taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805075/files/figure.png	https://doi.org/10.5281/zenodo.14805075	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805083/files/figure.png	https://doi.org/10.5281/zenodo.14805083	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805085/files/figure.png	https://doi.org/10.5281/zenodo.14805085	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805087/files/figure.png	https://doi.org/10.5281/zenodo.14805087	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805089/files/figure.png	https://doi.org/10.5281/zenodo.14805089	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805095/files/figure.png	https://doi.org/10.5281/zenodo.14805095	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805099/files/figure.png	https://doi.org/10.5281/zenodo.14805099	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6DB8534E7DFD8CC02897C6.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805101/files/figure.png	https://doi.org/10.5281/zenodo.14805101	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6BB8504F82FE95C6F79112.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805075/files/figure.png	https://doi.org/10.5281/zenodo.14805075	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6BB8504F82FE95C6F79112.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805087/files/figure.png	https://doi.org/10.5281/zenodo.14805087	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6BB8504F82FE95C6F79112.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B6BB8504F82FE95C6F79112.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805095/files/figure.png	https://doi.org/10.5281/zenodo.14805095	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805075/files/figure.png	https://doi.org/10.5281/zenodo.14805075	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805083/files/figure.png	https://doi.org/10.5281/zenodo.14805083	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805085/files/figure.png	https://doi.org/10.5281/zenodo.14805085	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805087/files/figure.png	https://doi.org/10.5281/zenodo.14805087	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805089/files/figure.png	https://doi.org/10.5281/zenodo.14805089	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805095/files/figure.png	https://doi.org/10.5281/zenodo.14805095	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B68B84E4C27F993C1249070.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805099/files/figure.png	https://doi.org/10.5281/zenodo.14805099	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B76B84F4C47F9C4C31B9071.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805095/files/figure.png	https://doi.org/10.5281/zenodo.14805095	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805077/files/figure.png	https://doi.org/10.5281/zenodo.14805077	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805083/files/figure.png	https://doi.org/10.5281/zenodo.14805083	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805085/files/figure.png	https://doi.org/10.5281/zenodo.14805085	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805097/files/figure.png	https://doi.org/10.5281/zenodo.14805097	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805099/files/figure.png	https://doi.org/10.5281/zenodo.14805099	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805089/files/figure.png	https://doi.org/10.5281/zenodo.14805089	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B75B84A4CE4FACCC39E947C.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805101/files/figure.png	https://doi.org/10.5281/zenodo.14805101	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C23FEF0C158913E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805077/files/figure.png	https://doi.org/10.5281/zenodo.14805077	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C23FEF0C158913E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C23FEF0C158913E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805083/files/figure.png	https://doi.org/10.5281/zenodo.14805083	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C23FEF0C158913E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805085/files/figure.png	https://doi.org/10.5281/zenodo.14805085	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C23FEF0C158913E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805089/files/figure.png	https://doi.org/10.5281/zenodo.14805089	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C23FEF0C158913E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C23FEF0C158913E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805097/files/figure.png	https://doi.org/10.5281/zenodo.14805097	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C23FEF0C158913E.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805093/files/figure.png	https://doi.org/10.5281/zenodo.14805093	Figure 10. Sculpture of Macrogyrus (Macrogyrus) albertisi. (A) metaventrite. Abbreviations: md, metaventral discrimen; tvs, transverse sulcus, scale bar = 1 mm; (B) elytra with canliculate microsculpture, scale bar = 300 µm; (C) canaliculate microsculpture, scale bar = 40 µm.	Figure 10. Sculpture of Macrogyrus (Macrogyrus) albertisi. (A) metaventrite. Abbreviations: md, metaventral discrimen; tvs, transverse sulcus, scale bar = 1 mm; (B) elytra with canliculate microsculpture, scale bar = 300 µm; (C) canaliculate microsculpture, scale bar = 40 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C2EF985C1A89182.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805077/files/figure.png	https://doi.org/10.5281/zenodo.14805077	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4C2EF985C1A89182.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805077/files/figure.png	https://doi.org/10.5281/zenodo.14805077	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805083/files/figure.png	https://doi.org/10.5281/zenodo.14805083	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805085/files/figure.png	https://doi.org/10.5281/zenodo.14805085	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805087/files/figure.png	https://doi.org/10.5281/zenodo.14805087	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805089/files/figure.png	https://doi.org/10.5281/zenodo.14805089	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805093/files/figure.png	https://doi.org/10.5281/zenodo.14805093	Figure 10. Sculpture of Macrogyrus (Macrogyrus) albertisi. (A) metaventrite. Abbreviations: md, metaventral discrimen; tvs, transverse sulcus, scale bar = 1 mm; (B) elytra with canliculate microsculpture, scale bar = 300 µm; (C) canaliculate microsculpture, scale bar = 40 µm.	Figure 10. Sculpture of Macrogyrus (Macrogyrus) albertisi. (A) metaventrite. Abbreviations: md, metaventral discrimen; tvs, transverse sulcus, scale bar = 1 mm; (B) elytra with canliculate microsculpture, scale bar = 300 µm; (C) canaliculate microsculpture, scale bar = 40 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805097/files/figure.png	https://doi.org/10.5281/zenodo.14805097	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805099/files/figure.png	https://doi.org/10.5281/zenodo.14805099	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B72B84A4F8FFC44C1D797C5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805101/files/figure.png	https://doi.org/10.5281/zenodo.14805101	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B73B8484F97FAC0C0949624.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805077/files/figure.png	https://doi.org/10.5281/zenodo.14805077	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B73B8484F97FAC0C0949624.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B73B8484F97FAC0C0949624.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805083/files/figure.png	https://doi.org/10.5281/zenodo.14805083	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B73B8484F97FAC0C0949624.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805087/files/figure.png	https://doi.org/10.5281/zenodo.14805087	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B73B8484F97FAC0C0949624.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B73B8484F97FAC0C0949624.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805093/files/figure.png	https://doi.org/10.5281/zenodo.14805093	Figure 10. Sculpture of Macrogyrus (Macrogyrus) albertisi. (A) metaventrite. Abbreviations: md, metaventral discrimen; tvs, transverse sulcus, scale bar = 1 mm; (B) elytra with canliculate microsculpture, scale bar = 300 µm; (C) canaliculate microsculpture, scale bar = 40 µm.	Figure 10. Sculpture of Macrogyrus (Macrogyrus) albertisi. (A) metaventrite. Abbreviations: md, metaventral discrimen; tvs, transverse sulcus, scale bar = 1 mm; (B) elytra with canliculate microsculpture, scale bar = 300 µm; (C) canaliculate microsculpture, scale bar = 40 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B73B8484F97FAC0C0949624.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805097/files/figure.png	https://doi.org/10.5281/zenodo.14805097	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B73B8484F97FAC0C0949624.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805099/files/figure.png	https://doi.org/10.5281/zenodo.14805099	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8484FB0FE8CC32095E5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805085/files/figure.png	https://doi.org/10.5281/zenodo.14805085	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805075/files/figure.png	https://doi.org/10.5281/zenodo.14805075	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805083/files/figure.png	https://doi.org/10.5281/zenodo.14805083	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805085/files/figure.png	https://doi.org/10.5281/zenodo.14805085	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805087/files/figure.png	https://doi.org/10.5281/zenodo.14805087	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805089/files/figure.png	https://doi.org/10.5281/zenodo.14805089	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805095/files/figure.png	https://doi.org/10.5281/zenodo.14805095	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805099/files/figure.png	https://doi.org/10.5281/zenodo.14805099	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805101/files/figure.png	https://doi.org/10.5281/zenodo.14805101	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B70B8494F90F9FCC0D695B4.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805097/files/figure.png	https://doi.org/10.5281/zenodo.14805097	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805077/files/figure.png	https://doi.org/10.5281/zenodo.14805077	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	Figure 2. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 2. Labels at node denote posterior probability, blue bars indicate 95% hpd for age.The green clades indicate members of Macrogyrus, with the lightest green showing the Macrogyrus s.s. subgenus, the next darkest members of the subgenus Cyclomimus, and the darkest green showing the subgenus Andogyrus stat. nov. Orange indicates the genus Enhydrus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805075/files/figure.png	https://doi.org/10.5281/zenodo.14805075	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	Figure 1. Phylogeny of the Dineutini based on Bayesian tip-dating analysis Part 1. Labels at node denote posterior probability, blue bars indicate 95% hpd for age. The blue clades indicate members of Dineutus, with lighter blue showing the subgenus Cyclous sensu nov. and the dark blue the Dineutus s.s. subgenus. Purple indicates the genus Porrorhynchus. Species are approximately to relative scale.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805079/files/figure.png	https://doi.org/10.5281/zenodo.14805079	Figure 3. Historical biogeography of the dineutine whirligig beetles.The Bayesian tip-dated tree is plotted as used in the biogeographic analysis. Blue bars indicate 95% hpd for age.The circle at the node shows the preferred ancestral state reconstruction from the BioGeoBears results (Figs S12, S13). The following abbreviations are used: DG R, De Geers route; Th R, Thulean route; EECO, Early Eocene Climatic Optimum; DPO, Drake’s Passage opening; MECO, Mid-Eocene Climatic Optimum. The map legend just below the tree shows the colour key for the ancestral state reconstructed. The palaeogeographic maps at bottom, © Colorado Plateau Geosystems used with permission, show continental positions in the major time slices (Blakey, 2008).	Figure 3. Historical biogeography of the dineutine whirligig beetles.The Bayesian tip-dated tree is plotted as used in the biogeographic analysis. Blue bars indicate 95% hpd for age.The circle at the node shows the preferred ancestral state reconstruction from the BioGeoBears results (Figs S12, S13). The following abbreviations are used: DG R, De Geers route; Th R, Thulean route; EECO, Early Eocene Climatic Optimum; DPO, Drake’s Passage opening; MECO, Mid-Eocene Climatic Optimum. The map legend just below the tree shows the colour key for the ancestral state reconstructed. The palaeogeographic maps at bottom, © Colorado Plateau Geosystems used with permission, show continental positions in the major time slices (Blakey, 2008).	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805081/files/figure.png	https://doi.org/10.5281/zenodo.14805081	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	Figure 4. Head capsules of dineutine species, anterior view. Scale bars = 1 mm. Abbreviations: lbr, labrum: cly, clypeus: frb, frontolateral bead. (A) Enhydrus sulcatus; (B) Macrogyrus (Macrogyrus) australis; (C) Dineutus (Cyclous) australis; (D) M. (Andogyrus) seriatopunctatus; (E) Porrorhynchus landaisi; (F) M. (Cyclomimus) purpurascens.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805083/files/figure.png	https://doi.org/10.5281/zenodo.14805083	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	Figure 5. Antennae of dineutine species: above, anterior view; below, posterior. Scale bar = 0.5 mm. (A) Macrogyrus (Macrogyrus) australis; (B) M. (Andogyrus) zimmermanni; (C) Porrorhynchus landaisi; (D) Enhydrus tibialis; (E) Dineutus (Cyclous) australis.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805085/files/figure.png	https://doi.org/10.5281/zenodo.14805085	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	Figure 6. Meso- and meta-ventrites of dineutine species and a gyrinine species, ventral view, scale bars = 1 mm, except (F). (A) Enhydrus sulcatus; (B) Macrogyrus (Andogyrus) colombicus; (C) Porrorhynchus marginatus; (D) Dineutus (Cyclous) carolinus; (E) Mesodineutes amurensis†; (F) Gyrinus maculiventris, scale bar = 0.5 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805087/files/figure.png	https://doi.org/10.5281/zenodo.14805087	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	Figure 7. Protrochanters of male dineutine species, ventral view. Abbreviations: ts, protrochanteric setae; wx, waxy spot; pt, protrochanteric setose patch. (A) Dineutus (Cyclous) australis, scale bar = 200 µm. pb, protrochanteric brush; (B) Dineutus (Cyclous) proximus, scale bar = 500 µm; (C) D. (Dineutus) ‘n. sp.’ Scale bar = 300 µm; (D) D. (Cyclous) serrulatus analis, scale bar = 300 µm; (E) Porrorhynchus marginatus, scale bar = 400 µm; (F) Macrogyrus (Macrogyrus) albertisi, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805089/files/figure.png	https://doi.org/10.5281/zenodo.14805089	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	Figure 8. Prolegs of male dineutine species. Abbreviations: sb, setose brush; asr, anterior row of profemoral setae; psr, posterior row of profemoral setae; sp, setigerous puncture. (A) Macrogyrus (Andogyrus) zimmermanni protibial apex, posterior view, scale bar = 400 µm; (B) Porrorhynchus marginatus protibia, posterior view, scale bar = 500 µm; (C) Dineutus (Cyclous) australis protrochanter and profemur, ventral view, scale bar = 500 µm; (D) M. (A.) zimmermanni protrochanter and profemur, ventral view, scale bar = 1 mm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805091/files/figure.png	https://doi.org/10.5281/zenodo.14805091	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	Figure 9. Protarsus of male dineutine species. Abbreviations: di, protarsal discus; sb, setose brush. (A) Macrogyrus (Andogyrus) zimmermanni, scale bar = 500 µm; (B) M. (Macrogyrus) sp., scale bar = 500 µm; (C) M. (M.) albertisi, scale bar = 500 µm; (D) Porrorhynchus marginatus, scale bar = 1 mm; (E) Dineutus (Dineutus) ‘n. sp.’, scale bar = 1 mm; (F) Enhydrus atratus, scale bar = 2 mm; (G) D. (Cyclous) australis, scale bar = 500 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805093/files/figure.png	https://doi.org/10.5281/zenodo.14805093	Figure 10. Sculpture of Macrogyrus (Macrogyrus) albertisi. (A) metaventrite. Abbreviations: md, metaventral discrimen; tvs, transverse sulcus, scale bar = 1 mm; (B) elytra with canliculate microsculpture, scale bar = 300 µm; (C) canaliculate microsculpture, scale bar = 40 µm.	Figure 10. Sculpture of Macrogyrus (Macrogyrus) albertisi. (A) metaventrite. Abbreviations: md, metaventral discrimen; tvs, transverse sulcus, scale bar = 1 mm; (B) elytra with canliculate microsculpture, scale bar = 300 µm; (C) canaliculate microsculpture, scale bar = 40 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805095/files/figure.png	https://doi.org/10.5281/zenodo.14805095	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	Figure 11. Female reproductive tracts, ventral view. Abbreviations: sp, spermatheca; fd, fertilization duct; ov, common oviduct; bu, bursa; lt, laterotergite; vs, vaginal shield; mp, medial apodeme of gonocoxa; gc, gonocoxa; scale bars = 1 mm. (A) Porrorhynchus landaisi; (B) Dineutus (Dineutus) tetracanthus; (C) D. (Cyclous) discolor; (D) D. (Rhomborhynchus) depressus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805097/files/figure.png	https://doi.org/10.5281/zenodo.14805097	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	Figure 12. Female reproductive tracts. Abbreviations: bg, bursal gland; scale bars = 1 mm. (A) Macrogyrus (Andogyrus) seriatopunctatus, ventral view; (B) gonocoxa of the same; (C) lateral view of the same; (D) M. (Macrogyrus) gouldii, ventral view; (E) gonocoxa of the same; (F) Enhydrus tibialis, ventral view; (G) gonocoxa of the same.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805099/files/figure.png	https://doi.org/10.5281/zenodo.14805099	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	Figure 13. Sperm of dineutine species, exhibiting primary conjugation via spermatostyles. (A) Enhydrus atratus, scale bar = 300 µm; (B) the same, scale bar = 50 µm; (C) the same with single sperm, scale bar = 20 µm; (D) Dineutus emarginatus, scale bar = 50 µm; (E) Macrogyrus (Macrogyrus) rivularis, scale bar = 100 µm; (F) Porrorhynchus marginatus, scale bar = 100 µm.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
03DF87AD3B46B87E4C47FF02C0FA9011.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/14805101/files/figure.png	https://doi.org/10.5281/zenodo.14805101	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	Figure 14. General distribution maps of dineutine genera. (A) Dineutus; (B) Porrorhynchus); (C) Enhydrus; (D) Macrogyrus.	2017-09-30	Gustafson, Grey T;Miller, Kelly B		Zenodo	biologists	Gustafson, Grey T;Miller, Kelly B			
