taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03D58787FFA3FFDBC2C1C3D3956F4D05.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832699/files/figure.png	https://doi.org/10.5281/zenodo.14832699	Figure 4. Phylogenetic relationships of Phyllonastes + Urkuphrynegen. nov.(= Noblella northern clade). Maximum likelihood tree obtained for genes 16S, 12S, TYR and RAG1 from a matrix of 3846 bp for 170 terminals.Support values are on the corresponding branches:aLRT values before the slash and non-parametric bootstrap after the slash; triangles on the branches represent aLRT or bootstrap support values of 100%. For each specimen the voucher number is shown, or in its absence, the GenBank accession number.The outgroup is not shown.Black bars represent species recovered by contrasting morphological and genetic information. Red boxes represent species recovered by bPTP analysis. Abbreviations:Morpho. = Morphology; Consens.= Consensus.Photography of Bahius bilineatus by Mauro Teixeira Jr.Photography of Phyllonastes coloma by Luis A.Coloma.Photography of Phyllonastes mindo by Matthijs Hollanders.Photography of Phyllonastes myrmecoides by Mauricio Ortega. Photography of Phyllonastes naturetrekii by Juan Pablo Reyes.Photography of Phyllonastes worleyae by David Brito.	Figure 4. Phylogenetic relationships of Phyllonastes + Urkuphrynegen. nov.(= Noblella northern clade). Maximum likelihood tree obtained for genes 16S, 12S, TYR and RAG1 from a matrix of 3846 bp for 170 terminals.Support values are on the corresponding branches:aLRT values before the slash and non-parametric bootstrap after the slash; triangles on the branches represent aLRT or bootstrap support values of 100%. For each specimen the voucher number is shown, or in its absence, the GenBank accession number.The outgroup is not shown.Black bars represent species recovered by contrasting morphological and genetic information. Red boxes represent species recovered by bPTP analysis. Abbreviations:Morpho. = Morphology; Consens.= Consensus.Photography of Bahius bilineatus by Mauro Teixeira Jr.Photography of Phyllonastes coloma by Luis A.Coloma.Photography of Phyllonastes mindo by Matthijs Hollanders.Photography of Phyllonastes myrmecoides by Mauricio Ortega. Photography of Phyllonastes naturetrekii by Juan Pablo Reyes.Photography of Phyllonastes worleyae by David Brito.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA3FFDBC2C1C3D3956F4D05.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA1FFDEC1D4C04990904829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832715/files/figure.png	https://doi.org/10.5281/zenodo.14832715	Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects.	Figure 10. Live and preserved specimens of Phyllonastes cerrogolondrinas. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 66090, adult male, SVL = 10.89 mm; C, paratype QCAZ 66087, adult female, SVL = 13.75 mm; and D, paratype QCAZ 66089, adult male, SVL = 12.17 mm.A, in life.B–D, in preservative. Scale bars are given only for preserved adults.The depigmentation visible in the specimens is attributable to preservation defects.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA1FFDEC1D4C04990904829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832717/files/figure.png	https://doi.org/10.5281/zenodo.14832717	Figure 11. Palmar (A) and plantar (B) surfaces of Phyllonastes cerrogolondrinas. Photographs of right hand and right foot of the holotype QCAZ 66090. Finger IV of the hand shows some raised skin at the terminal end.	Figure 11. Palmar (A) and plantar (B) surfaces of Phyllonastes cerrogolondrinas. Photographs of right hand and right foot of the holotype QCAZ 66090. Finger IV of the hand shows some raised skin at the terminal end.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA1FFDEC1D4C04990904829.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA4FFD3C2FEC52B96174DEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832721/files/figure.png	https://doi.org/10.5281/zenodo.14832721	Figure 12. Live and preserved specimens of Phyllonastes ecuadoriensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 57064, adult female, SVL = 18.07 mm; and C, paratype QCAZ 48916, subadult female.A, in life.B, C, in preservative.A scale bar is given only for preserved adults. The depigmentation visible in both specimens is attributable to preservation defects.	Figure 12. Live and preserved specimens of Phyllonastes ecuadoriensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 57064, adult female, SVL = 18.07 mm; and C, paratype QCAZ 48916, subadult female.A, in life.B, C, in preservative.A scale bar is given only for preserved adults. The depigmentation visible in both specimens is attributable to preservation defects.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA4FFD3C2FEC52B96174DEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832725/files/figure.png	https://doi.org/10.5281/zenodo.14832725	Figure 13.Palmar (A) and plantar (B) surfaces of Phyllonastes ecuadoriensis. Photographs of left hand and right foot of the holotype QCAZ 57064.	Figure 13.Palmar (A) and plantar (B) surfaces of Phyllonastes ecuadoriensis. Photographs of left hand and right foot of the holotype QCAZ 57064.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA4FFD3C2FEC52B96174DEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832727/files/figure.png	https://doi.org/10.5281/zenodo.14832727	Figure 14.Skulls of Phyllonastes species. A, Phyllonastes ecuadoriensis QCAZ 57064. B, Phyllonastes heyeri QCAZ 24875. C, Phyllonastes macuma QCAZ 40180.	Figure 14.Skulls of Phyllonastes species. A, Phyllonastes ecuadoriensis QCAZ 57064. B, Phyllonastes heyeri QCAZ 24875. C, Phyllonastes macuma QCAZ 40180.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA4FFD3C2FEC52B96174DEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832729/files/figure.png	https://doi.org/10.5281/zenodo.14832729	Figure 15. Skeletons of Phyllonastes species. A, Phyllonastes ecuadoriensis QCAZ 57064, SVL = 18.07 mm. B, Phyllonastes heyeri QCAZ 24875, SVL = 14.11 mm. C, Phyllonastes macuma QCAZ 40180, SVL = 11.92 mm. D, Phyllonastes myrmecoides QCAZ 53583, SVL = 11.13 mm.	Figure 15. Skeletons of Phyllonastes species. A, Phyllonastes ecuadoriensis QCAZ 57064, SVL = 18.07 mm. B, Phyllonastes heyeri QCAZ 24875, SVL = 14.11 mm. C, Phyllonastes macuma QCAZ 40180, SVL = 11.92 mm. D, Phyllonastes myrmecoides QCAZ 53583, SVL = 11.13 mm.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA4FFD3C2FEC52B96174DEC.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA9FFD6C033C0EF96A54EBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832731/files/figure.png	https://doi.org/10.5281/zenodo.14832731	Figure 16. Preserved specimens of Phyllonastes dicaprioi. Lateral, dorsal, and ventral views of: A, holotype DHMECN 13729, adult male, SVL = 10.7 mm; B, paratype DHMECN 13723, adult female, SVL = 12.84 mm; C, paratype DHMECN 13725, adult male, SVL = 12.85 mm; and D, paratype DHMECN 13730, adult female, SVL = 13.56 mm.	Figure 16. Preserved specimens of Phyllonastes dicaprioi. Lateral, dorsal, and ventral views of: A, holotype DHMECN 13729, adult male, SVL = 10.7 mm; B, paratype DHMECN 13723, adult female, SVL = 12.84 mm; C, paratype DHMECN 13725, adult male, SVL = 12.85 mm; and D, paratype DHMECN 13730, adult female, SVL = 13.56 mm.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA9FFD6C033C0EF96A54EBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832735/files/figure.png	https://doi.org/10.5281/zenodo.14832735	Figure 17. Palmar (A) and plantar (B) surfaces of Phyllonastes dicaprioi. Photographs of right hand and left foot of the holotype DHMECN 13729.	Figure 17. Palmar (A) and plantar (B) surfaces of Phyllonastes dicaprioi. Photographs of right hand and left foot of the holotype DHMECN 13729.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFA9FFD6C033C0EF96A54EBD.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFACFFE8C01CC3BF90C44ECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832727/files/figure.png	https://doi.org/10.5281/zenodo.14832727	Figure 14.Skulls of Phyllonastes species. A, Phyllonastes ecuadoriensis QCAZ 57064. B, Phyllonastes heyeri QCAZ 24875. C, Phyllonastes macuma QCAZ 40180.	Figure 14.Skulls of Phyllonastes species. A, Phyllonastes ecuadoriensis QCAZ 57064. B, Phyllonastes heyeri QCAZ 24875. C, Phyllonastes macuma QCAZ 40180.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFACFFE8C01CC3BF90C44ECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832729/files/figure.png	https://doi.org/10.5281/zenodo.14832729	Figure 15. Skeletons of Phyllonastes species. A, Phyllonastes ecuadoriensis QCAZ 57064, SVL = 18.07 mm. B, Phyllonastes heyeri QCAZ 24875, SVL = 14.11 mm. C, Phyllonastes macuma QCAZ 40180, SVL = 11.92 mm. D, Phyllonastes myrmecoides QCAZ 53583, SVL = 11.13 mm.	Figure 15. Skeletons of Phyllonastes species. A, Phyllonastes ecuadoriensis QCAZ 57064, SVL = 18.07 mm. B, Phyllonastes heyeri QCAZ 24875, SVL = 14.11 mm. C, Phyllonastes macuma QCAZ 40180, SVL = 11.92 mm. D, Phyllonastes myrmecoides QCAZ 53583, SVL = 11.13 mm.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFACFFE8C01CC3BF90C44ECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832740/files/figure.png	https://doi.org/10.5281/zenodo.14832740	Figure 18.Preserved specimens of Phyllonastes macuma. Lateral, dorsal, and ventral views of: A, holotype QCAZ 40180, adult male, SVL = 11.92 mm; B, paratype QCAZ 46352, adult female, SVL = 14.49 mm; and C, paratype QCAZ 40181, juvenile. Scale bars are given only for preserved adults. Paratype QCAZ 46352 has a missing shank and foot.	Figure 18.Preserved specimens of Phyllonastes macuma. Lateral, dorsal, and ventral views of: A, holotype QCAZ 40180, adult male, SVL = 11.92 mm; B, paratype QCAZ 46352, adult female, SVL = 14.49 mm; and C, paratype QCAZ 40181, juvenile. Scale bars are given only for preserved adults. Paratype QCAZ 46352 has a missing shank and foot.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFACFFE8C01CC3BF90C44ECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832744/files/figure.png	https://doi.org/10.5281/zenodo.14832744	Figure 19. Palmar (A) and plantar (B) surfaces of Phyllonastes macuma. Photographs of left hand and left foot of the holotype QCAZ 40180.	Figure 19. Palmar (A) and plantar (B) surfaces of Phyllonastes macuma. Photographs of left hand and left foot of the holotype QCAZ 40180.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FFACFFE8C01CC3BF90C44ECA.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF92FFEBC2F4C011965B4CF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832746/files/figure.png	https://doi.org/10.5281/zenodo.14832746	Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults.	Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF92FFEBC2F4C011965B4CF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832748/files/figure.png	https://doi.org/10.5281/zenodo.14832748	Figure 21. Palmar (A) and plantar (B) surfaces of Phyllonastes plateadensis. Photographs of right hand and right foot of the holotype QCAZ 65015.	Figure 21. Palmar (A) and plantar (B) surfaces of Phyllonastes plateadensis. Photographs of right hand and right foot of the holotype QCAZ 65015.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF92FFEBC2F4C011965B4CF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832750/files/figure.png	https://doi.org/10.5281/zenodo.14832750	Figure 22. Skulls of Phyllonastes species.A, Phyllonastes myrmecoides QCAZ 53583. B, Phyllonastes plateadensis QCAZ 65015. C, Phyllonastes sardinayacu, QCAZ 58821.	Figure 22. Skulls of Phyllonastes species.A, Phyllonastes myrmecoides QCAZ 53583. B, Phyllonastes plateadensis QCAZ 65015. C, Phyllonastes sardinayacu, QCAZ 58821.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF92FFEBC2F4C011965B4CF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832752/files/figure.png	https://doi.org/10.5281/zenodo.14832752	Figure 23. Skeletons of Phyllonastes species. A, Phyllonastes plateadensis QCAZ 65015, SVL = 15.70 mm. B, Phyllonastes sardinayacu QCAZ 58821, SVL = 14.32 mm. C, Phyllonastes personinus QCAZ 58818, SVL = 16.91 mm.	Figure 23. Skeletons of Phyllonastes species. A, Phyllonastes plateadensis QCAZ 65015, SVL = 15.70 mm. B, Phyllonastes sardinayacu QCAZ 58821, SVL = 14.32 mm. C, Phyllonastes personinus QCAZ 58818, SVL = 16.91 mm.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF92FFEBC2F4C011965B4CF8.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF91FFEFC2E5C2B890BB4C97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832750/files/figure.png	https://doi.org/10.5281/zenodo.14832750	Figure 22. Skulls of Phyllonastes species.A, Phyllonastes myrmecoides QCAZ 53583. B, Phyllonastes plateadensis QCAZ 65015. C, Phyllonastes sardinayacu, QCAZ 58821.	Figure 22. Skulls of Phyllonastes species.A, Phyllonastes myrmecoides QCAZ 53583. B, Phyllonastes plateadensis QCAZ 65015. C, Phyllonastes sardinayacu, QCAZ 58821.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF91FFEFC2E5C2B890BB4C97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832752/files/figure.png	https://doi.org/10.5281/zenodo.14832752	Figure 23. Skeletons of Phyllonastes species. A, Phyllonastes plateadensis QCAZ 65015, SVL = 15.70 mm. B, Phyllonastes sardinayacu QCAZ 58821, SVL = 14.32 mm. C, Phyllonastes personinus QCAZ 58818, SVL = 16.91 mm.	Figure 23. Skeletons of Phyllonastes species. A, Phyllonastes plateadensis QCAZ 65015, SVL = 15.70 mm. B, Phyllonastes sardinayacu QCAZ 58821, SVL = 14.32 mm. C, Phyllonastes personinus QCAZ 58818, SVL = 16.91 mm.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF91FFEFC2E5C2B890BB4C97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832754/files/figure.png	https://doi.org/10.5281/zenodo.14832754	Figure 24. Live and preserved specimens of Phyllonastes sardinayacu. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 58822, adult female, SVL = 14.10 mm; and C, paratype QCAZ 58821, adult female, SVL = 14.32 mm. A, in life.B, C, in preservative.Scale bars are given for preserved adults.	Figure 24. Live and preserved specimens of Phyllonastes sardinayacu. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 58822, adult female, SVL = 14.10 mm; and C, paratype QCAZ 58821, adult female, SVL = 14.32 mm. A, in life.B, C, in preservative.Scale bars are given for preserved adults.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF91FFEFC2E5C2B890BB4C97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832756/files/figure.png	https://doi.org/10.5281/zenodo.14832756	Figure 25. Palmar (A) and plantar (B) surfaces of Phyllonastes sardinayacu. Photographs of left hand and left foot of the holotype QCAZ 58822.	Figure 25. Palmar (A) and plantar (B) surfaces of Phyllonastes sardinayacu. Photographs of left hand and left foot of the holotype QCAZ 58822.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF91FFEFC2E5C2B890BB4C97.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF9AFFE0C1CCC69091364965.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832758/files/figure.png	https://doi.org/10.5281/zenodo.14832758	Figure 26.Live and preserved specimens of Phyllonastes personinus. A, QCAZ 59437. B, QCAZ 71453.C, QCAZ 31118. A, B, in life. C, in preservative.	Figure 26.Live and preserved specimens of Phyllonastes personinus. A, QCAZ 59437. B, QCAZ 71453.C, QCAZ 31118. A, B, in life. C, in preservative.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF9AFFE0C1CCC69091364965.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832760/files/figure.png	https://doi.org/10.5281/zenodo.14832760	Figure 27. Skeletons of Phyllonastes personinus. A, QCAZ 58818. B, QCAZ 31115.	Figure 27. Skeletons of Phyllonastes personinus. A, QCAZ 58818. B, QCAZ 31115.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF9AFFE0C1CCC69091364965.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF9AFFE2C52FC49990A44C84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832699/files/figure.png	https://doi.org/10.5281/zenodo.14832699	Figure 4. Phylogenetic relationships of Phyllonastes + Urkuphrynegen. nov.(= Noblella northern clade). Maximum likelihood tree obtained for genes 16S, 12S, TYR and RAG1 from a matrix of 3846 bp for 170 terminals.Support values are on the corresponding branches:aLRT values before the slash and non-parametric bootstrap after the slash; triangles on the branches represent aLRT or bootstrap support values of 100%. For each specimen the voucher number is shown, or in its absence, the GenBank accession number.The outgroup is not shown.Black bars represent species recovered by contrasting morphological and genetic information. Red boxes represent species recovered by bPTP analysis. Abbreviations:Morpho. = Morphology; Consens.= Consensus.Photography of Bahius bilineatus by Mauro Teixeira Jr.Photography of Phyllonastes coloma by Luis A.Coloma.Photography of Phyllonastes mindo by Matthijs Hollanders.Photography of Phyllonastes myrmecoides by Mauricio Ortega. Photography of Phyllonastes naturetrekii by Juan Pablo Reyes.Photography of Phyllonastes worleyae by David Brito.	Figure 4. Phylogenetic relationships of Phyllonastes + Urkuphrynegen. nov.(= Noblella northern clade). Maximum likelihood tree obtained for genes 16S, 12S, TYR and RAG1 from a matrix of 3846 bp for 170 terminals.Support values are on the corresponding branches:aLRT values before the slash and non-parametric bootstrap after the slash; triangles on the branches represent aLRT or bootstrap support values of 100%. For each specimen the voucher number is shown, or in its absence, the GenBank accession number.The outgroup is not shown.Black bars represent species recovered by contrasting morphological and genetic information. Red boxes represent species recovered by bPTP analysis. Abbreviations:Morpho. = Morphology; Consens.= Consensus.Photography of Bahius bilineatus by Mauro Teixeira Jr.Photography of Phyllonastes coloma by Luis A.Coloma.Photography of Phyllonastes mindo by Matthijs Hollanders.Photography of Phyllonastes myrmecoides by Mauricio Ortega. Photography of Phyllonastes naturetrekii by Juan Pablo Reyes.Photography of Phyllonastes worleyae by David Brito.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF9AFFE2C52FC49990A44C84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832707/files/figure.png	https://doi.org/10.5281/zenodo.14832707	Figure 7. Ancestral trait reconstruction for discrete morphological characters for Phyllonastes and related genera. The five synapomorphic characters of Phyllonastes are shown. Stars indicate statistically significant support for the most likely character state.	Figure 7. Ancestral trait reconstruction for discrete morphological characters for Phyllonastes and related genera. The five synapomorphic characters of Phyllonastes are shown. Stars indicate statistically significant support for the most likely character state.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF9AFFE2C52FC49990A44C84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832709/files/figure.png	https://doi.org/10.5281/zenodo.14832709	Figure 8. Ancestral trait reconstruction for the terminal expansion of distal phalanges for Phyllonastes and related genera. The characters analysed are the ratio between the terminal expansion and the length of the distal phalanx of the third finger and the fourth toe.	Figure 8. Ancestral trait reconstruction for the terminal expansion of distal phalanges for Phyllonastes and related genera. The characters analysed are the ratio between the terminal expansion and the length of the distal phalanx of the third finger and the fourth toe.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF9AFFE2C52FC49990A44C84.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832746/files/figure.png	https://doi.org/10.5281/zenodo.14832746	Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults.	Figure 20.Live and preserved specimens of Phyllonastes plateadensis. Lateral, dorsal, and ventral views of: A, B, holotype QCAZ 65015, adult female, SVL = 15.70 mm; and C, D, paratype QCAZ 65016, juvenile, SVL = 9.27 mm. A, C, in life.B, D, in preservative. Scale bar is given only for preserved adults.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF99FFFCC039C24D95CA4865.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832762/files/figure.png	https://doi.org/10.5281/zenodo.14832762	Figure 28. Holotype of Urkuphryne merinoi. Lateral, dorsal, and ventral views of holotype, QCAZ 66078, adult female, SVL = 19.53 mm. A, in life.B, in preservative. Scale bar is given only for specimen in preservative.The depigmentation visible in the specimen is attributable to preservation defects.	Figure 28. Holotype of Urkuphryne merinoi. Lateral, dorsal, and ventral views of holotype, QCAZ 66078, adult female, SVL = 19.53 mm. A, in life.B, in preservative. Scale bar is given only for specimen in preservative.The depigmentation visible in the specimen is attributable to preservation defects.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF99FFFCC039C24D95CA4865.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832764/files/figure.png	https://doi.org/10.5281/zenodo.14832764	Figure 29. Palmar (A) and plantar (B) surfaces Urkuphryne merinoi. Photographs of right hand and right foot of the holotype QCAZ 66078.	Figure 29. Palmar (A) and plantar (B) surfaces Urkuphryne merinoi. Photographs of right hand and right foot of the holotype QCAZ 66078.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF99FFFCC039C24D95CA4865.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832766/files/figure.png	https://doi.org/10.5281/zenodo.14832766	Figure 30. Live and preserved specimens of Urkuphryne merinoi. Lateral, dorsal, and ventral views of: A, B, QCAZ 66082, adult male, SVL = 18.33 mm; C, D, QCAZ 66083, adult female, SVL = 19.40 mm; and E, F, QCAZ 66085, adult male, SVL = 18.16 mm. A, C, E, in life. B, D, F, in preservative.Scale bars are given only for preserved adults.	Figure 30. Live and preserved specimens of Urkuphryne merinoi. Lateral, dorsal, and ventral views of: A, B, QCAZ 66082, adult male, SVL = 18.33 mm; C, D, QCAZ 66083, adult female, SVL = 19.40 mm; and E, F, QCAZ 66085, adult male, SVL = 18.16 mm. A, C, E, in life. B, D, F, in preservative.Scale bars are given only for preserved adults.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF99FFFCC039C24D95CA4865.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832768/files/figure.png	https://doi.org/10.5281/zenodo.14832768	Figure 31. Skull of Urkuphryne merinoi holotype QCAZ 66078. The skull is shown in dorsal (A), ventral (B), lateral (C), and frontal (D) views.	Figure 31. Skull of Urkuphryne merinoi holotype QCAZ 66078. The skull is shown in dorsal (A), ventral (B), lateral (C), and frontal (D) views.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF99FFFCC039C24D95CA4865.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832770/files/figure.png	https://doi.org/10.5281/zenodo.14832770	Figure 32. Computed tomography scan images of the skeleton of Urkuphryne merinoi holotype QCAZ 66078, SVL = 19.53 mm, in dorsal (A), ventral (B), and lateral (C) views.	Figure 32. Computed tomography scan images of the skeleton of Urkuphryne merinoi holotype QCAZ 66078, SVL = 19.53 mm, in dorsal (A), ventral (B), and lateral (C) views.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF99FFFCC039C24D95CA4865.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832772/files/figure.png	https://doi.org/10.5281/zenodo.14832772	Figure 33. Computed tomography scan images of the hand and foot osteology of Urkuphryne merinoi holotype QCAZ 66078. The right hand is shown in ventral (A) and dorsal (B) views. The left foot is shown in ventral (plantar) (C) and dorsal (D) views.	Figure 33. Computed tomography scan images of the hand and foot osteology of Urkuphryne merinoi holotype QCAZ 66078. The right hand is shown in ventral (A) and dorsal (B) views. The left foot is shown in ventral (plantar) (C) and dorsal (D) views.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF99FFFCC039C24D95CA4865.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832695/files/figure.png	https://doi.org/10.5281/zenodo.14832695	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	Figure 3.Collection localities ofNoblella northern clade and southern clade species.The map showsthe geographicalgap between thetwo clades. The dashedlines indicatethe southernmostboundary of thenorthern clade and thenorthernmost boundaryof thesouthern clade.Thequestion marks denote individualsidentified asPhyllonastes myrmecoides whoseidentification has notbeen confirmed; theirlocalities are notassigned toeither clade owingto uncertainty regarding their truespecies assignment.Localities arebased onspecimens deposited at Museode Zoologíaof Pontificia Universidad Católicadel Ecuador(QCAZ), InstitutoNacional deBiodiversidad (DHMECN),Museo deZoología of Universidad SanFrancisco de Quito(ZSFQ),and from Lynch(1976, 1986), Hoogmoed and Lescure (1984),Reichle et al.(2004), Dela Rivaet al.(2008a), Guayasaminand Terán- Valdez (2009), Lehr and Catenazzi (2009a), Harvey et al.(2013), Catenazziet al.(2015), Catenazzi and Ttito (2016, 2018, 2019), Reyes-Puig et al. (2019,2020),Santa Cruz et al.(2019),Reyes-Puig et al.(2021), Frost(2023), and GBIF.org (2023). Abbreviations:H,holotype; NC, notconfirmed.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
03D58787FF99FFFCC039C24D95CA4865.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14832774/files/figure.png	https://doi.org/10.5281/zenodo.14832774	Figure 34.Early development of Urkuphryne merinoi and Phyllonastes personinus. A, U. merinoi, clutch of female QCAZ 66078. B, P. personinus, clutch of couple QCAZ 71452–71453. Abbreviations: e, eye; ecd, endolymphatic calcium deposits; et, egg tooth; t, tail; v, fin vasculature.All scales are 1 mm unless otherwise specified.	Figure 34.Early development of Urkuphryne merinoi and Phyllonastes personinus. A, U. merinoi, clutch of female QCAZ 66078. B, P. personinus, clutch of couple QCAZ 71452–71453. Abbreviations: e, eye; ecd, endolymphatic calcium deposits; et, egg tooth; t, tail; v, fin vasculature.All scales are 1 mm unless otherwise specified.	2025-01-13	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.		Zenodo	biologists	Ortega, Jhael A.;Cisneros-Heredia, Diego F.;Camper, Jeffrey D.;Romero-Carvajal, Andrés;Negrete, Leonardo;Ron, Santiago R.			
