taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03D08783FF9F7E145D1585A1FBE16100.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906474/files/figure.png	https://doi.org/10.5281/zenodo.16906474	FIG. 8. Western North American 20-Stamen Black-Fruited Hawthorns (Crataegus Section Douglasianae). A. Crataegus ×suksdorfii (Sarg.) Kruschke; TRT00014409, Heckel,M.& Talent N.NT565, 15 May 2010, triploid,Castlegar,British Columbia.Photo: N.Talent; used with permission.B. Crataegus gaylussacia A.Heller;TRT00002016,Shiller,J., Tusha,J.,Dickinson,T.A.& Heckel, M.PORE-509-1, 26 Apr 2010,triploid,Point Reyes National Seashore, California. Collected under National Park Service permit PORE-2010-SCI-0010. Photo: T.A. Dickinson. C. Crataegus rhodamae-loveae sp. nov. Photo © Keir Morse 2008, Deer Creek Center, Selma, Oregon; used with permission (https://calphotos.berkeley.edu/cgi/img_query?enlar ge=0000+0000+0208+0944).	FIG. 8. Western North American 20-Stamen Black-Fruited Hawthorns (Crataegus Section Douglasianae). A. Crataegus ×suksdorfii (Sarg.) Kruschke; TRT00014409, Heckel,M.& Talent N.NT565, 15 May 2010, triploid,Castlegar,British Columbia.Photo: N.Talent; used with permission.B. Crataegus gaylussacia A.Heller;TRT00002016,Shiller,J., Tusha,J.,Dickinson,T.A.& Heckel, M.PORE-509-1, 26 Apr 2010,triploid,Point Reyes National Seashore, California. Collected under National Park Service permit PORE-2010-SCI-0010. Photo: T.A. Dickinson. C. Crataegus rhodamae-loveae sp. nov. Photo © Keir Morse 2008, Deer Creek Center, Selma, Oregon; used with permission (https://calphotos.berkeley.edu/cgi/img_query?enlar ge=0000+0000+0208+0944).	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9F7E145D1585A1FBE16100.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906462/files/figure.png	https://doi.org/10.5281/zenodo.16906462	FIG. 3. Map of western North America showing the locations at which 233 Crataegus individuals were sampled for DNA extraction and amplification microsatellite loci (Appendix 2; Coughlan et al. 2017b; Han 2013; Han et al. 2013).Taxa sampled at each site indicated by symbol shape and color. Figure produced using R package ggmap (Kahle & Wickham 2013) and Google Maps (Map data © Google).	FIG. 3. Map of western North America showing the locations at which 233 Crataegus individuals were sampled for DNA extraction and amplification microsatellite loci (Appendix 2; Coughlan et al. 2017b; Han 2013; Han et al. 2013).Taxa sampled at each site indicated by symbol shape and color. Figure produced using R package ggmap (Kahle & Wickham 2013) and Google Maps (Map data © Google).	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9F7E145D1585A1FBE16100.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906458/files/figure.png	https://doi.org/10.5281/zenodo.16906458	FIG. 1. PrincipalComponents (PCA;A–C) andCanonical Discriminant (CA;D)Analyses of a Crataegus morphology dataset (N = 246;Appendix 1) with six descriptors (used earlier in Dickinson et al.2008) transformed by ranging to a (0,1) interval:THNL,Thorn length (mm);THND,Thorn diameter at base (mm);relX,Leaf blade length above the widest point divided by leafwidth at the widest point;relZ,Leaf blade lengthbelow the widest point divided by leaf width at the widest point;STAM,Number of stamens per flower (fruit);STYL,Number of styles per flower (fruit).A. PCA biplot showing the component-descriptorcorrelations as vectors.Convexhulls enclosepoints representing specimens of C.suksdorfii sensu lato (suk)with 20 stamens per flower,specimens of C.gaylussacia (gay) with 20 stamens per flower fromMarin and Sonoma countiesin California,and specimens ofC.douglasii(dou) with 10 stamens per flower (Appendix 1). B.Cytotype differentiation of the sample as givenin the inset key.Convexhulls enclose points representing specimens of the different cytotypes.C.Positions of type specimens included in the sample:C.suksdorfii sensu stricto,W.N.Suksdorf numbers 4034, 4419,5026,5031,and5040(holotype andisosyntypes);Br,C.punctatabrevispinacollected byDavidDouglas;G,C.gaylussacia;D,C.douglasii specimens and specimens grown from seed collected by David Douglas;Ba, C.douglasii forma badia; and T,C. tennoWana (the latter two synonymized with C. douglasii. In C,the convex hulls enclose points representing specimens of C.gaylussacia, C.douglasii, and the diploid and polyploid components of C. suksdorfii sensu lato.Specimensrepresenting W.N.Suksdorf4419, theholotype of C.suksdorfii proposed below,are also enclosedwithin a convex hull. D. Canonical Discriminant Analysis of five 20-stamen groups of specimens in our sample:gay,C.gaylussacia; iso4419,W.N.Suksorf 4419, C.suksdorfii sensu lato isosyntype collected in 1905 in ‘Falcon Valley,’Klickitat Co.WA (Love 1998);isoBingen,W.N. Suksorf isosyntypes (4034, 5026, 5031, 5040) collected in1905along theColumbia RiverinBingenWA;suk,allopolyploidC.suksdorfii sensulato;andsuk2x,diploidC.suksdorfiisensu lato.Asterisks indicate that the ordination axis corresponds to a significant eigenvalue according to the broken-stick criterion (PCA) or other criterion (likelihood ratio for canonical axes;Friendly & Fox 2017); axes not associated with eigenvalues judged to be significant in these ways should be compared with the equidistribution among all axes of the total sample variance.	FIG. 1. PrincipalComponents (PCA;A–C) andCanonical Discriminant (CA;D)Analyses of a Crataegus morphology dataset (N = 246;Appendix 1) with six descriptors (used earlier in Dickinson et al.2008) transformed by ranging to a (0,1) interval:THNL,Thorn length (mm);THND,Thorn diameter at base (mm);relX,Leaf blade length above the widest point divided by leafwidth at the widest point;relZ,Leaf blade lengthbelow the widest point divided by leaf width at the widest point;STAM,Number of stamens per flower (fruit);STYL,Number of styles per flower (fruit).A. PCA biplot showing the component-descriptorcorrelations as vectors.Convexhulls enclosepoints representing specimens of C.suksdorfii sensu lato (suk)with 20 stamens per flower,specimens of C.gaylussacia (gay) with 20 stamens per flower fromMarin and Sonoma countiesin California,and specimens ofC.douglasii(dou) with 10 stamens per flower (Appendix 1). B.Cytotype differentiation of the sample as givenin the inset key.Convexhulls enclose points representing specimens of the different cytotypes.C.Positions of type specimens included in the sample:C.suksdorfii sensu stricto,W.N.Suksdorf numbers 4034, 4419,5026,5031,and5040(holotype andisosyntypes);Br,C.punctatabrevispinacollected byDavidDouglas;G,C.gaylussacia;D,C.douglasii specimens and specimens grown from seed collected by David Douglas;Ba, C.douglasii forma badia; and T,C. tennoWana (the latter two synonymized with C. douglasii. In C,the convex hulls enclose points representing specimens of C.gaylussacia, C.douglasii, and the diploid and polyploid components of C. suksdorfii sensu lato.Specimensrepresenting W.N.Suksdorf4419, theholotype of C.suksdorfii proposed below,are also enclosedwithin a convex hull. D. Canonical Discriminant Analysis of five 20-stamen groups of specimens in our sample:gay,C.gaylussacia; iso4419,W.N.Suksorf 4419, C.suksdorfii sensu lato isosyntype collected in 1905 in ‘Falcon Valley,’Klickitat Co.WA (Love 1998);isoBingen,W.N. Suksorf isosyntypes (4034, 5026, 5031, 5040) collected in1905along theColumbia RiverinBingenWA;suk,allopolyploidC.suksdorfii sensulato;andsuk2x,diploidC.suksdorfiisensu lato.Asterisks indicate that the ordination axis corresponds to a significant eigenvalue according to the broken-stick criterion (PCA) or other criterion (likelihood ratio for canonical axes;Friendly & Fox 2017); axes not associated with eigenvalues judged to be significant in these ways should be compared with the equidistribution among all axes of the total sample variance.	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9F7E145D1585A1FBE16100.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906460/files/figure.png	https://doi.org/10.5281/zenodo.16906460	FIG.2.Boxplots contrastingthorn andpetioledimensionsbetween Crataegus gaylussaciaand Crataegussuksdorfii sensulato (andC.douglasii;compare Fig.1; Appendix 1). A,width of thorn at its base inmm.B, lengthof thorn in mm;C,petiole length in mm;note difference inscales between A,B,and C.Data are mean dimensions for samples of five thorns per each of 198 specimens,or a similar number of petioles for 38 specimens.	FIG.2.Boxplots contrastingthorn andpetioledimensionsbetween Crataegus gaylussaciaand Crataegussuksdorfii sensulato (andC.douglasii;compare Fig.1; Appendix 1). A,width of thorn at its base inmm.B, lengthof thorn in mm;C,petiole length in mm;note difference inscales between A,B,and C.Data are mean dimensions for samples of five thorns per each of 198 specimens,or a similar number of petioles for 38 specimens.	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9F7E145D1585A1FBE16100.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906466/files/figure.png	https://doi.org/10.5281/zenodo.16906466	FIG. 5.Boxplots contrasting the density of leaf teeth between diploid and autopolyploid,andallopolyploid,Crataegus suksdorfii sensu lato.Data from isosyntypes (Bingen,Falcon Valley)of C.douglasiivar.suksdorfii Sarg.demonstrate their similarity to theallopolyploids (Appendix 3).Theisosyntypes were collected byW.N.Suksdorf at Bingen,Washington along theColumbia River(WNS4034,5026,5031,and 5040),and at his farm in“Falcon Valley” (WNS4419).Data are mean densities for five leaves for each of the numbers of specimens shown.	FIG. 5.Boxplots contrasting the density of leaf teeth between diploid and autopolyploid,andallopolyploid,Crataegus suksdorfii sensu lato.Data from isosyntypes (Bingen,Falcon Valley)of C.douglasiivar.suksdorfii Sarg.demonstrate their similarity to theallopolyploids (Appendix 3).Theisosyntypes were collected byW.N.Suksdorf at Bingen,Washington along theColumbia River(WNS4034,5026,5031,and 5040),and at his farm in“Falcon Valley” (WNS4419).Data are mean densities for five leaves for each of the numbers of specimens shown.	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9C7E135D3D84F1FD4E62ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906474/files/figure.png	https://doi.org/10.5281/zenodo.16906474	FIG. 8. Western North American 20-Stamen Black-Fruited Hawthorns (Crataegus Section Douglasianae). A. Crataegus ×suksdorfii (Sarg.) Kruschke; TRT00014409, Heckel,M.& Talent N.NT565, 15 May 2010, triploid,Castlegar,British Columbia.Photo: N.Talent; used with permission.B. Crataegus gaylussacia A.Heller;TRT00002016,Shiller,J., Tusha,J.,Dickinson,T.A.& Heckel, M.PORE-509-1, 26 Apr 2010,triploid,Point Reyes National Seashore, California. Collected under National Park Service permit PORE-2010-SCI-0010. Photo: T.A. Dickinson. C. Crataegus rhodamae-loveae sp. nov. Photo © Keir Morse 2008, Deer Creek Center, Selma, Oregon; used with permission (https://calphotos.berkeley.edu/cgi/img_query?enlar ge=0000+0000+0208+0944).	FIG. 8. Western North American 20-Stamen Black-Fruited Hawthorns (Crataegus Section Douglasianae). A. Crataegus ×suksdorfii (Sarg.) Kruschke; TRT00014409, Heckel,M.& Talent N.NT565, 15 May 2010, triploid,Castlegar,British Columbia.Photo: N.Talent; used with permission.B. Crataegus gaylussacia A.Heller;TRT00002016,Shiller,J., Tusha,J.,Dickinson,T.A.& Heckel, M.PORE-509-1, 26 Apr 2010,triploid,Point Reyes National Seashore, California. Collected under National Park Service permit PORE-2010-SCI-0010. Photo: T.A. Dickinson. C. Crataegus rhodamae-loveae sp. nov. Photo © Keir Morse 2008, Deer Creek Center, Selma, Oregon; used with permission (https://calphotos.berkeley.edu/cgi/img_query?enlar ge=0000+0000+0208+0944).	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9C7E135D3D84F1FD4E62ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906480/files/figure.png	https://doi.org/10.5281/zenodo.16906480	FIG.10.Crataegusrhodamae-loveaesp.nov.;one offoursimilar smallflowering shortshoots collectedbyDavid DouglasinApril1825,“neartheconfluence of theColumbia” (Appendix 1; K000370425);represents thepubescententity ignoredby Lindley (who,in any case,was describingthe speciesgrown in the gardenof the Horticultural Society).Hypanthiumlongpubescent,calyx lobes with entire margins,short (<2mm).Twenty-fivepollen grains from this specimen (piece 3,at the bottom right of K000370425,were found to be doubly-stained when treated with Alexander’s stain (Fig.6); no empty grains wereseen.Image© The Board of Trustees of the Royal Botanic Gardens,Kew.Reproduced with theconsent of theRoyal Botanic Gardens,Kew.	FIG.10.Crataegusrhodamae-loveaesp.nov.;one offoursimilar smallflowering shortshoots collectedbyDavid DouglasinApril1825,“neartheconfluence of theColumbia” (Appendix 1; K000370425);represents thepubescententity ignoredby Lindley (who,in any case,was describingthe speciesgrown in the gardenof the Horticultural Society).Hypanthiumlongpubescent,calyx lobes with entire margins,short (<2mm).Twenty-fivepollen grains from this specimen (piece 3,at the bottom right of K000370425,were found to be doubly-stained when treated with Alexander’s stain (Fig.6); no empty grains wereseen.Image© The Board of Trustees of the Royal Botanic Gardens,Kew.Reproduced with theconsent of theRoyal Botanic Gardens,Kew.	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9C7E135D3D84F1FD4E62ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906482/files/figure.png	https://doi.org/10.5281/zenodo.16906482	FIG. 11.Crataegus rhodamae-loveae sp.nov.(a) pre-anthesisflowering branch;(b) anthesis;(c) fruitingbranch;(d) short shoot leaf spectrum;(e)variation in thorn size;(f) flower,pre-anthesis (Kew 000370425);(g) flower at anthesis;(h) median longisection of flower at anthesis;(i) mature fruit;(j) median longisection of mature fruit;(k) median transverse section of mature fruit;(l) individual pyrene.Scale bars = 1.0 cm.	FIG. 11.Crataegus rhodamae-loveae sp.nov.(a) pre-anthesisflowering branch;(b) anthesis;(c) fruitingbranch;(d) short shoot leaf spectrum;(e)variation in thorn size;(f) flower,pre-anthesis (Kew 000370425);(g) flower at anthesis;(h) median longisection of flower at anthesis;(i) mature fruit;(j) median longisection of mature fruit;(k) median transverse section of mature fruit;(l) individual pyrene.Scale bars = 1.0 cm.	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9C7E135D3D84F1FD4E62ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906484/files/figure.png	https://doi.org/10.5281/zenodo.16906484	FIG. 12.Holotype of Crataegus rhodamae-loveae sp.nov.:U.S.A.Oregon.Jackson Co.:N side of Lampman Road above Rogue River,approx.80 m NE of the junction with Highway 99,Aug 2011,J.C.Coughlan, H.Moothoo & C.ShaW JC039 (TRT00020284).	FIG. 12.Holotype of Crataegus rhodamae-loveae sp.nov.:U.S.A.Oregon.Jackson Co.:N side of Lampman Road above Rogue River,approx.80 m NE of the junction with Highway 99,Aug 2011,J.C.Coughlan, H.Moothoo & C.ShaW JC039 (TRT00020284).	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9C7E135D3D84F1FD4E62ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906462/files/figure.png	https://doi.org/10.5281/zenodo.16906462	FIG. 3. Map of western North America showing the locations at which 233 Crataegus individuals were sampled for DNA extraction and amplification microsatellite loci (Appendix 2; Coughlan et al. 2017b; Han 2013; Han et al. 2013).Taxa sampled at each site indicated by symbol shape and color. Figure produced using R package ggmap (Kahle & Wickham 2013) and Google Maps (Map data © Google).	FIG. 3. Map of western North America showing the locations at which 233 Crataegus individuals were sampled for DNA extraction and amplification microsatellite loci (Appendix 2; Coughlan et al. 2017b; Han 2013; Han et al. 2013).Taxa sampled at each site indicated by symbol shape and color. Figure produced using R package ggmap (Kahle & Wickham 2013) and Google Maps (Map data © Google).	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9C7E135D3D84F1FD4E62ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906458/files/figure.png	https://doi.org/10.5281/zenodo.16906458	FIG. 1. PrincipalComponents (PCA;A–C) andCanonical Discriminant (CA;D)Analyses of a Crataegus morphology dataset (N = 246;Appendix 1) with six descriptors (used earlier in Dickinson et al.2008) transformed by ranging to a (0,1) interval:THNL,Thorn length (mm);THND,Thorn diameter at base (mm);relX,Leaf blade length above the widest point divided by leafwidth at the widest point;relZ,Leaf blade lengthbelow the widest point divided by leaf width at the widest point;STAM,Number of stamens per flower (fruit);STYL,Number of styles per flower (fruit).A. PCA biplot showing the component-descriptorcorrelations as vectors.Convexhulls enclosepoints representing specimens of C.suksdorfii sensu lato (suk)with 20 stamens per flower,specimens of C.gaylussacia (gay) with 20 stamens per flower fromMarin and Sonoma countiesin California,and specimens ofC.douglasii(dou) with 10 stamens per flower (Appendix 1). B.Cytotype differentiation of the sample as givenin the inset key.Convexhulls enclose points representing specimens of the different cytotypes.C.Positions of type specimens included in the sample:C.suksdorfii sensu stricto,W.N.Suksdorf numbers 4034, 4419,5026,5031,and5040(holotype andisosyntypes);Br,C.punctatabrevispinacollected byDavidDouglas;G,C.gaylussacia;D,C.douglasii specimens and specimens grown from seed collected by David Douglas;Ba, C.douglasii forma badia; and T,C. tennoWana (the latter two synonymized with C. douglasii. In C,the convex hulls enclose points representing specimens of C.gaylussacia, C.douglasii, and the diploid and polyploid components of C. suksdorfii sensu lato.Specimensrepresenting W.N.Suksdorf4419, theholotype of C.suksdorfii proposed below,are also enclosedwithin a convex hull. D. Canonical Discriminant Analysis of five 20-stamen groups of specimens in our sample:gay,C.gaylussacia; iso4419,W.N.Suksorf 4419, C.suksdorfii sensu lato isosyntype collected in 1905 in ‘Falcon Valley,’Klickitat Co.WA (Love 1998);isoBingen,W.N. Suksorf isosyntypes (4034, 5026, 5031, 5040) collected in1905along theColumbia RiverinBingenWA;suk,allopolyploidC.suksdorfii sensulato;andsuk2x,diploidC.suksdorfiisensu lato.Asterisks indicate that the ordination axis corresponds to a significant eigenvalue according to the broken-stick criterion (PCA) or other criterion (likelihood ratio for canonical axes;Friendly & Fox 2017); axes not associated with eigenvalues judged to be significant in these ways should be compared with the equidistribution among all axes of the total sample variance.	FIG. 1. PrincipalComponents (PCA;A–C) andCanonical Discriminant (CA;D)Analyses of a Crataegus morphology dataset (N = 246;Appendix 1) with six descriptors (used earlier in Dickinson et al.2008) transformed by ranging to a (0,1) interval:THNL,Thorn length (mm);THND,Thorn diameter at base (mm);relX,Leaf blade length above the widest point divided by leafwidth at the widest point;relZ,Leaf blade lengthbelow the widest point divided by leaf width at the widest point;STAM,Number of stamens per flower (fruit);STYL,Number of styles per flower (fruit).A. PCA biplot showing the component-descriptorcorrelations as vectors.Convexhulls enclosepoints representing specimens of C.suksdorfii sensu lato (suk)with 20 stamens per flower,specimens of C.gaylussacia (gay) with 20 stamens per flower fromMarin and Sonoma countiesin California,and specimens ofC.douglasii(dou) with 10 stamens per flower (Appendix 1). B.Cytotype differentiation of the sample as givenin the inset key.Convexhulls enclose points representing specimens of the different cytotypes.C.Positions of type specimens included in the sample:C.suksdorfii sensu stricto,W.N.Suksdorf numbers 4034, 4419,5026,5031,and5040(holotype andisosyntypes);Br,C.punctatabrevispinacollected byDavidDouglas;G,C.gaylussacia;D,C.douglasii specimens and specimens grown from seed collected by David Douglas;Ba, C.douglasii forma badia; and T,C. tennoWana (the latter two synonymized with C. douglasii. In C,the convex hulls enclose points representing specimens of C.gaylussacia, C.douglasii, and the diploid and polyploid components of C. suksdorfii sensu lato.Specimensrepresenting W.N.Suksdorf4419, theholotype of C.suksdorfii proposed below,are also enclosedwithin a convex hull. D. Canonical Discriminant Analysis of five 20-stamen groups of specimens in our sample:gay,C.gaylussacia; iso4419,W.N.Suksorf 4419, C.suksdorfii sensu lato isosyntype collected in 1905 in ‘Falcon Valley,’Klickitat Co.WA (Love 1998);isoBingen,W.N. Suksorf isosyntypes (4034, 5026, 5031, 5040) collected in1905along theColumbia RiverinBingenWA;suk,allopolyploidC.suksdorfii sensulato;andsuk2x,diploidC.suksdorfiisensu lato.Asterisks indicate that the ordination axis corresponds to a significant eigenvalue according to the broken-stick criterion (PCA) or other criterion (likelihood ratio for canonical axes;Friendly & Fox 2017); axes not associated with eigenvalues judged to be significant in these ways should be compared with the equidistribution among all axes of the total sample variance.	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9C7E135D3D84F1FD4E62ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906478/files/figure.png	https://doi.org/10.5281/zenodo.16906478	FIG. 9. Holotype designated here of Crataegus suksdorfii (Sarg.) Kruschke,W.N. Suksdorf 4419, 10 Aug 1905, Falcon Valley, Klickitat Co., Oregon (B; A00018057, the three fragments at the lower right side of the sheet with black fruit). Image courtesy of the Herbarium of the Arnold Arboretum of Harvard University,Cambridge,Massachusetts,USA.Sixty percent of 375 pollen grains from another specimen of this tree(W.N.Suksdorf 4419, 7 Jun 1909;WS142845) were found to be doubly-stained when treated with Alexander’s stain (Fig.6).	FIG. 9. Holotype designated here of Crataegus suksdorfii (Sarg.) Kruschke,W.N. Suksdorf 4419, 10 Aug 1905, Falcon Valley, Klickitat Co., Oregon (B; A00018057, the three fragments at the lower right side of the sheet with black fruit). Image courtesy of the Herbarium of the Arnold Arboretum of Harvard University,Cambridge,Massachusetts,USA.Sixty percent of 375 pollen grains from another specimen of this tree(W.N.Suksdorf 4419, 7 Jun 1909;WS142845) were found to be doubly-stained when treated with Alexander’s stain (Fig.6).	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
03D08783FF9C7E135D3D84F1FD4E62ED.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/16906468/files/figure.png	https://doi.org/10.5281/zenodo.16906468	FIG. 6. Boxplots contrasting pollen fertility in relation to ploidy level and taxon in Crataegus subg.Americanae and Sanguineae (Appendix 4).Fertility indicatedby uptake of all componentsof Alexander’sstain (MalachiteGreen,AcidFuchsin,Orange G;Alexander 1969;see text for details) versusstaining of pollen walls only by Malachite Green.Dataare percentage fully stained grains per specimenin the sites and taxaindicated.Data for the“Falcon Valley”WA holotypeindividual of C.suksdorfii comes from W.N.Suksdorf 4419,7 Jun 1909 (WS142845,which is thesame collection as HUH00018055, Table 2; compare Fig.9).	FIG. 6. Boxplots contrasting pollen fertility in relation to ploidy level and taxon in Crataegus subg.Americanae and Sanguineae (Appendix 4).Fertility indicatedby uptake of all componentsof Alexander’sstain (MalachiteGreen,AcidFuchsin,Orange G;Alexander 1969;see text for details) versusstaining of pollen walls only by Malachite Green.Dataare percentage fully stained grains per specimenin the sites and taxaindicated.Data for the“Falcon Valley”WA holotypeindividual of C.suksdorfii comes from W.N.Suksdorf 4419,7 Jun 1909 (WS142845,which is thesame collection as HUH00018055, Table 2; compare Fig.9).	2023-07-21	Dickinson, Timothy A.;Han, Shery		Zenodo	biologists	Dickinson, Timothy A.;Han, Shery			
