange-brown to black; submedian and lateral sigilla black, separated by narrow orange-brown to dark brown areas; lateral margins sometimes orange-brown; cruciform elevation pale brown to pale orange-brown, black centrally, ridges pale brown; scutal depression black; wing grooves mainly dark orange-brown and black. Metanotum dark brown to black. Thorax below mainly orange-brown or dark brown.</p><p>Legs brown with dark brown to black markings variable between individuals. Coxae about half brown, half dark brown or black; fore femora with inner surface usually black but sometimes dark orange-brown, the outer surface predominantly dull orange-brown, femoral spines dark brown to black; mid and hind femora dark brown to black on anterior sides, paler brown on posterior; fore tibiae dark brown; mid and hind tibiae orange-brown; tarsi orange-brown or brown suffused with black; meracantha black on about basal half, otherwise pale brown.</p><p>Wings with forewing costal margin a mixture of orange-brown and dark orange-brown; venation generally dark brown to orange-brown; basal membrane light orange-brown. Hind wing venation dark brown; plagas and adjacent area surrounding vein 3a brown.</p><p>Opercula dark brown to black basally, plates pale brown; approximately 1.5x larger than adjacent hind coxae (Plate 2B).</p><p>Timbals (Plate 2A) with five long ribs; ribs 1–4 attached to basal spur, separated ventrally and spanning the timbal membrane, rib 5 short, unattached, occupying dorsal half of timbal only; timbal plate with well developed, elongate dome bearing a distinct point of apodeme attachment near centre.</p><p>Abdomen (Plate 3A–B) with tergite 1 black; tergite 2 predominantly black, sometimes with diffuse orange areas sublaterally on posterior margin, curved to follow outer margin of exposed timbals; tergites 3–7 predominantly pale orange-brown each with a narrow, rectangular, black mark on dorsal midline, this mark being broadest on tergite 3 and narrowing slightly on each subsequent tergite towards the posterior, tergite 3 often with a diffuse dark smudge on lateral areas (noted consistently on Parramatta specimens, Plate 3B), highly variable in extent between individuals; tergite 8 black, tending dark orange-brown towards posterior margin; epipleurites flexed ventrally, pale orange-brown, diffusely smudged with black. Sternite I orange-brown; sternite II orange with dark-brown to black areas medially and laterally; sternites III–VII pale orange-brown with a small black spot on VII at centre of the posterior segment; sternite VIII orange-brown.</p><p>Genitalia (Fig. 3) with pygofer upper lobe of moderate size, gradually tapering to a broadly rounded apex (Fig. 3A); basal pygofer lobe small weakly developed, broadly rounded; median lobe of uncus enlarged, very long (1.5– 2mm), concave ventrally, in lateral view narrow basally, expanded around mid-length to widest width, thereafter evenly tapering to a bluntly pointed apex, with ratio of length from base divided by widest width being&gt;5.0 (Fig. 3B); claspers robust, claw-like, partially meeting, in ventral view their inner margins diverging with an indentation midway along the length and their outer margins gently turned outwards; aedeagus recurved distally through 180 degrees, the vesica weakly toothed either side subapically, apex terminating in a short, slender ‘beak’ downturned&gt;45 o when viewed laterally, with a pair of very small flat ear-like lobes pointing inwards, minutely toothed posteriorly; pseudoparameres very long, nearly reaching most distal part of theca, very slender, hair-like and weakly knobbed apically (Figs. 3C–D).</p><p>Description of adult female (Plate 1C–D, Plate 4A).</p><p>Head, thorax, wings and legs similar to male, except with greater amounts of ochraceous-brown fascia around ridges on mesonotum.</p><p>Abdomen with more pronounced annular markings on tergites (Plate 4A). Tergite 1 orange, black medially and diffusely black along anterior margin of lateral third; tergites 2–7 predominantly dark orange-brown each with a distinct black marking on the dorsal midline, broadest on tergite 3, extending laterally over the anterior half of each tergite and blending into dark orange-brown along the posterior half with a thin lighter orange-brown posterior margin; tergite 8 black over anterior half, tending dark orange-brown over remainder, with a black spot on lateral aspect. Sternite I black; sternites II–VII pale orange-brown with a black triangular marking along the midline, base widest anteriorly, apex posteriorly; sternite VIII darker orange-brown; abdominal segment 9 pale brown over dorsal two-thirds, darker brown ventrally, dorsal beak black, a distinct black longitudinal marking either side of dorsal midline, extending laterally along the anterior margin, enclosing a pale brown longitudinal triangle along the midline; a small black spot laterally near the posterior apex; ovipositor sheath dark brown, black posteriorly, ovipositor dark brown, becoming black posteriorly, extending 1–1.5 mm beyond abdominal segment 9 (Plate 4A).</p><p>Measurements (in mm; range with mean in parentheses for 12 males and 12 females, including smallest and largest specimens). Length of body including head: male, 19.7–22.1 (21.5); female, 23.5–25.8 (24.6). Length of forewing: male, 21.6–26.1 (24.5); female 28.5–32.4 (30.0). Width of forewing: male, 7.7–8.9 (8.0); female, 8.5–10.2 (9.4). Width of head (including eyes): male, 5.3–6.2 (5.8); female, 6.1–7.1 (6.6). Width of pronotum (across lateral angles): male, 5.4–6.2 (5.8); female, 5.8–7.6 (6.7). Width of abdomen: male, 5.4–6.4 (5.8); female, 6.2–6.7 (6.4). Length of ovipositor: female, 7.5–9.3 (8.3).</p><p>Morphological variation within and between populations. Both male and female specimens exhibit a colour range from medium brown to dark brown to black over the head and pronotum. The dorsal abdominal markings of all females captured to date range from black to grey.</p></div>	https://treatment.plazi.org/id/03D60217CC4BBA6F17876862FD66250E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Emery, David L.;Emery, Nathan J.;Popple, Lindsay W.	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC42BA7617876CAEFD6727CE.text	03D60217CC42BA7617876CAEFD6727CE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yoyetta psammitica Emery & Emery & Popple 2025	<div><p>Yoyetta psammitica sp. nov.</p><p>(Plates 2C–D, 3C–D, 4B, 5; Figs. 1, 2C–D, 6–9)</p><p>Types.  Holotype 1♂, Windamere Dam, Mudgee, NSW, 32 o 43’32’’S 149 o 46’19’’E, 8.i.2022, N. &amp; D. Emery &amp; C. Foster (EME0490 -017) (K624931, AM).</p><p>Paratypes. NEW SOUTH WALES. 1♂ 1♀, same data as holotype, (EME0490-030, -031) (K.924932, K.924936, AM);  2♂, same location as holotype, 2.i.2022, S. &amp; T. Corbin (EME0490-001 – 2; genitalia prep 490-1),  11♂ 1♀, same data as holotype (EME0490-003 – 014);  16♂ 2♀, same location as holotype, 8.i.2022, N. &amp; D. Emery &amp; C. Foster (EME0490-017 – 029; 032–035);   2♂, Goulburn R., nr <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.96028&amp;materialsCitation.latitude=-32.32861" title="Search Plazi for locations around (long 149.96028/lat -32.32861)">Wollar</a>, 32 o 19’43’’S 149 o 57’37’’E, 3.i.2022, T. Corbin (EME0490-036 – 7) ;   4♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.93361&amp;materialsCitation.latitude=-32.0375" title="Search Plazi for locations around (long 149.93361/lat -32.0375)">Cassilis</a> rest area, NSW, 32 o 02’15’’S 149 o 56’01’’E, 6.i.2013, N. C. &amp; D. Emery (EME0490-038 – 041; genitalia preps 490-3, 490-4) ;  3♂, same location, 15.xii, 2020, C. &amp; D. Emery (EME0490- 042 – 044);  5♂, same location, 23.i.2023, D. Emery (EME0490-058 – 062);   1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.94362&amp;materialsCitation.latitude=-32.888615" title="Search Plazi for locations around (long 149.94362/lat -32.888615)">Clandulla State Forest</a>, 32 o 53’19’’S 149 o 56’37’’E, 7.i.2010, D. Emery &amp; L. Popple (EME0490-045) ;   3♂ 2♀, Cooper Drive, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.81667&amp;materialsCitation.latitude=-32.9075" title="Search Plazi for locations around (long 149.81667/lat -32.9075)">Clandulla</a>, 32 o 54’27’’S 149 o 49’00’’E, 26.xi.2020, D. Emery (EME0490-046 – 050; genitalia prep 490-5) ;  4♂ 1♀, same location, 15.i.2021, N. &amp; D. Emery (EME0490-051 – 055);   1♂ 1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.71194&amp;materialsCitation.latitude=-32.791943" title="Search Plazi for locations around (long 149.71194/lat -32.791943)">Bocoble</a>, 32 o 47’31’’S 149 o 42’43’’E, 1050m, 11.xii.2021, H. Ottersdorf (EME0490-056 – 057) ;   1♂,  Denman, 5.xi.2007, R. Chin (EME0490-058) ;   4♂, Whitegum Lookout, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.0389&amp;materialsCitation.latitude=-31.2925" title="Search Plazi for locations around (long 149.0389/lat -31.2925)">Warrumbungle</a> NPk, 31 o 17’33’’S 149 o 02’20’’E, 16.xi.2013, N. &amp; D. Emery (EME0717-059 – 062) ;  3♂, 14.xi.2013, N. &amp; D. Emery (EME0490-063 – 065; genitalia prep 490-2);  1♂, same location, 12.i.2014, N., C. &amp; D. Emery (EME0490-066);  1♂ 1♀, same location, 12.i.2014, N., C. &amp; D. Emery, (EME0490-067 – 068; genitalia prep 490-6);  1♂, same location, 8.xii.2015, C. &amp; D. Emery (EME0490-069);  2♂, same location, 5.xi.13, C. &amp; D. Emery (EME0490-070 – 071);  1♀, same location, 8.xii.2015, N. &amp; D. Emery (EME0490-072) (DE);  1♂, same location as previous, 14.xi.2013, N. &amp;. D. Emery;  1♂, same data as previous, 30.xi.2013;   1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.93361&amp;materialsCitation.latitude=-32.0375" title="Search Plazi for locations around (long 149.93361/lat -32.0375)">Cassilis</a> rest area, NSW, 32 o 02’15’’S 149 o 56’01’’E, 6.i.2013, N. C. &amp; D. Emery ;  1♂, same location as previous, 15.xi.2020, C. &amp; D. Emery;   1♂, Napolean Reef area via <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.7564&amp;materialsCitation.latitude=-33.43889" title="Search Plazi for locations around (long 149.7564/lat -33.43889)">Bathurst</a>, 33°26’20’’S 149°45’23’’E, 8.i.2010, Popple &amp; Emery, open woodland ;  2♂, same location as holotype, 2.i.2022, S. &amp; T. Corbin (LWP);  2♂, same data as holotype (EME0490-015 – 6);   1♂, Glen Davis Rd. between Airly Creek Rd. and <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.06778&amp;materialsCitation.latitude=-33.134445" title="Search Plazi for locations around (long 150.06778/lat -33.134445)">Pinegrove Rd.</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=150.06778&amp;materialsCitation.latitude=-33.134445" title="Search Plazi for locations around (long 150.06778/lat -33.134445)">Capertee</a>, NSW, 33°08’04’’S 150°04’04”E, 13.xii.2022, J. Poyitt, (HEMC490.001) ;  4♂, same location as holotype, 11.i.2023, J. Poyitt (HEMC490.004 – 007);  4♂, same location as holotype, 17.i.2023, J. Poyitt, (HEMC490.008 – 011) (JP) .</p><p>Other material.   QUEENSLAND. 1♂, Carnarvon Stn, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.753&amp;materialsCitation.latitude=-24.809" title="Search Plazi for locations around (long 147.753/lat -24.809)">Homestd</a>, 24.809°S 147.753°E, 747 m, 13.x.2014, J. Buchan, mv lamp, 37156, T199410 ;   1♀, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.753&amp;materialsCitation.latitude=-24.809" title="Search Plazi for locations around (long 147.753/lat -24.809)">Carnarvon Stn</a>, nr homestead, 24.809°S 147.753°E, 747 m, 13.x.2014, B. Hawkins, hand collected, flowering flat, T199411 ;   1♂,  Mt Moffatt NP, top shelter shed, 1000 m, 10–12.xii.1987, Monteith, Thompson, Yeates (QM)  .</p><p>Distribution, habitat and seasonality. The majority of specimens of  Yoyetta psammitica sp. nov. have been found along the western slopes of the Great Dividing Range in New South Wales from around Capertee and Wollemi National Park, north to around Merriwa and Cassilis and further northwest into the Warrumbungle Range National Park (NPk; Fig. 1). An isolated population also occurs in the Carnarvon Range in central Queensland. In New South Wales, this species appears to favour smooth barked eucalypts, particularly  Eucalyptus cannonii (Capertee Stringybark) around Mudgee (Windamere Dam and northern Cudgegong River) and Clandulla (Fig. 2C–D). At Cassilis Park rest area on the Golden Highway, specimens are also attracted to narrow-leaved ironbark trees ( E. crebra), while in the Warrumbungle NPk, they are often found on white gums ( E. rossii). Individuals call in flight and while moving constantly around the tree trunks. Single populations are mostly present at Cassilis rest area and Windamere Dam, but at Clandulla and Bocoble,  Y. psammitica sp. nov. co-occurs with  Y. humphreyae,  Yoyetta timothyi Emery, Emery &amp; Popple and two additional undescribed  Yoyetta species. In the Warrumbungle NPk,  Y. psammitica sp. nov. cohabits with  Y. humphreyae and  Y. australicta Popple &amp; Emery. Most specimens of  Y. psammitica sp. nov. have been collected from mid-November to late-January. It has been noted that since 2015, two years after wildfires burnt out over 80% of Warrumbungle NPk, specimens of this species have been difficult to find, with those of  Y. australicta and  Y. humphreyae being much more common. The reason(s) for this change in prevalence is not clear.</p><p>Etymology. The term  “psammitica ” (adjective, “sandstone”) reflects the distribution of this species throughout the open sandstone woodlands of the western slopes of the Great Dividing Range of NSW and southern Queensland.</p><p>Description of adult male</p><p>(Plates 2C–D, 3C–D, 5A–D, G; Figs. 2C–D, 6)</p><p>Head slightly narrower than the lateral margins of pronotum; mostly light to dark orange-brown with extensive black colouration between and surrounding the ocelli, sometimes meeting medial margins of eyes, a triangular orange-brown fascia with a base on the posterior border narrows to an apex midway between the ocelli; supra-antennal plates black, edged dull orange-brown around base of antennae; ocelli dull red; eyes dull red to light brown; postclypeus orange-brown dorsally with a black horizontal stripe along the anterior margin, ventral surface black with an orange medial line and orange coloration over half of lateral margins; anteclypeus shiny black; lorum black, gena black; rostrum reaching anterior margins of hind coxae, dark brown proximally, becoming dark brown to black distally; antennae black.</p><p>Thorax mostly dark brown to black. Pronotum with a narrow orange-brown fascia along midline surrounded by black, resembling a vintage key lock; interior of pronotum orange to dark brown, with irregular black areas, especially along paramedian and lateral fissures and adjacent ridges, also bordering the lateral halves of the pronotal collar; pronotal collar black on anterior margin, orange-brown to dark brown over remainder. Mesonotum variably black with orange-brown intrusions; submedian and lateral sigilla black, intervening depressions orange-brown to dark brown; lateral margins sometimes orange-brown; scutal depressions black; cruciform elevation orange-brown, variably black along midline and into anterior depression, lateral depressions striated black transversely; wing grooves mainly dark orange-brown and black. Metanotum dark brown to black; black central “dot” with orange-brown ridges. Thorax below mainly orange-brown or dark brown.</p><p>Legs brown with variable longitudinal orange-brown to black markings between specimens. Coxae about half orange-brown and black; fore femora with inner surface black but sometimes striped dark orange-brown, the outer surface predominantly dull orange-brown, femoral spines dark brown basally and black tipped, upright; mid and hind femora dark brown to black anteriorly, paler orange-brown posteriorly; fore tibiae dark brown; mid and hind tibiae orange-brown; tarsi orange-brown to brown; claws black; meracantha variably black on base around medial half, remainder pale cream-brown, reaching anterior margin of opercula.</p><p>Wings with forewing costal margin orange-brown to dark orange-brown; venation generally dark brown to orange-brown; basal cell translucent, tinged orange; basal membranes orange; smoky infusions throughout a1–a8; infuscations of variable intensity along crossveins r and r-m on the majority of individuals. Hind wing venation dark brown; v-shaped plagas tinged orange-grey visible along veins 2a and 3a and within anal cells.</p><p>Opercula (Plate 2D) variably black basally, plates pale yellow-brown, spatulate, curved medially, flexed ventrally, not meeting; approximately 1.5x larger than adjacent hind coxae.</p><p>Timbals (Plate 2C) with five long ribs; ribs 1-4 attached to basal spur, separated ventrally and spanning the timbal membrane, rib 5 unattached, extending over central half of membrane; timbal plate with well developed, elongate dome bearing a distinct point of apodeme attachment near centre.</p><p>Abdomen (Plate 3C–D) with tergite 1 orange centrally, with symmetrical, black diagonal striations posteriorly, black laterally; tergite 2 variably black along anterior third and laterally, becoming orange posteriorly, curved around outer margin of exposed timbals; tergites 3–7 predominantly pale orange-brown each with a distinct black mark on dorsal midline, this mark broadest on tergite 3 and equivalently narrower on tergites 4–7, tergites 3–5 sometimes also with a diffuse dark smudge on lateral sides; tergite 8 black over anterior half, tending dark orange-brown over remainder, with a black spot on lateral aspect; epipleurites orange, flexed ventrally. Sternite I orange-brown to black; sternite II orange with dark-brown to black areas medially and laterally; sternites III–VII pale orange-brown; sternite VIII darker orange-brown.</p><p>Genitalia (Fig. 6) with pygofer black, upper lobe of moderate size with a broadly rounded apex, in ventral view angled medially (Fig. 6A); basal pygofer lobe small weakly developed, broadly rounded; median lobe of uncus enlarged, variably elongated with length around 1.0– 1.4mm, concave ventrally, in lateral view narrow basally, expanded around mid-length to its widest width, thereafter evenly tapering to a rounded apex with ratio of length from base divided by widest width ranging from 2.6–4.5; claspers robust, claw-like in lateral view (Fig. 6B), partially meeting, in ventral view their inner margins diverging with a “notch” around aedeagus and their outer margins gently turned outwards to a bluntly pointed apex; aedeagus recurved distally through 180 degrees with ornamentation distally from the recurvature, the vesica broadened and weakly toothed either side towards apically, apex terminating in a short, slender ‘beak’ pointing outwards, downturned&gt;45 o viewed laterally, with a pair of very small flat ear-like flanges directed medially and minutely toothed; a “viking helmet” shape viewed laterally; pseudoparameres very long, reaching the distal recurvature of the aedeagus, very slender, hair-like, each with a rounded apical knob (Figs. 6C–D).</p><p>Description of adult female (Plate 4B, 5E–F, H). Head, thorax, wings and legs similar to male.</p><p>Abdomen. Tergite 1 orange, black laterally along posterior margin; tergite 2 orange suffused variably with black coloration, most prominent along posterior margin, a black spot on each lateral side; tergites 3–7 predominantly dark orange-brown each with a distinct black “double triangle” on dorsal midline, broadest on each anterior margin, narrowing centrally to an apex and joined to the apex of a posterior triangle with its base on the posterior margin of each tergite, most pronounced on tergites 3–5, reducing on successive tergite 4–7; a distinct black marking centrally located on each ventrolateral aspect; tergite 8 black over anterior half, tending dark orange-brown over remainder, with a black spot on lateral aspect. Sternite I black; sternites II–VII pale orange-brown with black infusion along midline; sternite VIII darker orange-brown; abdominal segment 9 dark ochraceous over dorsal two-thirds, orange-brown ventrally, dorsal beak black, a distinct black longitudinal marking either side of dorsal midline extending laterally along the anterior margin; a small black spot laterally near the posterior apex; ovipositor sheath dark brown, black posteriorly, ovipositor dark brown, becoming black posteriorly, extending &lt;1 mm beyond abdominal segment 9.</p><p>Measurements (in mm; range with mean in parentheses for 26 males and 10 females, including smallest and largest specimens). Length of body including head: male, 18.9–24.2 (22.5); female, 20.2–25.1 (23.7). Length of forewing: male, 25.0–29.6 (27.8); female, 26.1–32.4 (29.9). Width of forewing: male, 7.8–9.6 (9.1); female, 8.3–9.9 (9.5). Width of head (including eyes): male, 5.3–6.5 (6.1); female, 5.9–7.5 (6.7). Width of pronotum (across lateral angles): male, 5.7–6.5 (6.2); female, 6.2–7.6 (6.8). Width of abdomen: male, 5.6–6.6 (6.3); female, 5.7–6.8 (6.4). Length of ovipositor: female, 6.3–8.6 (7.5).</p><p>Morphological variation within and between populations. The morphological appearances of  Y. psammitica sp. nov. specimens are consistent within and between the populations around Clandulla, Cassilis and the Warrumbungle NPk. However, males from the northern sites (Warrumbungle NPk and Carnarvon Range, Queensland) have a smaller body length (BL; 18.9–22.4 mm; n=11) than those from Clandulla and Cassilis (BL&gt; 22.4 mm; n=15). Females from the northern sites are also substantially smaller (BL 20.2–22.9 mm; n=3) than those from the two southernmost populations (BL&gt; 24.1 mm; n=7).</p><p>Morphological distinguishing features. Male specimens of  Y. psammitica sp. nov. most closely resemble  Y. corindi,  Y. humphreyae,  Y. ignita,  Y. robertsonae and  Y. robusta,  Y. darug sp. nov. and  Y. fumea sp. nov., which are all similar in size and have brown hind wing plagas. They can be distinguished by the following combination of characters: (1) median lobe of uncus well developed but not particularly elongate (extending ventrally to the same extent as claspers; Fig 6B), (2) fore wings with a weak to moderate smoky infusion over distal apical cells, and (3) fore wing length&gt; 26 mm. Notably, in the Warrumbungles, male specimens of  Y. psammitica sp. nov. can only be confidently differentiated from cohabiting  Y. humphreyae males by their shorter median lobe of the uncus. Females can be distinguished from all other  Yoyetta species apart from  Y. fumea sp. nov. by having (1) fore wings with a weak to moderate smoky infusion over distal apical cells, (2) ventral side of abdominal segment 9 orange-brown, (3) ovipositor extending &lt;1 mm beyond abdominal segment 9 and (4) more pronounced black markings along the midline of the sternites. Females of  Y. psammitica sp. nov. can be distinguished from  Y. fumea sp. nov. by their slightly longer ovipositor, which extends &lt;1 mm beyond the apex of abdominal segment 9 (cf. not extending noticeably beyond the anal styles in  Y. fumea sp. nov.).</p><p>Calling song (Figs. 7–9). The male calling song of  Y. psammitica sp. nov. has three modes. The first mode (“clicking” mode) comprises a series of repetitive clicking, which is produced whilst in flight and occasionally when stationary on the trunk or branches of  Eucalyptus species. The second mode is the same as the first mode but includes a short soft buzz after each click (“click and buzz” mode) and only occurs while stationary. The third mode is a variant of the second mode that commences with a buzz followed by a series of clicks (“buzz and click” mode; Fig. 9).</p><p>Closer examination of the clicking mode (all statistics, n =9 recordings from the Warrumbungles, Mudgee and Clandulla localities) shows each click is a syllable as illustrated in Figs 7–9. Both in flight and at rest, each short syllable (0.012 – 0.023 s, 8–9 pulses, sometimes with 1-2 preceding soft pulses) is separated by a gap of roughly half a second in duration.</p><p>The click and buzz mode is illustrated in Fig. 8, and is described as follows (all statistics, n =5 recordings from the type locality). Following a syllable that is equivalent to that in the clicking mode is a short gap (0.04– 0.16 s), then a short soft buzz (0.07– 0.18 s), followed by a longer gap (0.32– 0.50 s). The third “buzz and click” mode is illustrated in Fig 9 (n =1) has only been recorded in the Warrumbungles, though it has also been observed at Napoleon Reef near Bathurst. The song begins with a long buzz (8.0 s) followed by a series of short syllables.</p><p>The frequency plateau is consistent between modes. It ranges broadly from approximately 6.5 to 12.5 kHz, with a dominant frequency between 7 and 11.5 kHz.</p><p>Y. humphreyae co-occurs with  Y. psammitica sp. nov., but  Y. humphreyae produces fewer (5–6) pulses (cf. 8–9 pulses in  Y. psammitica sp. nov.). The song is also close to  Y. corindi (which lacks the “long buzz” component) and the clicking structure may be indistinguishable from  Y. darug sp. nov., although these species do not overlap in geographical distribution with  Y. psammitica sp. nov.</p></div>	https://treatment.plazi.org/id/03D60217CC42BA7617876CAEFD6727CE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Emery, David L.;Emery, Nathan J.;Popple, Lindsay W.	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC5DBA7B1787682BFF522562.text	03D60217CC5DBA7B1787682BFF522562.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yoyetta fumea Emery & Emery & Popple 2025	<div><p>Yoyetta fumea sp. nov.</p><p>(Plates 2E–F, 3E, 4C, 6; Figs. 1, 2E–F, 10–12)</p><p>Types.   Holotype 1♂, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.77444&amp;materialsCitation.latitude=-35.09611" title="Search Plazi for locations around (long 149.77444/lat -35.09611)">Cullala Rd</a>, Lower Broughton, 35 o 05’46’’S 149 o 46’28’’E, 10.i.2021, N., C. &amp; D. Emery (EME0717-037) (K. 624937, AM).</p><p>Paratypes. NEW SOUTH WALES: 1♂, Jacqua Rd., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.90805&amp;materialsCitation.latitude=-34.96417" title="Search Plazi for locations around (long 149.90805/lat -34.96417)">Windellama</a>, 34 o 57’51’’S 149 o 54’29’’E, 10.i.2021, N., C. &amp; D. Emery (EME0717-027) ;   1♀, Jerrara Rd., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.9611&amp;materialsCitation.latitude=-34.786945" title="Search Plazi for locations around (long 149.9611/lat -34.786945)">Bungonia</a>, 34 o 47’13’’S 149 o 57’40’’E, 9.i.2023, C. &amp; D. Emery (EME0717-039) (K. 624938–939, AM) ;   2♂, Yarrawal Rd., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.91695&amp;materialsCitation.latitude=-34.973335" title="Search Plazi for locations around (long 149.91695/lat -34.973335)">Windellama</a>, NSW, 34 o 58’24’’S 149 o 55’01’’E, 4.i.2019, S. &amp; T. Corbin, D. Emery (EME0717-025 – 026; genitalia prep 717-2) ;   1♂ 4♀, Jacqua Rd., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.90805&amp;materialsCitation.latitude=-34.96417" title="Search Plazi for locations around (long 149.90805/lat -34.96417)">Windellama</a>, 34 o 57’51’’S 149 o 54’29’’E, 10.i.2021, N., C. &amp; D. Emery (EME0717-028 – 033) ;  3♂ 1♀, same data as holotype (EME0717-034 – 036; genitalia preps 717-3, 717-4);   1♀, Jerrara Rd., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.9611&amp;materialsCitation.latitude=-34.786945" title="Search Plazi for locations around (long 149.9611/lat -34.786945)">Bungonia</a>, 34 o 47’13’’S 149 o 57’40’’E, 9.i.2023, C. &amp; D. Emery (EME0717-042) (DE) ;  1♂, same location as holotype, 30.xi.2013, N. &amp; D. Emery;   1♂, Yarrawal Rd., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.91695&amp;materialsCitation.latitude=-34.973335" title="Search Plazi for locations around (long 149.91695/lat -34.973335)">Windellama</a>, NSW, 34 o 58’24’’S 149 o 55’01’’E, 4.i.2019, S. &amp; T. Corbin, D. Emery ;   1♂, Jacqua Rd, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.90805&amp;materialsCitation.latitude=-34.96417" title="Search Plazi for locations around (long 149.90805/lat -34.96417)">Windellama</a>, 34 o 57’51’’S 149 o 54’29’’E, 10.i.2021, N., C. &amp; D. Emery (LWP) ;   2♂, Jacqua Rd., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.90805&amp;materialsCitation.latitude=-34.96417" title="Search Plazi for locations around (long 149.90805/lat -34.96417)">Windellama</a>, 34 o 57’51’’S 149 o 54’29’’E, 10.i.2021, N., C. &amp; D. Emery (EME0717-040 – 041) ;   2♂, Jerrara Rd. 600m north of Tickner Valley Rd., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=149.96056&amp;materialsCitation.latitude=-34.78528" title="Search Plazi for locations around (long 149.96056/lat -34.78528)">Marulan</a>, NSW, 34°47’07”S 149°57’38”E, 28.xii.2022, J. Poyitt, (HEMC 717.001,002) (JP)  .</p><p>Distribution, habitat and seasonality. Specimens of  Yoyetta fumea sp. nov. are currently known only from around Windellama and Tarago in the Southern Highlands (Fig. 1). Individuals mainly favour the trunks of black she-oaks ( Allocasuarina littoralis), where they call while stationary or moving around the trunk at heights of 2–5 m from the ground. Around Windellama, the black and drooping she-oaks ( Allocasuarina littoralis and  A. verticillata, respectively), are the main floral preferences (Figs 2E–F). This is a notable difference between this species and the closely related  Y. humphreyae, which is primarily associated with eucalypts. Adults have been found between November and January.</p><p>Etymology. The name  “ fumea ” (Latin, adjective, feminine) reflects the prominent smoky infusion of the apical cells of the forewing, which is more pronounced in this species than in closely related  Yoyetta species.</p><p>Description of adult male (Plates 2E–F, 3E, 4C, 6; Fig. 10).</p><p>Head slightly narrower than the lateral margins of pronotum; extensively black with orange brown fascia along posterior half of midline, between ocelli and eyes and surrounding pedicles; supra-antennal plates black in all specimens, occasionally edged dull orange-brown; ocelli pale pink to yellow; eyes dark brown; postclypeus dark orange-brown dorsally, sometimes with black infusions along the midline, anterior margin and ventral areas black, with lateral margins sometimes dark orange-brown, transverse ridges black, lorum black, gena black; anteclypeus mainly black, sometimes smudged orange-black anteriorly; rostrum reaching bases of hind coxae, brown basally becoming dark brown or black apically; antennae black.</p><p>Thorax black with irregular orange-brown marking areas, surrounding paramedian and lateral fissures; pronotal collar black, sometimes with a central orange-brown “V” with apex anterior and extending into pronotum anteriorly. Mesonotum mainly black; submedian and lateral sigilla black; scutal depressions black; cruciform elevation black with dark orange infusion with faint black transverse striations in the lateral depressions, the symmetrical orange-brown markings extending forward along anterior ridges and ending lateral to the scutal depressions; wing grooves mainly dark orange-brown and black. Metanotum black with orange-brown central section. Thorax below mainly black with variable orange markings.</p><p>Legs dark brown to black markings variable between individuals. Coxae mainly black, coxal membranes bright orange-red, trochanter black; fore femora with inner surface usually black, the outer surface predominantly dull orange-brown, femoral spines black, erect; mid and hind femora striped black on anterior sides and paler brown on posterior; fore tibiae black anteriorly, dark brown posteriorly; mid and hind tibiae dark orange-brown; tarsi orange-brown or brown suffused with black; claws dark orange-brown, becoming black at tips; meracantha black on basal half, cream over remainder, not reaching opercula.</p><p>Wings with forewing costal margin black to dark orange-brown; venation generally dark brown to orange-brown; basal cell translucent, tinged green-orange; basal membranes orange; smoky infusions throughout a1–a8; infuscations of variable intensity along crossveins r and r-m. Hind wing venation dark brown; v-shaped plagas tinged smoky-grey visible along veins 2a and 3a and within anal cells.</p><p>Opercula (Plate 2F), pale black basally, plates dull cream-brown, spatulate, curved medially, flexed ventrally, not meeting; approximately 1.5x larger than adjacent hind coxae.</p><p>Timbals (Plate 2E) with five long ribs; ribs 1–4 attached to basal spur, separated ventrally and spanning the timbal membrane, rib 5 short, unattached, occupying dorsal central half of timbal only; timbal plate with well developed, prominent elongate and elevated dome bearing a distinct point of apodeme attachment near centre.</p><p>Abdomen with tergite 1 black, posterior margins dark orange anterior to timbal cavity; tergite 2 predominantly black with dark orange spot either side of posterior midline, sometimes meeting along posterior margin; tergites 3–7 predominantly pale orange-brown each with a narrow black mark on dorsal midline, this mark being broadest on tergite 3 and narrowing slightly on each subsequent tergite posteriorly, also with a diffuse dark smudge on lateral sides resolving as a black dot on the postero-ventral aspect; tergite 8 black, tending dark orange-brown towards posterior margin, with a distinct dark orange marking on the antero-lateral aspect (Plate 3E); epipleurites flexed ventrally, a small black dot on the posterior margins. Sternite I black, orange-brown posteriorly; sternite II orange with dark-brown to black areas medially and laterally; sternites III–VII pale orange-brown, diffuse, with light black infusions on lateral halves; sternite VIII mainly orange-brown, dark brown at base, paler brown at apex.</p><p>Genitalia (Fig. 10) with pygofer upper lobe of moderate size, gradually tapering to a peaked but rounded apex, in ventral view projecting outwards; basal pygofer lobe small and weakly developed, broadly rounded (Fig 10A); median lobe of uncus enlarged, long, in lateral view (Fig 10B) almost uniformly wide along length (1.6–2 mm), evenly tapering to a rounded apex, with ratio of length from base divided by widest width around 3.0, extending distally beyond the claspers, although shorter than the distance between the dorsal beak and upper pygofer lobe; claspers robust, claw-like, partially meeting, in ventral view their bases closely applied to aedeagus, with inner margins curving laterally and their outer margins turned outwards to a blunted pointed apex; aedeagus recurved distally through 180 degrees, the vesica consistently toothed either side from the curvature, apex terminating in a short, slender ‘beak’ pointing outwards and a pair of very small flat ear-like flanges directed medially and minutely toothed; a “viking helmet” shape viewed laterally, with apex either straight or slightly downturned (&lt;30 o dorsally) in lateral view; pseudoparameres very long, reaching the distal recurvature of the aedeagus, very slender, each with a rounded apical knob (Fig 10C–D).</p><p>Description of adult female (Plates 4C, 6C–D, F). Head mainly similar to male, larger amounts of orange-brown areas between ocelli and eyes, a black spot on the posterio-medial margin of the eye; ocelli colour ranging from bright red to dull pink; postclypeus dark orange-brown dorsally, sometimes with black infusions along the midline, anterior margin black, ventral aspect black in most specimens, lateral margins dark orange-brown, lighter specimens with orange-brown midline, transverse ridges black, lorum black, gena black.</p><p>Thorax and legs similar to male, with coloration ranging from mainly black to principally dark orange-brown.</p><p>Wings similar to male, smoky infusions and cross-vein infuscations slightly lighter. Basal membranes either bright orange in orange-brown specimens or muddy orange in darker individuals.</p><p>Abdomen. (Plates 4C, 7C–D). Tergite 1 orange, black laterally along posterior margin, a central black spot on darker specimens; tergite 2 orange black over anterior third, orange to dark orange-brown over remainder, black markings around lateral extremity; tergite 3 with black coloration either restricted to the anterior margin or diffusely broadened laterally, a narrow black mark on dorsal midline, wider on darker females; tergites 4–7 predominantly dark orange-brown each with a distinct black “parallelogram” on dorsal midline, broadest on each anterior margin and widest on darker specimens, most pronounced on tergites 3–5, reducing slightly in width on successive tergites 4–7, variable black smudging laterally on each tergite, culminating in a darker black marking on the posterior half of the lateral margins; tergite 8 black over anterior half, more extensively black in darker individuals, tending dark orange-brown over remainder, with a black spot on lateral aspect. Sternite I black; sternites II–VII pale orange-brown with very faint black smudging along midline; sternite VIII darker orange-brown, a black spot either side of posterior midline; abdominal segment 9 dark ochraceous over dorsal two-thirds, darker brown ventrally, dorsal beak black, in darker specimens a black line extends from the dorsal beak anteriorly for a short distance along midline, a distinct black longitudinal marking either side of dorsal midline extends laterally along the anterior margin, a small black spot laterally near the posterior apex; ovipositor sheath dark brown, black posteriorly; ovipositor dark brown, becoming black posteriorly, not extending noticeably beyond the anal styles.</p><p>Measurements (in mm; range with mean in parentheses for 12 males and 5 females, including smallest and largest specimens). Length of body including head: male, 21.3–22.4 (21.9); female, 20.7–23.3 (21.4). Length of forewing: male, 22.8–28.6 (27.0); female, 27.7–29.6 (28.3). Width of forewing: male, 8.5–9.8 (9.0); female, 8.9–9.7 (9.2). Width of head (including eyes): male, 5.5–6.2 (5.9); female, 6.3–6.8 (6.5). Width of pronotum (across lateral angles): male, 5.3–6.0 (5.7); female, 6.0–6.5 (6.3). Width of abdomen: male, 5.4–6.2 (5.9); female, 5.5–6.1 (5.9). Length of ovipositor: female, 5.5–7.5 (6.1).</p><p>Morphological variation within and between populations. Specimens of this species are consistent morphologically, likely reflecting their current limited distribution.</p><p>Morphological distinguishing features. The presence of infuscations on fore wing crossveins r and r-m is a trait that consistently distinguishes  Y. fumea sp. nov. from all other described species of  Yoyetta, apart from  Y. psammitica sp. nov. and  Y. humphreyae . Males of  Y. fumea sp. nov. can be distinguished from  Y. psammitica sp. nov. by having a more elongate median uncal lobe that extends posteriorly beyond the claspers (a feature that is visible without dissection). Notably, the median lobe of the uncus in  Y. fumea sp. nov. is shorter than the distance between the dorsal beak and the apex of upper pygofer lobe, a feature that distinguishes it not only from  Y. humphreyae, but also from other species that possess a more elongate uncus, including  Y. darug sp. nov. and  Y. corindi (the uncus in these three species is about as long as the distance between the dorsal beak and apex of the pygofer lobe). Females of  Y. fumea sp. nov. can be distinguished from  Y. psammitica sp. nov. by (1) their shorter ovipositor, which does not extend conspicuously beyond the anal styles (cf. extending &lt;0.5 mm beyond the anal styles in  Y. psammitica sp. nov.), and (2) their lack of distinct black markings along the midline of the tergites.</p><p>Calling song. (Figs. 11–12). The male calling song of  Y. fumea sp. nov. has two modes. One mode (combination mode) has a combination of buzzes and clicks. The second mode (clicking mode) comprises only clicks. Both modes can be produced when the male is stationary; however, only the clicking mode is produced in flight. Both modes can be produced for an extended duration&gt;60 seconds.</p><p>The combination mode contains phrases that have two variants, which can be produced in an alternating fashion by the same male as Fig. 11 illustrates.This mode and its variants are described as follows (all statistics, n =7 recordings from Windellama, Clandulla and the type locality). Each phrase begins with an echeme typically between 0.315 and 0.829 s (but can be maintained for at least 50 s duration) that increases in amplitude as it progresses, followed by a short gap (0.05 s). After this is either a repeated series of clicking and short buzzing (variant 1) or clicking (variant 2; similar to the clicking mode as shown in Fig.12). Further analysis of variant 1 reveals a alternating series of short syllables (0.010 –0.014 s) and short macrosyllables (0.097 –0.147 s). This may be repeated up to 4 times before the phrase finishes with an echeme (0.154 –0.242 s). In one recording from Windellama, the phrase was extended to include an additional short gap (0.123 s) followed by an echeme (0.395 s) that increased in amplitude, then a longer gap (0.345 s), and finishing with a short echeme (0.142 s) (Figure 11C). Further recordings would be required to determine how often these extended phrases are produced. For variant 2, each click is a syllable that is equivalent in structure and duration to those in variant 1; but each syllable is separated by a gap (0.205 –0.292 s). In available recordings (n= 4), males produced between 6 and 23 syllables (in succession) in variant 2.</p><p>The clicking mode consists of repeated single syllables as illustrated in Fig 12 and are equivalent in duration to the syllables in the combination mode. A gap of varying duration (0.135 – 0.275 s) punctuates each syllable (all statistics, n =3 from the Windellama locality). When producing this mode, males fly between trees at canopy height making it difficult for acoustic recording, and especially difficult to isolate the song from other co-occurring cicada calling songs. It is hypothesized that females respond with wing-flicks following each syllable in this mode as has been recorded for other  Yoyetta species.</p><p>The frequency plateau is generally consistent between modes. It ranges broadly from approximately 5.5 to 12.5 kHz, with a dominant frequency between 7 and 11 kHz.</p><p>The calling song and singing behaviour of  Y. fumea sp. nov. is closest to  Y. humphreyae in terms of complexity and structure.  Y. fumea sp. nov. produces syllables that often contain more pulses (6–7 pulses, cf. 5–6 pulses in  Y. humphreyae). It has also been noted in the field that the singing behaviour of  Y. humphreyae and  Y. fumea sp. nov. differs, with the latter producing more complex alternating click and buzz variants, as described above. In comparison, although complex, the call of  Y. humphreyae has a more predictable sequence as illustrated by Moulds and Popple (2021).</p><p>The song is also similar to those of  Y. corindi (illustrated in Popple and Emery, 2022) and  Y. psammitica sp. nov. The long buzz produced by  Y. fumea sp. nov. has not been recorded in the song of  Y. corindi and the two species are not known to co-occur.  Y. psammitica sp. nov. produces clicks with 8–9 pulses (cf. 6–7 pulses in  Y. fumea sp. nov.).</p></div>	https://treatment.plazi.org/id/03D60217CC5DBA7B1787682BFF522562	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Emery, David L.;Emery, Nathan J.;Popple, Lindsay W.	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC55BA7817876917FB9C2781.text	03D60217CC55BA7817876917FB9C2781.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Yoyetta Moulds 2012	<div><p>Morphological key to species of  Yoyetta cicadas</p><p>Specimens must be set with wings spread to begin using this key.A vernier calliper is required to check measurements. Male specimens must often be examined microscopically and may require dissection in some instances.</p><p>Key to males</p><p>1. Hind wing plagas white................................................................................ 2</p><p>- Hind wing plagas pale brownish, pinkish or grey........................................................... 19</p><p>2. Abdominal tergites 2–8 and sternites II–VII uniformly black or brown without orange-brown or yellow-brown markings; intersegmental membranes dark and inconspicuous.......................................................... 3</p><p>- Abdominal tergites 2–8 mainly black with contrasting orange-brown or yellow-brown markings, or contrasting intersegmental membranes; sternites II–VII mainly yellow-brown or orange................................................... 5</p><p>3. Fore wing basal membranes orange or pale orange-white...............................................  Y. timothyi</p><p>- Fore wing basal membranes red.......................................................................... 4</p><p>4. When viewed from ventral side, tip of aedeagus strongly bifurcate, with apical arms splayed laterally, sometimes in a weak “v” shape, at an angle of 150–180 degrees.............................................................  Y. denisoni</p><p>- When viewed from ventral side, tip of aedeagus undivided or weakly divided, without distinct lateral arms......  Y. kershawi</p><p>5. Tergites 5–7 partly black with contrasting, orange or yellow markings........................................... 6</p><p>- Tergites 5–7 entirely dark brown to black with yellow or orange coloration restricted to the intersegmental membranes.... 11</p><p>6. Opercula mainly pale grey to dark grey-brown.............................................................. 7</p><p>- Opercula mainly red or orange.......................................................................... 10</p><p>7. Dorsal surface of head, pronotum and mesonotum typically covered with dense gold pubescence; fore wing basal membranes orange or pink........................................................................................ 8</p><p>- Head, pronotum and mesonotum not covered in conspicuous pubescence; fore wing basal membranes pale grey.......... 9</p><p>8. Hind wing plagas narrow on ac2(v) (along vein 3); apex of theca blunt and club-like.....................  Y. abdominalis</p><p>- Hind wing plagas broad on ac2 (along vein 3), though not quite as broad as on jugum; apex of theca tapered and angled 30° ventrally....................................................................................  Y. loftyensis</p><p>9. Lateral sides of tergites 3–7 mainly orange; tergite 8 with orange markings on anterodorsal side.................  Y. aaede</p><p>- Lateral sides of tergites 3–7 mainly black (with yellow dorsolateral markings); anterodorsal side of tergite 8 black.....................................................................................................  Y. serrata</p><p>10. Body length &lt;23 mm; sternites III–VII bright reddish-orange without a continuous dark brown to black central marking...............................................................................................  Y. spectabilis</p><p>- Body length&gt; 23 mm; sternites II–VII yellowish-orange or reddish-brown with a continuous dark brown to black central marking.......................................................................................  Y. regalis</p><p>11. Fore wing length ≤ 26 mm ............................................................................. 12</p><p>- Fore wing length&gt; 26 mm ............................................................................. 14</p><p>12. Opercula mainly black on basal half, pale dull yellow over remainder; meracantha orange.....................  Y. incepta</p><p>- Opercula almost entirely pale reddish-brown or pale grey; meracantha pale brown................................. 13</p><p>13. Pronotum dark brown to black with contrasting yellow-brown central marking; fore wing basal membranes orange....................................................................................................  Y. electrica</p><p>- Pronotum dark brown to black without contrasting yellow-brown central marking; fore wing basal membranes grey to smoky orange....................................................................................  Y. hunterorum</p><p>14. Dorsal surface of head, pronotum and mesonotum with an inconspicuous, sparse covering of pubescence; lateral depressions adjacent to cruciform elevation conspicuous, brown to pale brown.......................................  Y. grandis</p><p>- Dorsal surface of head, pronotum and mesonotum typically with dense, black pubescence; lateral depressions adjacent to cruciform elevation inconspicuous, typically dark brown to black.............................................. 15</p><p>15. Hind wing plagas broad; with opaque white area expressed equally in cells ac3 and ac2(v) (either side of the jugal fold)...............................................................................................  Y. subalpina</p><p>- Hind wing plagas narrow or predominantly expressed in ac3, with a narrow extent on ac2(v) (along vein 3a)............ 16</p><p>16. Tergite 8 black with a brown, reddish-brown or dark reddish-brown posterior lateral marking on each side; apex of theca distinctly club-shaped; transparent flange along margin of recurvature not quite as broad as thecal shaft...........  Y. regalis</p><p>- Tergite 8 entirely dark brown to black without posterior lateral markings; apex of theca not club-shaped but narrow with ornamentation; transparent flange along margin of recurvature much broader than thecal shaft....................... 17</p><p>17. Hind wing plagas broad and occupying the jugum...................................................  Y. douglasi</p><p>- Hind wing plagas narrow with jugum mainly hyaline........................................................ 18</p><p>18. Fore wings conspicuously broad, with bend in ambient vein about equal to bend in costa; sternites III–VII typically with dark brown to black central markings.................................................................  Y. ngarabal</p><p>- Fore wings not conspicuously broad, with bend in ambient vein more pronounced than bend in costa; sternites III–VII usually without dark brown to black central markings........................................................  Y. verrens</p><p>19. Median lobe of uncus long, protruding conspicuously further ventrally than the claspers............................ 20</p><p>- Median lobe of uncus short, not protruding conspicuously further ventrally than the claspers......................... 23</p><p>20. Abdomen with sternite VIII entirely orange-brown or light orange brown.............................  Y. darug sp. nov.</p><p>- abdomen with sternite VIII dark brown to black at base grading to orange brown, becoming paler towards apex......... 21</p><p>21. Fore wings with weak to moderate infuscations along crossveins r and r-m; median lobe of uncus shorter than distance between dorsal beak and apex of upper pygofer lobe....................................................  Y. fumea sp. nov.</p><p>- Fore wings with or without infuscations along crossveins r and r-m (a weak infuscation may be present); median lobe of uncus about as long as the distance between the apices of the upper pygofer lobe and dorsal beak.......................... 22</p><p>22. Median lobe of uncus narrow at base, widest around mid length and tapering towards apex................  Y. humphreyae</p><p>- Median lobe of uncus narrow, slightly expanding distally, with rounded apex...............................  Y. corindi</p><p>23. Head width &lt;5 mm ................................................................................... 24</p><p>- Head width ≥ 5 mm ................................................................................... 33</p><p>24. Thorax mainly brown................................................................................. 25</p><p>- Thorax mainly black.................................................................................. 27</p><p>25. Tergite 1 pale brown; pronotum mainly pale brown......................................................  Y. celis</p><p>- Tergite 1 black or dark brown; pronotum mainly brown or black............................................... 26</p><p>26. Fore wing length&gt; 20 mm; cruciform elevation and surrounding areas adjacent to wing grooves all much paler and brighter than remainder of thorax..........................................................................  Y. tristrigata</p><p>- Fore wing length &lt;20 mm; cruciform elevation and surrounding areas adjacent to wing grooves not all distinctly paler than remainder of thorax...........................................................................  Y. fluviatilis</p><p>27. Pronotum mainly brown, contrasting with mesonotum..................................  Y. tristrigata (dark specimen)</p><p>- Pronotum mainly dark brown to black, not distinctly paler than mesonotum...................................... 28</p><p>28. Lateral edges of tergites orange, with dark markings along posterior margin (broad or narrow, often diffuse)............ 29</p><p>- Lateral edges of tergites orange, without dark markings on posterior margin...................................... 30</p><p>29. Fore wing length usually&gt; 20 mm; theca recurved distally through 180 degrees; apex weakly upturned..........  Y. delicata</p><p>- Fore wing length &lt;20 mm; theca recurved distally through 120 degrees; apex downturned....................  Y. fluviatilis</p><p>30. Theca recurved distally through 180 degrees; apex straight or weakly upturned................................... 31</p><p>- Theca recurved distally through 120 degrees; apex straight or downturned....................................... 32</p><p>31. Fore wing length typically&gt; 20 mm; apex of theca weakly upturned......................................  Y. delicata</p><p>- Fore wing length &lt;20 mm; apex of theca straight................................................  Y. landsboroughi</p><p>32. Fore wing length&gt; 20 mm; timbal long rib 4 appearing discontinuous, with dorsal and ventral areas connected by only a very narrow medial area........................................................................  Y. nigrimontana</p><p>- Fore wing length &lt;20 mm; timbal long rib 4 clearly continuous, narrowing only slightly in medial area.........  Y. fluviatilis</p><p>33. Timbal with four long ribs; fourth (anteriormost) long rib fused to the anterior cuticle..........................  Y. ignita</p><p>- Timbal with five long ribs, all separate from anterior cuticle and surrounded by pale timbal membrane................. 34</p><p>34. Timbal long rib 5 distinctly narrower than widest portion of adjacent long rib 4............................  Y. repetens</p><p>- Timbal long rib 5 about equal in width to adjacent long rib 4.................................................. 35</p><p>35. Thorax mainly brown to dark brown..................................................................... 36</p><p>- Thorax mainly black.................................................................................. 40</p><p>36. Fore wing length&gt; 26 mm ............................................................................. 37</p><p>- Fore wing length ≤ 26 mm ............................................................................. 38</p><p>37. Fore wings hyaline, without smoky infusions or infuscations............................................  Y. robusta</p><p>- Fore wings with smoky infusion over distal apical cells, often with an infuscation present on crossveins r and r-m............................................................................................  Y. psammitica sp. nov.</p><p>38. Cruciform elevation and surrounding areas adjacent to wing grooves all much paler and brighter than remainder of thorax.............................................................................................  Y. tristrigata</p><p>- Cruciform elevation and surrounding areas adjacent to wing grooves not all distinctly paler than remainder of thorax..... 39</p><p>39. Length of body typically&gt; 18 mm; tergite 2 black medially.........................................  Y. cumberlandi</p><p>- Length of body &lt;18 mm; tergite 2 orange with black restricted to anterior margin..............................  Y. celis</p><p>40. Tergite 2 mainly orange..........................................................  Y. tristrigata (dark specimen)</p><p>- Tergite 2 mainly black................................................................................ 41</p><p>41. Opercula relatively small, each &lt;1.9 mm wide.......................................................  Y. delicata</p><p>- Opercula not relatively small, each ≥2.0 mm wide.......................................................... 42</p><p>42. Apex of theca very short lanceolate with a pair of upright basal lobes........................................... 43</p><p>- Apex of theca produced as a long spine with ornamentation and without a pair of upright basal lobes..........  Y. australicta</p><p>43. Pronotal collar brown; tergites 3–7 yellow-brown with black markings.......................  Y. robusta (dark specimen)</p><p>- Pronotal collar dark brown to black; tergites 3–7 orange with black markings............................  Y. robertsonae</p><p>Key to females (excluding  Y. enigmatica for which the female is unknown)</p><p>1. Hind wing plagas white................................................................................ 2</p><p>- Hind wing plagas cream, brown or orange to pink.......................................................... 17</p><p>2. Ovipositor sheath extends &lt;1 mm beyond apex of abdominal segment 9.......................................... 3</p><p>- Ovipositor sheath extends ≥ 1 mm beyond apex of abdominal segment 9......................................... 13</p><p>3. Pronotum mainly black or dark brown with a contrasting yellow-brown central marking............................. 4</p><p>- Pronotum mainly black with central marking inconspicuous or similar in colour to other parts of pronotum, or brown with or without a pale brown central marking..................................................................... 9</p><p>4. Head width &lt;6 mm .................................................................................... 5</p><p>- Head width&gt; 6 mm .................................................................................... 7</p><p>5. Tergites 6 and 7 orange anteriorly with broad black bands extending to posterior margins......................  Y. aaede</p><p>- Tergites 6 and 7 black or dark brown anteriorly and orange or orange-brown posteriorly............................. 6</p><p>6. Fore wing length&gt; 21 mm ......................................................................  Y. electrica</p><p>- Fore wing length &lt;21 mm .......................................................................  Y. incepta</p><p>7. Fore wings with basal membranes pale whitish-grey; tergite 8 mainly black, without a marking that is similar in shape to the yellow markings on tergites 5–7...................................................................  Y. serrata</p><p>- Fore wings with basal membranes orange, light orange or pink; tergite 8 with distinct orange lateral marking that is similar in shape to yellow markings on tergites 5–7.................................................................. 8</p><p>8. Dorsal surface of head, pronotum and mesonotum typically covered with dense gold pubescence...........  Y. abdominalis</p><p>- Dorsal surface of head, pronotum and mesonotum typically with sparse, black pubescence or without conspicuous pubescence...................................................................................  Y. loftensis</p><p>9. Fore wing length &lt;21 mm .......................................................................  Y. incepta</p><p>- Fore wing length&gt; 21 mm ............................................................................. 10</p><p>10. Tergites 5–7 black with distinctive, yellow triangular markings on dorsolateral sides, widening towards posterior margins.............................................................................................  Y. spectabilis</p><p>- Tergites 5–7 mainly brown to reddish-brown, or black....................................................... 11</p><p>11. Fore wing basal membranes red or pink.................................................  Y. denisoni or  Y. kershawi</p><p>- Fore wing basal membranes pale orange-white or orange..................................................... 12</p><p>12. Tergites 3–8 brown, a combination of brown (or reddish-brown) and black or entirely black...................  Y. timothyi</p><p>- Tergites 3–8 black with orange coloration on dorsolateral posterior margins, with the orange most conspicuous on tergite 6............................................................................................  Y. subalpina</p><p>13. Ovipositor sheath extends ≥ 2 mm beyond apex of abdominal segment 9......................................... 14</p><p>- Ovipositor sheath extends &lt;2 mm beyond apex of abdominal segment 9......................................... 16</p><p>14. Hind wing plagas narrow, restricted to basal margins of anal cell 3 and vein 2A.............................  Y. verrens</p><p>- Hind wing plagas broad, extending to cover almost the entire jugum............................................ 15</p><p>15. Ovipositor sheath extends approximately 3 mm beyond apex of abdominal segment 9.....................  Y. hunterorum</p><p>- Ovipositor sheath extends approximately 3 mm beyond apex of abdominal segment 9.......................  Y. douglasi</p><p>16. Fore wings distinctly angulated at costal nodes; costal veins mainly reddish-brown or orange-brown; dorsolateral sides of tergites mainly reddish-brown.....................................................................  Y. regalis</p><p>- Fore wings gradually curved at node; costal veins mainly dark brown; dorsolateral sides of tergites mainly dull brown to black........................................................................................  Y. grandis</p><p>17. Ovipositor sheath extends&gt; 2 mm beyond apex of abdominal segment 9..................................  Y. ngarabal</p><p>- Ovipositor sheath extends &lt;2 mm beyond apex of abdominal segment 9......................................... 18</p><p>18. Ovipositor sheath extends ≥ 1 mm beyond apex of abdominal segment 9......................................... 19</p><p>- Ovipositor sheath extends &lt;1 mm beyond apex of abdominal segment 9......................................... 20</p><p>19. Dorsal side of body mainly pale brown................................................................  Y. celis</p><p>- Dorsal side of body mainly brown............................................................  Y. darug sp. nov.</p><p>20. Thorax mainly black.................................................................................. 21</p><p>- Thorax mainly brown (with black sigilla)................................................................. 28</p><p>21. Fore wing length&gt; 27 mm ............................................................................. 22</p><p>- Fore wing length &lt;27 mm ............................................................................. 24</p><p>22. Tergites 3–7 predominantly black..............................................................  Y. robertsonae</p><p>- Tergites 3–7 predominantly orange...................................................................... 23</p><p>23. Specimen from South Australia .................................................  Y. australicta (larger specimens)</p><p>- Specimen from New South Wales ...................................................................  Y. ignita</p><p>24. Fore wing length &lt;22 mm ............................................................................. 25</p><p>- Fore wing length&gt; 22 mm ............................................................................. 27</p><p>25. Legs with fore and mid tibiae mainly dark ochraceous to black.....................................  Y. nigrimontana</p><p>- Legs with fore and mid tibiae brown to orange-brown........................................................26</p><p>26. Eyes clearly red or dull red......................................................................  Y. fluviatilis</p><p>- Eyes not clearly red or dull red; instead black, brown, grey or faded.....................  Y. landsboroughi,  Y. fluviatilis *</p><p>27. Head width &lt;5.2 mm ......................................................................  Y. nigrimontana</p><p>- Head width&gt; 5.2 mm .....................................................  Y. australicta,  Y. delicata,  Y. repetens *</p><p>28. Fore wings with weak to moderate infuscations along crossveins r and r-m....................................... 29</p><p>- Fore wings without infuscations along crossveins r and r-m................................................... 30</p><p>29. Ovipositor sheath extends conspicuously beyond anal styles..................................  Y. psammitica sp. nov. Ovipositor sheath does not extend noticeably beyond anal styles...................................  Y. fumea sp. nov.</p><p>30. Fore wings with a weak to moderate smoky infusion over distal apical cells...................................... 31</p><p>- Fore wings without a smoky infusion over distal apical cells.................................................. 32</p><p>31. Ventral side of abdominal segment 9 dark brown to black..........................................  Y. humphreyae</p><p>- Ventral side of abdominal segment 9 orange-brown.........................................  Y. psammitica sp. nov.</p><p>32. Sternites with dark triangular central markings (prominent or diffuse)........................................... 33</p><p>- Sternites without dark triangular central markings.......................................................... 34</p><p>33. Ovipositor sheath extends &lt;0.5 mm beyond apex of abdominal segment 9.................................  Y. robusta</p><p>- Ovipositor sheath extends 0.5 mm –&lt;1 mm beyond apex of abdominal segment 9..............  Y. corindi,  Y. humphreyae *</p><p>34. Ovipositor sheath extends 0.5 mm –&lt;1 mm beyond apex of abdominal segment 9...............  Y. corindi,  Y. humphreyae *</p><p>- Ovipositor sheath extends &lt;0.5 mm beyond apex of abdominal segment 9....................................... 35</p><p>35. Fore wing length&gt; 27 mm .........................................................................  Y. ignita</p><p>- Fore wing length &lt;27 mm ............................................................................. 36</p><p>36. Body length &lt;17 mm, fore wing length &lt;21 mm .....................................................  Y. fluviatilis</p><p>- Body length&gt; 17 mm, fore wing length&gt;21............................................................... 37</p><p>37. Thorax with cruciform elevation and surrounding areas adjacent to wing grooves distinctly paler and brighter than remainder of mesonotum...............................................................................  Y. tristrigata</p><p>- Thorax with cruciform elevation and surrounding areas adjacent to wing grooves not noticeably paler than remainder of mesonotum......................................................................................... 38</p><p>38. Abdominal tergites 3–7 with black lateral markings clearly defined...................................  Y. cumberlandi</p><p>- Abdominal tergites 3–7 with dull brown lateral markings diffuse and faded...............................  Y. repetens</p><p>* These species cannot be reliably differentiated with female specimens alone. Male specimens are required. In some cases, checking information on geographical distributions may be helpful.</p></div>	https://treatment.plazi.org/id/03D60217CC55BA7817876917FB9C2781	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Emery, David L.;Emery, Nathan J.;Popple, Lindsay W.	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
