taxonID	type	description	language	source
03D60217CC48BA6517876987FD3D23BE.taxon	diagnosis	The diagnosis follows Moulds (2012) with modifications of Emery et al. (2015). Type species: Cicadetta celis Moulds, 1988.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC4BBA6F17876862FD66250E.taxon	description	(Plate 1; Plate 2 A – B; Plate 3 A – B; Plate 4 A; Figs. 1, 2 C – D, 3 – 5). Types. Holotype ♂, Silverdale NSW, 33 o 52 ’ 08 ’’ S 150 o 37 ’ 11 ’’ E, 14. xi. 2007, D. Emery (EME 0489 - 002) (K. 624928, AM). Paratypes. NEW SOUTH WALES. 1 ♂ 1 ♀, Bourke St., Parramatta, 33 ° 47 ’ 35 ’’ S 151 ° 00 ’ 00 ’’ E, 6. xii. 2022, D. Emery (EME 0489 - 017, - 018) (K. 624929 – 930, AM); 1 ♂, same location as holotype, 7. xi. 2007, D. Emery & L. Popple (EME 0489 - 001; genitalia prep 489 - 1); 1 ♂, Castlereagh Nature Reserve, 33 o 40 ’ 52 ’’ S 150 o 44 ’ 39 ’’ E, 18. x. 2015, N. Emery (EME 0489 - 003); 1 ♂, same location, 27. i. 2019, N. Emery (EME 0489 - 004); 1 ♂, same location, 22. xi. 2017, N. Emery & C. Foster (EME 0489 - 005); 1 ♂, same location, 23. x. 2020, D. Emery (EME 0489 - 006); 1 ♂, Lake Parramatta Reserve, Sth James Ruse Drive, 33 o 793 ’ S 150 o 010 ’ E, 31. x. 2020, J. Poyitt (EME 0489 - 007); 1 ♂, Longneck Lagoon, Scheyville NPk, 33 ° 34 ’ 25 ’’ S 150 ° 33 ’ 37 ’’ E, 27. xii. 2018, J. Poyitt (EME 0489 - 008, genitalia prep 489 - 2); 2 ♂, Carysfield Park, Bass Hill, 33 ° 54 ’ 13 ” S 150 ° 59 ’ 53 ” E, 6. xii. 2022, D. Emery (EME 0489 - 019 – 020); 1 ♂, Bourke St., Parramatta, 33 ° 47 ’ 35 ’’ S 151 ° 00 ’ 00 ’’ E, 11. xi. 2022, D. Emery (EME 0489 - 009); 2 ♂, same location, 24. xii. 2022, D. Emery (EME 0489 - 010 – 011); 2 ♂ 2 ♀, same location, 28. x 1.2022, D. Emery (EME 0489 - 012 – 014; 014 & 015 mating); 1 ♂, same location, 4. xi. 2022, D. Emery (EME 0489 - 016); 2 ♂ same location, 6. xii. 2022, D. Emery (EME 489 - 019 – 020); 3 ♂ 2 ♀, same location 11. xii. 2022, D. Emery (EME 489 - 021 – 024; 021 & 022 mating); 1 ♂, Nattai, 33 ° 04 ’ 12 ’’ S 150 ° 26 ’ 35 ’’ E, 20. xii. 2020, D. Emery (EME 0489 - 025); 1 ♀, Burragorang Nature Res., 33 ° 14 ’ 12 ’’ S 150 ° 26 ’ 35 ’’ E, 29. xii. 2010, D. Emery (EME 0489 - 026); 1 ♀, Avon Dam Rd., 34 ° 20 ’ 57 ’’ S 150 ° 37 ’ 30 ’’ E, 23. xii. 1998, D. Emery (EME 0489 - 027); 1 ♂, Mt Annan Botanical Gardens, 34 ° 04 ’ 00 ’’ S 150 ° 46 ’ 01 ’’ E 12. xi. 2024, N. Emery (EME 0489 - 028) (DE); 3 ♂, Silverdale, Nepean Basin, 2. xii. 2003, L. Popple & D. Emery, 489 - 0001 to 489 - 0003 (1 M genitalia prep 489 - 01); 1 ♀, Bourke St., Parramatta, 33 ° 47 ’ 57 ’’ S 151 ° 00 ’ 00 ’’ E, 11. xi. 2022, D. Emery (EME 0489 - 015, mating with - 014); 1 ♂, Bass Hill, xii. 2009, D. Emery (LWP); 1 ♂, Sth. of James Ruse Drive, Lake Parramatta Reserve, NSW, 33 ° 47 ’ 37 ” S 151 ° 0 ’ 36 ” E, 17. ii. 2018, J. Poyitt, (HEMC 489.001); 2 ♂ 1 ♀, same location, 29. xii, 2018, J. Poyitt (HEMC 489.002, 011, 012); 1 ♂, same location, 31. x. 2020, J. Poyitt (HEMC 489.009); 1 ♂, same location, 14. i. 2021, J. Poyitt (HEMC 489.007); 4 ♂, same location, 22 – 30. x. 2022, J. Poyitt (HEMC 489.017 – 020); 1 ♂, same location, 19. xi. 2022, J. Poyitt (HEMC 489.044); 1 ♀, same location, 28. xi. 2022, J. Poyitt (HEMC 489.022); 1 ♂, Lake Parramatta Reserve, NSW, 33 ° 47 ’ 37 ” S 151 ° 0 ’ 36 ” E, 26. xii. 2018, S. Jones (HEMC 489.016); 1 ♂, same location, 24. xi. 2019, J. Poyitt, (HEMC 489.010); 1 ♂, same location, 8. xii. 2019, J. Poyitt, (HEMC 489.009); 2 ♂, same location, 10. i. 2024, J. Poyitt (HEMC 489.044, - 045); 1 ♂, Castlereagh Nature Reserve, 33 o 40 ’ 55 ’’ S 150 o 45 ’ 52 ’’ E, 1. xii. 2018, J. Poyitt, (HEMC 489.006); 1 ♂, Longneck Lagoon, Scheyville NPk, 33 ° 34 ’ 25 ’’ S 150 ° 33 ’ 37 ’’ E, 27. xii. 2018, J. Poyitt (HEMC 489.003); 1 ♂, Norwest Blvd, 33 ° 47 ’ 08 ’’ S 150 ° 57 ’ 41 ’’ E, 10. xii. 2018, J. Poyitt (HEMC 489.014); 1 ♂, Fred Caterson Res., NSW, 33 ° 43 ’ 14 ’’ S 150 ° 59 ’ 18 ’’ E, 15. xi. 2020, J. Poyitt (HEMC 489.013); 1 ♂, Lookout Trail, Cattai NPk., 33 ° 32 ’ 56 ’’ S 150 ° 53 ’ 41 ’’ E, 30. i. 2021, J. Poyitt (HEMC 489.005); 1 ♂, Box Hill fire Stn., 33 ° 38 ’ 19 ’’ S 151 ° 54 ’ 02 ’’ E, 30. x. 2022, J. Poyitt (HEMC 489.021); 1 ♂, Redeemer Baptist School, 33 ° 47 ’ 38 ’’ S 151 ° 00 ’ 48 ’’ E, 30. xi. 2022, J. Poyitt (HEMC 489.045); 3 ♂ 2 ♀, Century Cct., Norwest, NSW, 33 ° 47 ’ 38 ’’ S 151 ° 00 ’ 48 ’’ E, 5 – 10. xii. 2022, J. Poyitt (emerging on Eucalyptus crebra; HEMC 489.025, 027, 029, 030, 032); 4 ♂ 1 ♀, Solent Cct., Norwest, NSW, 33 ° 44 ’ 10 ’’ S 150 ° 57 ’ 38 ’’ E, 10 – 16. xii. 2022, J. Poyitt (emerging on Eucalyptus crebra; HEMC 489.034 – 038); 2 ♂, Ted Horwood Res., Baulkham Hills, 33 ° 45 ’ 38 ’’ S 151 ° 00 ’ 18 ’’ E, 2. i. 2023, J. Poyitt (HEMC 489.039, 040); 1 ♂, Castle Hill Showground, 33 ° 43 ’ 35 ’’ S 150 ° 59 ’ 10 ’’ E, 9. i. 2023, J. Poyitt (HEMC 489.041); 1 ♂, Douglas Park Drive, Douglas Park, 34 ° 11 ’ 46 ’’ S 150 ° 42 ’ 25 ’’ E, 10. i. 2023, J. Poyitt (HEMC 489.042); 1 ♂, Cataract Dam Picnic Area, NSW, 34 ° 15 ’ 41 ’’ S 150 ° 48 ’ 29 ’’ E, 10. i. 2023, J. Poyitt (HEMC 489.043) (JP).	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC4BBA6F17876862FD66250E.taxon	distribution	Distribution, habitat and seasonality. Restricted to central eastern New South Wales, where it is known subcoastally from around Illawarra and Burragorang regions, north to the Hunter River area (Fig. 1). It is most prevalent in Greater Western Sydney. Adults occur in open forest, mainly on rough-barked eucalypts, especially ironbarks (Eucalyptus siderophyloia, E. fibrosa in Western Sydney and E. crebra around Parramatta and the Hills District) and stringybarks (e. g. Eucalyptus obliqua) (Figs. 2 A – B). Females actively fly to ironbarks and secrete themselves vertically along the grooves in the rough bark on the main trunk at around 2 – 5 m, from which they wingflick to attract males (two mating couples were captured in these grooves). In this position they are extremely well camouflaged and difficult to dislodge when attempts are made by human observers to capture them using a longhandled net. Males fly to the locations of the females and continuously move, tick and buzz, leaving if no further encouragement occurs. Males respond to simulated wing-flicks (e. g. finger snapping) and playback of their song recordings, but are quite wary. This species has been encountered from late October to early February and co-occurs with several related and similar looking Yoyetta species, including Yoyetta humphreyae, Yoyetta repetens and Yoyetta cumberlandi.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC4BBA6F17876862FD66250E.taxon	etymology	Etymology. The specific epithetic honours a language of the traditional peoples of the Sydney Basin bioregion where this species is found (a noun in apposition).	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC4BBA6F17876862FD66250E.taxon	description	Description of adult male (Plate 1 A – B, E – G; Plate 2 A – B; Plate 3 A – B; Plate 4 A; Fig. 3). Head slightly narrower than the lateral margins of pronotum; mostly brown to orange-brown with extensive black colouration between and surrounding the ocelli; supra-antennal plates black, occasionally edged dull orange-brown; ocelli pale red; eyes dark brown to black; postclypeus mainly black, dorsal side dark orange-brown, ventral surface with narrow orange lateral margins and medial line; anteclypeus shiny black; rostrum reaching bases of hind coxae, brown basally becoming dark brown or black apically; antennae black. Thorax mostly black. Pronotum with a narrow orange-brown fascia along midline surrounded by black; interior of pronotum brown to dark brown, with irregular black areas, especially along paramedian and lateral fissures; pronotal collar black on anterior margin, orange-brown to dark brown over remainder, black around margins of lateral angles. Mesonotum orange-brown to black; submedian and lateral sigilla black, separated by narrow orange-brown to dark brown areas; lateral margins sometimes orange-brown; cruciform elevation pale brown to pale orange-brown, black centrally, ridges pale brown; scutal depression black; wing grooves mainly dark orange-brown and black. Metanotum dark brown to black. Thorax below mainly orange-brown or dark brown. Legs brown with dark brown to black markings variable between individuals. Coxae about half brown, half dark brown or black; fore femora with inner surface usually black but sometimes dark orange-brown, the outer surface predominantly dull orange-brown, femoral spines dark brown to black; mid and hind femora dark brown to black on anterior sides, paler brown on posterior; fore tibiae dark brown; mid and hind tibiae orange-brown; tarsi orange-brown or brown suffused with black; meracantha black on about basal half, otherwise pale brown. Wings with forewing costal margin a mixture of orange-brown and dark orange-brown; venation generally dark brown to orange-brown; basal membrane light orange-brown. Hind wing venation dark brown; plagas and adjacent area surrounding vein 3 a brown. Opercula dark brown to black basally, plates pale brown; approximately 1.5 x larger than adjacent hind coxae (Plate 2 B). Timbals (Plate 2 A) with five long ribs; ribs 1 – 4 attached to basal spur, separated ventrally and spanning the timbal membrane, rib 5 short, unattached, occupying dorsal half of timbal only; timbal plate with well developed, elongate dome bearing a distinct point of apodeme attachment near centre. Abdomen (Plate 3 A – B) with tergite 1 black; tergite 2 predominantly black, sometimes with diffuse orange areas sublaterally on posterior margin, curved to follow outer margin of exposed timbals; tergites 3 – 7 predominantly pale orange-brown each with a narrow, rectangular, black mark on dorsal midline, this mark being broadest on tergite 3 and narrowing slightly on each subsequent tergite towards the posterior, tergite 3 often with a diffuse dark smudge on lateral areas (noted consistently on Parramatta specimens, Plate 3 B), highly variable in extent between individuals; tergite 8 black, tending dark orange-brown towards posterior margin; epipleurites flexed ventrally, pale orange-brown, diffusely smudged with black. Sternite I orange-brown; sternite II orange with dark-brown to black areas medially and laterally; sternites III – VII pale orange-brown with a small black spot on VII at centre of the posterior segment; sternite VIII orange-brown. Genitalia (Fig. 3) with pygofer upper lobe of moderate size, gradually tapering to a broadly rounded apex (Fig. 3 A); basal pygofer lobe small weakly developed, broadly rounded; median lobe of uncus enlarged, very long (1.5 – 2 mm), concave ventrally, in lateral view narrow basally, expanded around mid-length to widest width, thereafter evenly tapering to a bluntly pointed apex, with ratio of length from base divided by widest width being> 5.0 (Fig. 3 B); claspers robust, claw-like, partially meeting, in ventral view their inner margins diverging with an indentation midway along the length and their outer margins gently turned outwards; aedeagus recurved distally through 180 degrees, the vesica weakly toothed either side subapically, apex terminating in a short, slender ‘ beak’ downturned> 45 o when viewed laterally, with a pair of very small flat ear-like lobes pointing inwards, minutely toothed posteriorly; pseudoparameres very long, nearly reaching most distal part of theca, very slender, hair-like and weakly knobbed apically (Figs. 3 C – D). Description of adult female (Plate 1 C – D, Plate 4 A). Head, thorax, wings and legs similar to male, except with greater amounts of ochraceous-brown fascia around ridges on mesonotum. Abdomen with more pronounced annular markings on tergites (Plate 4 A). Tergite 1 orange, black medially and diffusely black along anterior margin of lateral third; tergites 2 – 7 predominantly dark orange-brown each with a distinct black marking on the dorsal midline, broadest on tergite 3, extending laterally over the anterior half of each tergite and blending into dark orange-brown along the posterior half with a thin lighter orange-brown posterior margin; tergite 8 black over anterior half, tending dark orange-brown over remainder, with a black spot on lateral aspect. Sternite I black; sternites II – VII pale orange-brown with a black triangular marking along the midline, base widest anteriorly, apex posteriorly; sternite VIII darker orange-brown; abdominal segment 9 pale brown over dorsal two-thirds, darker brown ventrally, dorsal beak black, a distinct black longitudinal marking either side of dorsal midline, extending laterally along the anterior margin, enclosing a pale brown longitudinal triangle along the midline; a small black spot laterally near the posterior apex; ovipositor sheath dark brown, black posteriorly, ovipositor dark brown, becoming black posteriorly, extending 1 – 1.5 mm beyond abdominal segment 9 (Plate 4 A). Measurements (in mm; range with mean in parentheses for 12 males and 12 females, including smallest and largest specimens). Length of body including head: male, 19.7 – 22.1 (21.5); female, 23.5 – 25.8 (24.6). Length of forewing: male, 21.6 – 26.1 (24.5); female 28.5 – 32.4 (30.0). Width of forewing: male, 7.7 – 8.9 (8.0); female, 8.5 – 10.2 (9.4). Width of head (including eyes): male, 5.3 – 6.2 (5.8); female, 6.1 – 7.1 (6.6). Width of pronotum (across lateral angles): male, 5.4 – 6.2 (5.8); female, 5.8 – 7.6 (6.7). Width of abdomen: male, 5.4 – 6.4 (5.8); female, 6.2 – 6.7 (6.4). Length of ovipositor: female, 7.5 – 9.3 (8.3). Morphological variation within and between populations. Both male and female specimens exhibit a colour range from medium brown to dark brown to black over the head and pronotum. The dorsal abdominal markings of all females captured to date range from black to grey.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC42BA7617876CAEFD6727CE.taxon	description	(Plates 2 C – D, 3 C – D, 4 B, 5; Figs. 1, 2 C – D, 6 – 9) Types. Holotype 1 ♂, Windamere Dam, Mudgee, NSW, 32 o 43 ’ 32 ’’ S 149 o 46 ’ 19 ’’ E, 8. i. 2022, N. & D. Emery & C. Foster (EME 0490 - 017) (K 624931, AM). Paratypes. NEW SOUTH WALES. 1 ♂ 1 ♀, same data as holotype, (EME 0490 - 030, - 031) (K. 924932, K. 924936, AM); 2 ♂, same location as holotype, 2. i. 2022, S. & T. Corbin (EME 0490 - 001 – 2; genitalia prep 490 - 1), 11 ♂ 1 ♀, same data as holotype (EME 0490 - 003 – 014); 16 ♂ 2 ♀, same location as holotype, 8. i. 2022, N. & D. Emery & C. Foster (EME 0490 - 017 – 029; 032 – 035); 2 ♂, Goulburn R., nr Wollar, 32 o 19 ’ 43 ’’ S 149 o 57 ’ 37 ’’ E, 3. i. 2022, T. Corbin (EME 0490 - 036 – 7); 4 ♂, Cassilis rest area, NSW, 32 o 02 ’ 15 ’’ S 149 o 56 ’ 01 ’’ E, 6. i. 2013, N. C. & D. Emery (EME 0490 - 038 – 041; genitalia preps 490 - 3, 490 - 4); 3 ♂, same location, 15. xii, 2020, C. & D. Emery (EME 0490 - 042 – 044); 5 ♂, same location, 23. i. 2023, D. Emery (EME 0490 - 058 – 062); 1 ♂, Clandulla State Forest, 32 o 53 ’ 19 ’’ S 149 o 56 ’ 37 ’’ E, 7. i. 2010, D. Emery & L. Popple (EME 0490 - 045); 3 ♂ 2 ♀, Cooper Drive, Clandulla, 32 o 54 ’ 27 ’’ S 149 o 49 ’ 00 ’’ E, 26. xi. 2020, D. Emery (EME 0490 - 046 – 050; genitalia prep 490 - 5); 4 ♂ 1 ♀, same location, 15. i. 2021, N. & D. Emery (EME 0490 - 051 – 055); 1 ♂ 1 ♀, Bocoble, 32 o 47 ’ 31 ’’ S 149 o 42 ’ 43 ’’ E, 1050 m, 11. xii. 2021, H. Ottersdorf (EME 0490 - 056 – 057); 1 ♂, Denman, 5. xi. 2007, R. Chin (EME 0490 - 058); 4 ♂, Whitegum Lookout, Warrumbungle NPk, 31 o 17 ’ 33 ’’ S 149 o 02 ’ 20 ’’ E, 16. xi. 2013, N. & D. Emery (EME 0717 - 059 – 062); 3 ♂, 14. xi. 2013, N. & D. Emery (EME 0490 - 063 – 065; genitalia prep 490 - 2); 1 ♂, same location, 12. i. 2014, N., C. & D. Emery (EME 0490 - 066); 1 ♂ 1 ♀, same location, 12. i. 2014, N., C. & D. Emery, (EME 0490 - 067 – 068; genitalia prep 490 - 6); 1 ♂, same location, 8. xii. 2015, C. & D. Emery (EME 0490 - 069); 2 ♂, same location, 5. xi. 13, C. & D. Emery (EME 0490 - 070 – 071); 1 ♀, same location, 8. xii. 2015, N. & D. Emery (EME 0490 - 072) (DE); 1 ♂, same location as previous, 14. xi. 2013, N. &. D. Emery; 1 ♂, same data as previous, 30. xi. 2013; 1 ♂, Cassilis rest area, NSW, 32 o 02 ’ 15 ’’ S 149 o 56 ’ 01 ’’ E, 6. i. 2013, N. C. & D. Emery; 1 ♂, same location as previous, 15. xi. 2020, C. & D. Emery; 1 ♂, Napolean Reef area via Bathurst, 33 ° 26 ’ 20 ’’ S 149 ° 45 ’ 23 ’’ E, 8. i. 2010, Popple & Emery, open woodland; 2 ♂, same location as holotype, 2. i. 2022, S. & T. Corbin (LWP); 2 ♂, same data as holotype (EME 0490 - 015 – 6); 1 ♂, Glen Davis Rd. between Airly Creek Rd. and Pinegrove Rd., Capertee, NSW, 33 ° 08 ’ 04 ’’ S 150 ° 04 ’ 04 ” E, 13. xii. 2022, J. Poyitt, (HEMC 490.001); 4 ♂, same location as holotype, 11. i. 2023, J. Poyitt (HEMC 490.004 – 007); 4 ♂, same location as holotype, 17. i. 2023, J. Poyitt, (HEMC 490.008 – 011) (JP). Other material. QUEENSLAND. 1 ♂, Carnarvon Stn, Homestd, 24.809 ° S 147.753 ° E, 747 m, 13. x. 2014, J. Buchan, mv lamp, 37156, T 199410; 1 ♀, Carnarvon Stn, nr homestead, 24.809 ° S 147.753 ° E, 747 m, 13. x. 2014, B. Hawkins, hand collected, flowering flat, T 199411; 1 ♂, Mt Moffatt NP, top shelter shed, 1000 m, 10 – 12. xii. 1987, Monteith, Thompson, Yeates (QM).	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC42BA7617876CAEFD6727CE.taxon	distribution	Distribution, habitat and seasonality. The majority of specimens of Yoyetta psammitica sp. nov. have been found along the western slopes of the Great Dividing Range in New South Wales from around Capertee and Wollemi National Park, north to around Merriwa and Cassilis and further northwest into the Warrumbungle Range National Park (NPk; Fig. 1). An isolated population also occurs in the Carnarvon Range in central Queensland. In New South Wales, this species appears to favour smooth barked eucalypts, particularly Eucalyptus cannonii (Capertee Stringybark) around Mudgee (Windamere Dam and northern Cudgegong River) and Clandulla (Fig. 2 C – D). At Cassilis Park rest area on the Golden Highway, specimens are also attracted to narrow-leaved ironbark trees (E. crebra), while in the Warrumbungle NPk, they are often found on white gums (E. rossii). Individuals call in flight and while moving constantly around the tree trunks. Single populations are mostly present at Cassilis rest area and Windamere Dam, but at Clandulla and Bocoble, Y. psammitica sp. nov. co-occurs with Y. humphreyae, Yoyetta timothyi Emery, Emery & Popple and two additional undescribed Yoyetta species. In the Warrumbungle NPk, Y. psammitica sp. nov. cohabits with Y. humphreyae and Y. australicta Popple & Emery. Most specimens of Y. psammitica sp. nov. have been collected from mid-November to late-January. It has been noted that since 2015, two years after wildfires burnt out over 80 % of Warrumbungle NPk, specimens of this species have been difficult to find, with those of Y. australicta and Y. humphreyae being much more common. The reason (s) for this change in prevalence is not clear.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC42BA7617876CAEFD6727CE.taxon	etymology	Etymology. The term “ psammitica ” (adjective, “ sandstone ”) reflects the distribution of this species throughout the open sandstone woodlands of the western slopes of the Great Dividing Range of NSW and southern Queensland.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC42BA7617876CAEFD6727CE.taxon	description	Description of adult male (Plates 2 C – D, 3 C – D, 5 A – D, G; Figs. 2 C – D, 6) Head slightly narrower than the lateral margins of pronotum; mostly light to dark orange-brown with extensive black colouration between and surrounding the ocelli, sometimes meeting medial margins of eyes, a triangular orange-brown fascia with a base on the posterior border narrows to an apex midway between the ocelli; supra-antennal plates black, edged dull orange-brown around base of antennae; ocelli dull red; eyes dull red to light brown; postclypeus orange-brown dorsally with a black horizontal stripe along the anterior margin, ventral surface black with an orange medial line and orange coloration over half of lateral margins; anteclypeus shiny black; lorum black, gena black; rostrum reaching anterior margins of hind coxae, dark brown proximally, becoming dark brown to black distally; antennae black. Thorax mostly dark brown to black. Pronotum with a narrow orange-brown fascia along midline surrounded by black, resembling a vintage key lock; interior of pronotum orange to dark brown, with irregular black areas, especially along paramedian and lateral fissures and adjacent ridges, also bordering the lateral halves of the pronotal collar; pronotal collar black on anterior margin, orange-brown to dark brown over remainder. Mesonotum variably black with orange-brown intrusions; submedian and lateral sigilla black, intervening depressions orange-brown to dark brown; lateral margins sometimes orange-brown; scutal depressions black; cruciform elevation orange-brown, variably black along midline and into anterior depression, lateral depressions striated black transversely; wing grooves mainly dark orange-brown and black. Metanotum dark brown to black; black central “ dot ” with orange-brown ridges. Thorax below mainly orange-brown or dark brown. Legs brown with variable longitudinal orange-brown to black markings between specimens. Coxae about half orange-brown and black; fore femora with inner surface black but sometimes striped dark orange-brown, the outer surface predominantly dull orange-brown, femoral spines dark brown basally and black tipped, upright; mid and hind femora dark brown to black anteriorly, paler orange-brown posteriorly; fore tibiae dark brown; mid and hind tibiae orange-brown; tarsi orange-brown to brown; claws black; meracantha variably black on base around medial half, remainder pale cream-brown, reaching anterior margin of opercula. Wings with forewing costal margin orange-brown to dark orange-brown; venation generally dark brown to orange-brown; basal cell translucent, tinged orange; basal membranes orange; smoky infusions throughout a 1 – a 8; infuscations of variable intensity along crossveins r and r-m on the majority of individuals. Hind wing venation dark brown; v-shaped plagas tinged orange-grey visible along veins 2 a and 3 a and within anal cells. Opercula (Plate 2 D) variably black basally, plates pale yellow-brown, spatulate, curved medially, flexed ventrally, not meeting; approximately 1.5 x larger than adjacent hind coxae. Timbals (Plate 2 C) with five long ribs; ribs 1 - 4 attached to basal spur, separated ventrally and spanning the timbal membrane, rib 5 unattached, extending over central half of membrane; timbal plate with well developed, elongate dome bearing a distinct point of apodeme attachment near centre. Abdomen (Plate 3 C – D) with tergite 1 orange centrally, with symmetrical, black diagonal striations posteriorly, black laterally; tergite 2 variably black along anterior third and laterally, becoming orange posteriorly, curved around outer margin of exposed timbals; tergites 3 – 7 predominantly pale orange-brown each with a distinct black mark on dorsal midline, this mark broadest on tergite 3 and equivalently narrower on tergites 4 – 7, tergites 3 – 5 sometimes also with a diffuse dark smudge on lateral sides; tergite 8 black over anterior half, tending dark orange-brown over remainder, with a black spot on lateral aspect; epipleurites orange, flexed ventrally. Sternite I orange-brown to black; sternite II orange with dark-brown to black areas medially and laterally; sternites III – VII pale orange-brown; sternite VIII darker orange-brown. Genitalia (Fig. 6) with pygofer black, upper lobe of moderate size with a broadly rounded apex, in ventral view angled medially (Fig. 6 A); basal pygofer lobe small weakly developed, broadly rounded; median lobe of uncus enlarged, variably elongated with length around 1.0 – 1.4 mm, concave ventrally, in lateral view narrow basally, expanded around mid-length to its widest width, thereafter evenly tapering to a rounded apex with ratio of length from base divided by widest width ranging from 2.6 – 4.5; claspers robust, claw-like in lateral view (Fig. 6 B), partially meeting, in ventral view their inner margins diverging with a “ notch ” around aedeagus and their outer margins gently turned outwards to a bluntly pointed apex; aedeagus recurved distally through 180 degrees with ornamentation distally from the recurvature, the vesica broadened and weakly toothed either side towards apically, apex terminating in a short, slender ‘ beak’ pointing outwards, downturned> 45 o viewed laterally, with a pair of very small flat ear-like flanges directed medially and minutely toothed; a “ viking helmet ” shape viewed laterally; pseudoparameres very long, reaching the distal recurvature of the aedeagus, very slender, hair-like, each with a rounded apical knob (Figs. 6 C – D). Description of adult female (Plate 4 B, 5 E – F, H). Head, thorax, wings and legs similar to male. Abdomen. Tergite 1 orange, black laterally along posterior margin; tergite 2 orange suffused variably with black coloration, most prominent along posterior margin, a black spot on each lateral side; tergites 3 – 7 predominantly dark orange-brown each with a distinct black “ double triangle ” on dorsal midline, broadest on each anterior margin, narrowing centrally to an apex and joined to the apex of a posterior triangle with its base on the posterior margin of each tergite, most pronounced on tergites 3 – 5, reducing on successive tergite 4 – 7; a distinct black marking centrally located on each ventrolateral aspect; tergite 8 black over anterior half, tending dark orange-brown over remainder, with a black spot on lateral aspect. Sternite I black; sternites II – VII pale orange-brown with black infusion along midline; sternite VIII darker orange-brown; abdominal segment 9 dark ochraceous over dorsal two-thirds, orange-brown ventrally, dorsal beak black, a distinct black longitudinal marking either side of dorsal midline extending laterally along the anterior margin; a small black spot laterally near the posterior apex; ovipositor sheath dark brown, black posteriorly, ovipositor dark brown, becoming black posteriorly, extending <1 mm beyond abdominal segment 9. Measurements (in mm; range with mean in parentheses for 26 males and 10 females, including smallest and largest specimens). Length of body including head: male, 18.9 – 24.2 (22.5); female, 20.2 – 25.1 (23.7). Length of forewing: male, 25.0 – 29.6 (27.8); female, 26.1 – 32.4 (29.9). Width of forewing: male, 7.8 – 9.6 (9.1); female, 8.3 – 9.9 (9.5). Width of head (including eyes): male, 5.3 – 6.5 (6.1); female, 5.9 – 7.5 (6.7). Width of pronotum (across lateral angles): male, 5.7 – 6.5 (6.2); female, 6.2 – 7.6 (6.8). Width of abdomen: male, 5.6 – 6.6 (6.3); female, 5.7 – 6.8 (6.4). Length of ovipositor: female, 6.3 – 8.6 (7.5). Morphological variation within and between populations. The morphological appearances of Y. psammitica sp. nov. specimens are consistent within and between the populations around Clandulla, Cassilis and the Warrumbungle NPk. However, males from the northern sites (Warrumbungle NPk and Carnarvon Range, Queensland) have a smaller body length (BL; 18.9 – 22.4 mm; n = 11) than those from Clandulla and Cassilis (BL> 22.4 mm; n = 15). Females from the northern sites are also substantially smaller (BL 20.2 – 22.9 mm; n = 3) than those from the two southernmost populations (BL> 24.1 mm; n = 7). Morphological distinguishing features. Male specimens of Y. psammitica sp. nov. most closely resemble Y. corindi, Y. humphreyae, Y. ignita, Y. robertsonae and Y. robusta, Y. darug sp. nov. and Y. fumea sp. nov., which are all similar in size and have brown hind wing plagas. They can be distinguished by the following combination of characters: (1) median lobe of uncus well developed but not particularly elongate (extending ventrally to the same extent as claspers; Fig 6 B), (2) fore wings with a weak to moderate smoky infusion over distal apical cells, and (3) fore wing length> 26 mm. Notably, in the Warrumbungles, male specimens of Y. psammitica sp. nov. can only be confidently differentiated from cohabiting Y. humphreyae males by their shorter median lobe of the uncus. Females can be distinguished from all other Yoyetta species apart from Y. fumea sp. nov. by having (1) fore wings with a weak to moderate smoky infusion over distal apical cells, (2) ventral side of abdominal segment 9 orange-brown, (3) ovipositor extending <1 mm beyond abdominal segment 9 and (4) more pronounced black markings along the midline of the sternites. Females of Y. psammitica sp. nov. can be distinguished from Y. fumea sp. nov. by their slightly longer ovipositor, which extends <1 mm beyond the apex of abdominal segment 9 (cf. not extending noticeably beyond the anal styles in Y. fumea sp. nov.). Calling song (Figs. 7 – 9). The male calling song of Y. psammitica sp. nov. has three modes. The first mode (“ clicking ” mode) comprises a series of repetitive clicking, which is produced whilst in flight and occasionally when stationary on the trunk or branches of Eucalyptus species. The second mode is the same as the first mode but includes a short soft buzz after each click (“ click and buzz ” mode) and only occurs while stationary. The third mode is a variant of the second mode that commences with a buzz followed by a series of clicks (“ buzz and click ” mode; Fig. 9). Closer examination of the clicking mode (all statistics, n = 9 recordings from the Warrumbungles, Mudgee and Clandulla localities) shows each click is a syllable as illustrated in Figs 7 – 9. Both in flight and at rest, each short syllable (0.012 – 0.023 s, 8 – 9 pulses, sometimes with 1 - 2 preceding soft pulses) is separated by a gap of roughly half a second in duration. The click and buzz mode is illustrated in Fig. 8, and is described as follows (all statistics, n = 5 recordings from the type locality). Following a syllable that is equivalent to that in the clicking mode is a short gap (0.04 – 0.16 s), then a short soft buzz (0.07 – 0.18 s), followed by a longer gap (0.32 – 0.50 s). The third “ buzz and click ” mode is illustrated in Fig 9 (n = 1) has only been recorded in the Warrumbungles, though it has also been observed at Napoleon Reef near Bathurst. The song begins with a long buzz (8.0 s) followed by a series of short syllables. The frequency plateau is consistent between modes. It ranges broadly from approximately 6.5 to 12.5 kHz, with a dominant frequency between 7 and 11.5 kHz. Y. humphreyae co-occurs with Y. psammitica sp. nov., but Y. humphreyae produces fewer (5 – 6) pulses (cf. 8 – 9 pulses in Y. psammitica sp. nov.). The song is also close to Y. corindi (which lacks the “ long buzz ” component) and the clicking structure may be indistinguishable from Y. darug sp. nov., although these species do not overlap in geographical distribution with Y. psammitica sp. nov.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC5DBA7B1787682BFF522562.taxon	description	(Plates 2 E – F, 3 E, 4 C, 6; Figs. 1, 2 E – F, 10 – 12) Types. Holotype 1 ♂, Cullala Rd, Lower Broughton, 35 o 05 ’ 46 ’’ S 149 o 46 ’ 28 ’’ E, 10. i. 2021, N., C. & D. Emery (EME 0717 - 037) (K. 624937, AM). Paratypes. NEW SOUTH WALES: 1 ♂, Jacqua Rd., Windellama, 34 o 57 ’ 51 ’’ S 149 o 54 ’ 29 ’’ E, 10. i. 2021, N., C. & D. Emery (EME 0717 - 027); 1 ♀, Jerrara Rd., Bungonia, 34 o 47 ’ 13 ’’ S 149 o 57 ’ 40 ’’ E, 9. i. 2023, C. & D. Emery (EME 0717 - 039) (K. 624938 – 939, AM); 2 ♂, Yarrawal Rd., Windellama, NSW, 34 o 58 ’ 24 ’’ S 149 o 55 ’ 01 ’’ E, 4. i. 2019, S. & T. Corbin, D. Emery (EME 0717 - 025 – 026; genitalia prep 717 - 2); 1 ♂ 4 ♀, Jacqua Rd., Windellama, 34 o 57 ’ 51 ’’ S 149 o 54 ’ 29 ’’ E, 10. i. 2021, N., C. & D. Emery (EME 0717 - 028 – 033); 3 ♂ 1 ♀, same data as holotype (EME 0717 - 034 – 036; genitalia preps 717 - 3, 717 - 4); 1 ♀, Jerrara Rd., Bungonia, 34 o 47 ’ 13 ’’ S 149 o 57 ’ 40 ’’ E, 9. i. 2023, C. & D. Emery (EME 0717 - 042) (DE); 1 ♂, same location as holotype, 30. xi. 2013, N. & D. Emery; 1 ♂, Yarrawal Rd., Windellama, NSW, 34 o 58 ’ 24 ’’ S 149 o 55 ’ 01 ’’ E, 4. i. 2019, S. & T. Corbin, D. Emery; 1 ♂, Jacqua Rd, Windellama, 34 o 57 ’ 51 ’’ S 149 o 54 ’ 29 ’’ E, 10. i. 2021, N., C. & D. Emery (LWP); 2 ♂, Jacqua Rd., Windellama, 34 o 57 ’ 51 ’’ S 149 o 54 ’ 29 ’’ E, 10. i. 2021, N., C. & D. Emery (EME 0717 - 040 – 041); 2 ♂, Jerrara Rd. 600 m north of Tickner Valley Rd., Marulan, NSW, 34 ° 47 ’ 07 ” S 149 ° 57 ’ 38 ” E, 28. xii. 2022, J. Poyitt, (HEMC 717.001,002) (JP).	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC5DBA7B1787682BFF522562.taxon	distribution	Distribution, habitat and seasonality. Specimens of Yoyetta fumea sp. nov. are currently known only from around Windellama and Tarago in the Southern Highlands (Fig. 1). Individuals mainly favour the trunks of black she-oaks (Allocasuarina littoralis), where they call while stationary or moving around the trunk at heights of 2 – 5 m from the ground. Around Windellama, the black and drooping she-oaks (Allocasuarina littoralis and A. verticillata, respectively), are the main floral preferences (Figs 2 E – F). This is a notable difference between this species and the closely related Y. humphreyae, which is primarily associated with eucalypts. Adults have been found between November and January.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC5DBA7B1787682BFF522562.taxon	etymology	Etymology. The name “ fumea ” (Latin, adjective, feminine) reflects the prominent smoky infusion of the apical cells of the forewing, which is more pronounced in this species than in closely related Yoyetta species.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC5DBA7B1787682BFF522562.taxon	description	Description of adult male (Plates 2 E – F, 3 E, 4 C, 6; Fig. 10). Head slightly narrower than the lateral margins of pronotum; extensively black with orange brown fascia along posterior half of midline, between ocelli and eyes and surrounding pedicles; supra-antennal plates black in all specimens, occasionally edged dull orange-brown; ocelli pale pink to yellow; eyes dark brown; postclypeus dark orange-brown dorsally, sometimes with black infusions along the midline, anterior margin and ventral areas black, with lateral margins sometimes dark orange-brown, transverse ridges black, lorum black, gena black; anteclypeus mainly black, sometimes smudged orange-black anteriorly; rostrum reaching bases of hind coxae, brown basally becoming dark brown or black apically; antennae black. Thorax black with irregular orange-brown marking areas, surrounding paramedian and lateral fissures; pronotal collar black, sometimes with a central orange-brown “ V ” with apex anterior and extending into pronotum anteriorly. Mesonotum mainly black; submedian and lateral sigilla black; scutal depressions black; cruciform elevation black with dark orange infusion with faint black transverse striations in the lateral depressions, the symmetrical orange-brown markings extending forward along anterior ridges and ending lateral to the scutal depressions; wing grooves mainly dark orange-brown and black. Metanotum black with orange-brown central section. Thorax below mainly black with variable orange markings. Legs dark brown to black markings variable between individuals. Coxae mainly black, coxal membranes bright orange-red, trochanter black; fore femora with inner surface usually black, the outer surface predominantly dull orange-brown, femoral spines black, erect; mid and hind femora striped black on anterior sides and paler brown on posterior; fore tibiae black anteriorly, dark brown posteriorly; mid and hind tibiae dark orange-brown; tarsi orange-brown or brown suffused with black; claws dark orange-brown, becoming black at tips; meracantha black on basal half, cream over remainder, not reaching opercula. Wings with forewing costal margin black to dark orange-brown; venation generally dark brown to orange-brown; basal cell translucent, tinged green-orange; basal membranes orange; smoky infusions throughout a 1 – a 8; infuscations of variable intensity along crossveins r and r-m. Hind wing venation dark brown; v-shaped plagas tinged smoky-grey visible along veins 2 a and 3 a and within anal cells. Opercula (Plate 2 F), pale black basally, plates dull cream-brown, spatulate, curved medially, flexed ventrally, not meeting; approximately 1.5 x larger than adjacent hind coxae. Timbals (Plate 2 E) with five long ribs; ribs 1 – 4 attached to basal spur, separated ventrally and spanning the timbal membrane, rib 5 short, unattached, occupying dorsal central half of timbal only; timbal plate with well developed, prominent elongate and elevated dome bearing a distinct point of apodeme attachment near centre. Abdomen with tergite 1 black, posterior margins dark orange anterior to timbal cavity; tergite 2 predominantly black with dark orange spot either side of posterior midline, sometimes meeting along posterior margin; tergites 3 – 7 predominantly pale orange-brown each with a narrow black mark on dorsal midline, this mark being broadest on tergite 3 and narrowing slightly on each subsequent tergite posteriorly, also with a diffuse dark smudge on lateral sides resolving as a black dot on the postero-ventral aspect; tergite 8 black, tending dark orange-brown towards posterior margin, with a distinct dark orange marking on the antero-lateral aspect (Plate 3 E); epipleurites flexed ventrally, a small black dot on the posterior margins. Sternite I black, orange-brown posteriorly; sternite II orange with dark-brown to black areas medially and laterally; sternites III – VII pale orange-brown, diffuse, with light black infusions on lateral halves; sternite VIII mainly orange-brown, dark brown at base, paler brown at apex. Genitalia (Fig. 10) with pygofer upper lobe of moderate size, gradually tapering to a peaked but rounded apex, in ventral view projecting outwards; basal pygofer lobe small and weakly developed, broadly rounded (Fig 10 A); median lobe of uncus enlarged, long, in lateral view (Fig 10 B) almost uniformly wide along length (1.6 – 2 mm), evenly tapering to a rounded apex, with ratio of length from base divided by widest width around 3.0, extending distally beyond the claspers, although shorter than the distance between the dorsal beak and upper pygofer lobe; claspers robust, claw-like, partially meeting, in ventral view their bases closely applied to aedeagus, with inner margins curving laterally and their outer margins turned outwards to a blunted pointed apex; aedeagus recurved distally through 180 degrees, the vesica consistently toothed either side from the curvature, apex terminating in a short, slender ‘ beak’ pointing outwards and a pair of very small flat ear-like flanges directed medially and minutely toothed; a “ viking helmet ” shape viewed laterally, with apex either straight or slightly downturned (<30 o dorsally) in lateral view; pseudoparameres very long, reaching the distal recurvature of the aedeagus, very slender, each with a rounded apical knob (Fig 10 C – D). Description of adult female (Plates 4 C, 6 C – D, F). Head mainly similar to male, larger amounts of orange-brown areas between ocelli and eyes, a black spot on the posterio-medial margin of the eye; ocelli colour ranging from bright red to dull pink; postclypeus dark orange-brown dorsally, sometimes with black infusions along the midline, anterior margin black, ventral aspect black in most specimens, lateral margins dark orange-brown, lighter specimens with orange-brown midline, transverse ridges black, lorum black, gena black. Thorax and legs similar to male, with coloration ranging from mainly black to principally dark orange-brown. Wings similar to male, smoky infusions and cross-vein infuscations slightly lighter. Basal membranes either bright orange in orange-brown specimens or muddy orange in darker individuals. Abdomen. (Plates 4 C, 7 C – D). Tergite 1 orange, black laterally along posterior margin, a central black spot on darker specimens; tergite 2 orange black over anterior third, orange to dark orange-brown over remainder, black markings around lateral extremity; tergite 3 with black coloration either restricted to the anterior margin or diffusely broadened laterally, a narrow black mark on dorsal midline, wider on darker females; tergites 4 – 7 predominantly dark orange-brown each with a distinct black “ parallelogram ” on dorsal midline, broadest on each anterior margin and widest on darker specimens, most pronounced on tergites 3 – 5, reducing slightly in width on successive tergites 4 – 7, variable black smudging laterally on each tergite, culminating in a darker black marking on the posterior half of the lateral margins; tergite 8 black over anterior half, more extensively black in darker individuals, tending dark orange-brown over remainder, with a black spot on lateral aspect. Sternite I black; sternites II – VII pale orange-brown with very faint black smudging along midline; sternite VIII darker orange-brown, a black spot either side of posterior midline; abdominal segment 9 dark ochraceous over dorsal two-thirds, darker brown ventrally, dorsal beak black, in darker specimens a black line extends from the dorsal beak anteriorly for a short distance along midline, a distinct black longitudinal marking either side of dorsal midline extends laterally along the anterior margin, a small black spot laterally near the posterior apex; ovipositor sheath dark brown, black posteriorly; ovipositor dark brown, becoming black posteriorly, not extending noticeably beyond the anal styles. Measurements (in mm; range with mean in parentheses for 12 males and 5 females, including smallest and largest specimens). Length of body including head: male, 21.3 – 22.4 (21.9); female, 20.7 – 23.3 (21.4). Length of forewing: male, 22.8 – 28.6 (27.0); female, 27.7 – 29.6 (28.3). Width of forewing: male, 8.5 – 9.8 (9.0); female, 8.9 – 9.7 (9.2). Width of head (including eyes): male, 5.5 – 6.2 (5.9); female, 6.3 – 6.8 (6.5). Width of pronotum (across lateral angles): male, 5.3 – 6.0 (5.7); female, 6.0 – 6.5 (6.3). Width of abdomen: male, 5.4 – 6.2 (5.9); female, 5.5 – 6.1 (5.9). Length of ovipositor: female, 5.5 – 7.5 (6.1). Morphological variation within and between populations. Specimens of this species are consistent morphologically, likely reflecting their current limited distribution. Morphological distinguishing features. The presence of infuscations on fore wing crossveins r and r-m is a trait that consistently distinguishes Y. fumea sp. nov. from all other described species of Yoyetta, apart from Y. psammitica sp. nov. and Y. humphreyae. Males of Y. fumea sp. nov. can be distinguished from Y. psammitica sp. nov. by having a more elongate median uncal lobe that extends posteriorly beyond the claspers (a feature that is visible without dissection). Notably, the median lobe of the uncus in Y. fumea sp. nov. is shorter than the distance between the dorsal beak and the apex of upper pygofer lobe, a feature that distinguishes it not only from Y. humphreyae, but also from other species that possess a more elongate uncus, including Y. darug sp. nov. and Y. corindi (the uncus in these three species is about as long as the distance between the dorsal beak and apex of the pygofer lobe). Females of Y. fumea sp. nov. can be distinguished from Y. psammitica sp. nov. by (1) their shorter ovipositor, which does not extend conspicuously beyond the anal styles (cf. extending <0.5 mm beyond the anal styles in Y. psammitica sp. nov.), and (2) their lack of distinct black markings along the midline of the tergites. Calling song. (Figs. 11 – 12). The male calling song of Y. fumea sp. nov. has two modes. One mode (combination mode) has a combination of buzzes and clicks. The second mode (clicking mode) comprises only clicks. Both modes can be produced when the male is stationary; however, only the clicking mode is produced in flight. Both modes can be produced for an extended duration> 60 seconds. The combination mode contains phrases that have two variants, which can be produced in an alternating fashion by the same male as Fig. 11 illustrates. This mode and its variants are described as follows (all statistics, n = 7 recordings from Windellama, Clandulla and the type locality). Each phrase begins with an echeme typically between 0.315 and 0.829 s (but can be maintained for at least 50 s duration) that increases in amplitude as it progresses, followed by a short gap (0.05 s). After this is either a repeated series of clicking and short buzzing (variant 1) or clicking (variant 2; similar to the clicking mode as shown in Fig. 12). Further analysis of variant 1 reveals a alternating series of short syllables (0.010 – 0.014 s) and short macrosyllables (0.097 – 0.147 s). This may be repeated up to 4 times before the phrase finishes with an echeme (0.154 – 0.242 s). In one recording from Windellama, the phrase was extended to include an additional short gap (0.123 s) followed by an echeme (0.395 s) that increased in amplitude, then a longer gap (0.345 s), and finishing with a short echeme (0.142 s) (Figure 11 C). Further recordings would be required to determine how often these extended phrases are produced. For variant 2, each click is a syllable that is equivalent in structure and duration to those in variant 1; but each syllable is separated by a gap (0.205 – 0.292 s). In available recordings (n = 4), males produced between 6 and 23 syllables (in succession) in variant 2. The clicking mode consists of repeated single syllables as illustrated in Fig 12 and are equivalent in duration to the syllables in the combination mode. A gap of varying duration (0.135 – 0.275 s) punctuates each syllable (all statistics, n = 3 from the Windellama locality). When producing this mode, males fly between trees at canopy height making it difficult for acoustic recording, and especially difficult to isolate the song from other co-occurring cicada calling songs. It is hypothesized that females respond with wing-flicks following each syllable in this mode as has been recorded for other Yoyetta species. The frequency plateau is generally consistent between modes. It ranges broadly from approximately 5.5 to 12.5 kHz, with a dominant frequency between 7 and 11 kHz. The calling song and singing behaviour of Y. fumea sp. nov. is closest to Y. humphreyae in terms of complexity and structure. Y. fumea sp. nov. produces syllables that often contain more pulses (6 – 7 pulses, cf. 5 – 6 pulses in Y. humphreyae). It has also been noted in the field that the singing behaviour of Y. humphreyae and Y. fumea sp. nov. differs, with the latter producing more complex alternating click and buzz variants, as described above. In comparison, although complex, the call of Y. humphreyae has a more predictable sequence as illustrated by Moulds and Popple (2021). The song is also similar to those of Y. corindi (illustrated in Popple and Emery, 2022) and Y. psammitica sp. nov. The long buzz produced by Y. fumea sp. nov. has not been recorded in the song of Y. corindi and the two species are not known to co-occur. Y. psammitica sp. nov. produces clicks with 8 – 9 pulses (cf. 6 – 7 pulses in Y. fumea sp. nov.).	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
03D60217CC55BA7817876917FB9C2781.taxon	description	Specimens must be set with wings spread to begin using this key. A vernier calliper is required to check measurements. Male specimens must often be examined microscopically and may require dissection in some instances.	en	Emery, David L., Emery, Nathan J., Popple, Lindsay W. (2025): Description of three new cicada species in the genus Yoyetta Moulds (Hemiptera: Cicadidae: Cicadettinae), from eastern Australia. Zootaxa 5590 (2): 151-184, DOI: 10.11646/zootaxa.5590.2.1, URL: https://doi.org/10.11646/zootaxa.5590.2.1
