identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03DDE642FFC0EF1EFB1D9222D9152168.text	03DDE642FFC0EF1EFB1D9222D9152168.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus brachyostomus Viets 1940	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus brachyostomus Viets, 1940 (Figs. 11A–C, Table 4) </p>
            <p>Morphology and notes</p>
            <p>Idiosomal lengths 275–280. Porose areolae and a pair of costae of dorsal plates with rosette pores. AD with three porose areolae and glp-1 on lateral margin of the plate. OC longer than wide, with two cornea. PD anteriorly ovate. Ds-1 on anterior edge of porose areaolae on AD. Pairs of ds-2 and ds-3 on striated integument. Glp lateral to costae of PD at the level of insertion of leg IV. Ds-4 and ds-5 on PD (Fig. 11A). AE with three setae and pair of epimeral pore. PE with one dorsal and two ventral setae. GP large almost rectangular, with a pair of genital acetabula and pair of setae (Fig. 11B). Gnathosoma longer than wide (1:0.6) with four segments (Fig. 11C). Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 3, 4, 5(1), 6; leg II: 1, 2, 3, 4, 4(1), 3; leg III: 1, 1, 2, 3, 5(1), 4; leg IV: 0, 1+2 (basifemur + telofemur), 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae ventromedially and one normal ventral setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Remarks</p>
            <p>  Bartsch (2001) reported  C. brachyostomus from the  Black Sea off the coast of Sinop, Türkiye  .   Later, Durucan (2019) recorded this species from Antalya (  Finike )   among macroalgae, and from Izmir (  Urla )  among oysters (Pinctata radiata). In the present study, the protonymph stage of the species is reported for the first time from Antalya .  The external morphological characteristics of the specimens match those previously described from Sinop, Türkiye . </p>
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	https://treatment.plazi.org/id/03DDE642FFC0EF1EFB1D9222D9152168	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC3EF1FFB3A9603D82F22A2.text	03DDE642FFC3EF1FFB3A9603D82F22A2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus cribosoma (Police 1909)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus cribosoma (Police, 1909) (Figs. 2A–O, 11D–G, Table 1) </p>
            <p>Redescription – Measurements presented in Table 1.</p>
            <p>Morphology and notes</p>
            <p>Female – All dorsal and ventral plates divided by narrow bands of densely striated cuticle and the plates with uniformly rosette pores numerous canaliculi (Figs. 2A, B). AD longer than wide (1.10); anterior margin ovate and with small frontal lamella. Dorsal setae small. Pair of ds-1 on AD posterior to pair of glp-1; ds- 2 in anteromedial margin of OC. Posterior end of OC pointed and almost extending to level of leg III. Gland pores (glp-2) small and lateral to OC. Pore canaliculus posterior to gland pore (Fig. 2C). PD wide, anteriorly truncate, posteriorly ovate. Pair of ds-3 just below the anterior margin of PD, ds-4 and ds-5 at 0.31 and 0.64 of PD's total length, respectively. Pair of ds-6 on anal cone. AE with three pairs of ventral setae and a pair of epimeral pores. Epimeral pores 3–4 wide. Epimeral processes I and II are small and lamellar (Fig. 2D). PE with one dorsolateral and three ventral setae. GA anterior part truncate with uniformly arranged with canaliculi. Ovipositor extending beyond GO but not reaching pgs-1. Three pairs of pgs arranged around GO. GO 54 long, 33 wide. Distance between edges of GA and GO 90, equalling 0.55 of GO length. Genital sclerites with a pair of sgs (Fig. 2E).</p>
            <p>Gnathosoma (Fig. 2H) 1.35 times longer than wide. Rostrum triangular, reaching end of P2. Tectum truncate, palps are slender and consist of four segments. P1 to P4 lengths 12, 36, 9, 34 respectively. P1 and P3 without setae, P2 with single seta, P4 with three setae at basal whorl.</p>
            <p>All legs (Figs. 2 I-O) slender and long but shorter than idiosoma. On the surface of telofemora, genua and tibiae I and II foveate with rosette pores. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 5, 4, 7(2), 6 5; leg II: 1, 2, 5, 4, 6(2), 3; leg III: 1, 2, 3, 3, 5(1), 4; leg IV: 0, 2, 3, 3, 5(1), 3. Tibiae I to IV with 2, 2, 1, 1 bipectinate setae. Tarsi I to IV with 3, 3, 4, 3 dorsal setae and 3, 0, 0, 0 ventral setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Male – Male is similar to female except for the genital plate. GO surrounded by 55–57 pgs (Fig. 2F). Spermatopositor large (78–85), but not extending anterior margin of GA (Fig. 2G). Genital sclerites with four pairs of sgs; anterior pairs hair-like, posterior ones spur-like. GO 40 long, 17 wide. Distance between edges of GA and GO 90, equalling 2.25 of GO length.</p>
            <p>Protonymph – AD with three porose areolae and glp-1 on lateral margin of the plate. OC longer than wide. PD anteriorly ovate. Ds-1 on anterior edge of porose areaolae on AD. Pairs of ds-2 and ds- 3 on striated integument. Ds-4 and ds-5 on PD. Ds-6 on anal cone (Fig. 11D). AE with three setae and pair of epimeral pore. GP large almost quadrangular with a pair of genital acetabula and pair of setae (Fig. 11E). Gnathosoma longer than wide (Fig. 11F). Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 3, 4, 5(1), 6; leg II: 1, 2, 3, 4, 4(1), 3; leg III: 1, 2, 2, 3, 5(1), 4; leg IV: 0, 1+2 (basifemur + telofemur) 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Remarks</p>
            <p>This species was originally described by Police (1909) from Naples, Italy (Tyrrhenian Sea, Mediterranean). Later, Morselli and Mari (1982) reported it from Salento, Italy (Ionian Sea, Mediterranean), in sandy sediment habitats. In this study, it is recorded for the first time from Antalya, Türkiye (Levantine Sea, Mediterranean). The external morphological characteristics of the specimens align with those from Salento, Italy.</p>
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	https://treatment.plazi.org/id/03DDE642FFC3EF1FFB3A9603D82F22A2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC2EF18FB1E95DBDF4C211A.text	03DDE642FFC2EF18FB1E95DBDF4C211A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus figeus Bartsch 1976	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus figeus Bartsch, 1976 (Figs. 3A–J; 4A–G, Table 2) </p>
            <p>Redescription – Measurements presented in Table 2.</p>
            <p>Morphology and notes</p>
            <p>Female – All dorsal and ventral plates divided by narrow bands of striated cuticle and the plates with typical rosette pores which have central ostium surrounded by canaliculi (Figs. 3A, B). AD longer than wide (1.15); anterior margin with small frontal lamella (Fig. 3D).</p>
            <p>AD with pair of rounded wide areolae; anterior areolae with 15–20 rosette pores, two middle areolae each with about 45–50 rosette pores (Fig. 3D). Dorsal setae small. Pair of ds-1 on AD posterior to pair of glp-1; ds- 2 in anteromedial margin of OC. Posterior end of OC rounded and extending to level of leg III. Raised areolae with 20–25 typical rosette pores. OC with two corneae. Gland pores small. Pore canaliculus slightly posterior to gland pore. (Fig. 3E); ds-3 near anterior margin of PD (Fig. 3F), ds-4 and ds-5 at 0.32 and 0.61 of its total length, respectively. Pair of ds-6 on anal cone. PD have two pair of costae which are prominent, but not join anteriorly. Each medial costae with 5-7 and each lateral costae 3–4 rosette pores wide. Panels between two raised costae are very week, cannot seen clearly. Gland pores inconspicuous.</p>
            <p>AE with typical rosette pores (Fig. 3H) but posterior middle part has no porose areas (Fig. 3B). AE with three pairs of ventral setae and a pair of epimeral pores. Epimeral pores 5–7 wide (see Fig. 3B). Epimeral processes I and II are large and lamellar (Fig. 3H). Posterior margin of AE and anterior margin of GA truncate. PE anteriad extending to the level of epimeral gland pores, posteriad almost to the level of posterior margin of GO. PE with one dorsolateral and three ventral setae, porose areolae typical rosette pores.</p>
            <p>GA mostly with typical rosette pores but some area without pores as shown in Figure 3B. Distance from anterior pgs to GO equalling 0.72 the GO's length. Ovipositor extending beyond GO but not reaching pgs-1. Genital sclerites with a pair of sgs. Three pairs of pgs arranged around GO (Fig. 3G).</p>
            <p>Gnathosoma (Figs. 4A–C) 0.67 times longer than wide; with a pair of basirostral setae. Gnathosomal base with pair of areolae distinctly. One pair of maxillary setae on gnathosomal base, other pair half-way along rostrum. Rostrum triangular, reaching end of P2. Tectum triangular almost reaching end of P-1, Rostrum as long as 0.44 of gnathosoma length. Rostral sulcus long, reaching up to 0.60 of rostral length. Palps are slender and consist of four segments. P1 to P4 lengths 10, 40, 10, 37 respectively. P1 and P3 without setae, P2 with single seta, P4 with three setae at basal whorl. P4 shorter than P2.</p>
            <p>All legs (Figs. 4D–G) slender and long but shorter than idiosoma. Telofemora, genua and tibiae with articular membranes; those of tibiae distinctly triangular and pointed. On the surface of telofemora, genua and tibiae I and II foveate. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 5, 4, 7(2), 65; leg II: 1, 2, 5, 4, 7(2), 3; leg III: 1, 2, 2, 3, 5(1), 4; leg IV: 0, 2, 2, 3, 5(1), 3. Telofemura I to IV 1.35, 1.56, 1.53 and 1.86 times longer than high, respectively. Tibiae I to IV with 2, 2, 1, 1 bipectinate setae. Tarsi I to IV with 3, 3, 4, 3 dorsal setae and 3, 0, 0, 0 ventral setae. All legs with two lateral claws with accessory process but delicate pectines (except for leg I) (oil immersion) and a small bidentate median claw. Fossa membrane present but small.</p>
            <p>Male – Male is similar to female except for the genital plate (Fig. 3C). GO surrounded by 31–33 pgs. Genital sclerites with five pairs of sgs; anterior pairs hair-like, posterior ones spur-like (Fig. 3J). GO 42 long, 30 wide. Distance between edges of GA and GO 85, equalling 0.48 of GO length. Spermatopositor large (109) but not extending anterior margin of GA (Fig. 3I).</p>
            <p>Juvenile – Unknown</p>
            <p>Remarks</p>
            <p>  This species was described by Bartsch (1976) based on specimens collected from the upper littoral zone among sediments in the Bay of Arcachon, France.  Specimens from France exhibit a sharper posterior margin on the ocular plate.  In this study, the species is reported for the first time from  Fethiye , Muğla, Türkiye (Levantine Sea, Mediterranean), far from its type locality  . </p>
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	https://treatment.plazi.org/id/03DDE642FFC2EF18FB1E95DBDF4C211A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC5EF19FB6D9690D80C273B.text	03DDE642FFC5EF19FB6D9690D80C273B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus gibbus (Trouessart 1889)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus gibbus (Trouessart, 1889) (Figs. 5A–L, 6A–H, 7A–E, Table 4) </p>
            <p>Redescription – Measurements presented in Table 4.</p>
            <p>Morphology and notes</p>
            <p>Female – All dorsal plates well developed. Porose areolae both dorsal and ventral plates with rosette pores (Fig. 5C). Idiosoma anteriorly with pointed protuberance. AD has “A” shaped internal sclerite. AD rectangular posteriorly. OC with two corneae. PD with single costae and with 3–5 rosette pores wide. Pair of ds-1 near lateral margin of AD. Setae ds-2 on OC. ds-3 to ds-5 on PD. Setae ds-5 long and stout (78 long). Ventral plates uniformly punctated. First epimeral process with prominent epimeral processes both female and male specimens. GA truncate anteriorly with three pairs of setae. Genital sclerites a pair of sgs. Ovipositor extending far beyond GO and pgs-1 (Fig. 5F).</p>
            <p>Gnathosoma (Fig. 5J) 1.38 times longer than wide. Gnathosomal base with pair of distinct areolae. One pair of maxillary setae on gnathosomal base, other pair half-way along rostrum. Rostrum triangular, reaching end of P2. Tectum with large ridge-like process. Rostrum as long as 0.48 of gnathosoma length. Rostral sulcus long, reaching up to 0.6 of rostral length. Palps are slender and consist of four segments. P1 to P4 lengths 13, 35, 10, 40 respectively. P1 and P3 without setae, P2 with single seta, P4 with three setae at basal whorl. P4 longer than P2.</p>
            <p>All legs with lamellae (Fig. 6A–F) Some of lamellae with reticulate patterns as illustrated. All basifemura and telofemora with ventrolateral lamellae. Genua I and II with lamellae, genua III and IV has no lamellae. All tibiae each with two lamellae ventrolaterally and ventromedially. Trochanter III and IV with triangular projection. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae excluded): leg I: 1, 2, 5, 4, 7(2), 65; leg II: 1, 2, 5, 4, 7(1), 3; leg III: 1, 2, 2, 3, 5(1), 3; leg IV: 0, 1, 2, 3, 5(1), 2. Tarsi I to IV with 3, 3, 4, 3 dorsal setae and 32, 0, 0, 0 ventral setae. Delicately pectinate setae exist on basifemora 2, 2, 1, 1, telofemora 1, 1, 0, 0, genua 1, 1, 1, 1 and tibiae 1, 1, 1, 1. All legs with two lateral claws with accessory process and a small bidentate median claw. Fossa membrane well-developed in tarsi I and II, while slender in tarsi III and IV.</p>
            <p>Male – Male is similar to female except for the genital plate (Fig. 5G). Spermatopositor large (115), extending beyond ring of pgs (Fig. 5I). GO 50 long, 38 wide. GO surrounded by 20 pgs. Four pairs of sgs are present; anterior pairs hair-like, posterior ones spur-like (Fig. 5H). Distance between edges of GA and GO 75, equalling 0.66 of GO length.</p>
            <p>Protonymph – Dorsal and ventral plates and porose areolae on plates less developed than adults (Figs. 5B, E). AD with three areolae and glp-1 on lateral margin of the plate. Pair of ds-1 on middle areolae. OC longer than wide, not tail like and posteriorly extending to almost level of leg III insertions. PD with a pair of costae which is two rosette pores wide. AE with three pairs of setae and a pair of epimeral pores. PE with one dorsal and two ventral setae. Genitoanal plate is almost rectangular with genital acetabula. No setae on genital plate. Gnathosoma longer than wide. Tectum triangular and with reticulate patterns. Palps four segmented (Fig. 5K).</p>
            <p>Leg chaetotaxy (Figs. 6G–H, 7A–B): leg I, 1, 2, 3, 4, 5(1), 6; leg II, 1, 2, 3, 4, 5(1), 3; leg III, 1, 1, 2, 3, 5(1), 4; leg IV, 0, 1+2(basifemur + telofemur), 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Larva – Dorsal plates smaller than protonymph. The larva is charecterized by its distinctive big reticulate patterns on dorsal plates. AD with pair of gland pores and pair of setae. OC large with a gland pore and pore canaliculus. PD anteriorly narrow and has one costae. Ds-4 and ds-5 at 0.37 and 0.7 of its total length, respectively. Pair of ds-6 has not seen. Glp-3 on PD situated between ds-4 and ds-5 (Fig. 5A). Gnathosoma longer than wide. Palps four segmented (Fig. 5L). Leg chatetotaxy (Fig. 7C–E); leg I, 1, 1+3, 4, 5, 3; leg II, 1, 1+3, 4, 5, 4; leg III, 0, 1+2, 3, 5, 4. Lamellae not developed (Fig. 7C).</p>
            <p>Remarks</p>
            <p> There are several natural groups within the genus  Copidognathus , one of which is the  gibbus group, comprising 50 valid species (Lee et al., 2023). Three species from this group (  C. gibbus ,  C. majusculatus , and  C. remipes ) have previously been reported from Türkiye (Durucan, 2019).  Copidognathus gibbus was first described by Trouessart (1889) from the littoral zones of the French Atlantic and French Mediterranean. The species was later recorded from fine sand sediments at a depth of 2 m in Antalya (Yakamoz Beach), based on adult specimens (Durucan, 2019). The protonymph stage of  C. gibbus was reported and illustrated by Morselli (1970) from Livorno, Italy (Tyrrhenian Sea). The morphological characteristics of the protonymph specimens from Türkiye correspond with those from Italy. </p>
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	https://treatment.plazi.org/id/03DDE642FFC5EF19FB6D9690D80C273B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC4EF1AFB6090B2DE04268F.text	03DDE642FFC4EF1AFB6090B2DE04268F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus humerosus (Trouessart 1896)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus humerosus (Trouessart, 1896) (Figs. 8A–G, 11H–M, Table 3) </p>
            <p>Redescription – Measurements presented in Table 3.</p>
            <p>Morphology and notes</p>
            <p>Male – All dorsal and ventral plates well developed and divided by narrow bands of densely striated cuticle and the plates with distinctly raised porose areolae with canaliculi (Figs. 8A, B). AD wider than long; anterior margin with small frontal lamella. AD has one large areolae in the middle AD has one large middle areolae with about 30–35 pores. Dorsal setae small. Pair of ds-1 on AD posterior to pair of glp-1; ds- 2 in anteromedial margin of OC. Posterior end of OC pointed and extending to level of leg III. Raised areolae with 10–12 rosette pores. OC with two corneae; longer than wide. Gland pores small. Pore canaliculus slightly posterior to gland pore. Ds-3 below anterior margin of PD, ds-4 and ds-5 at 0.42 and 0.65 of its total length, respectively. Pair of ds-6 on anal cone. PD longer than wide, has only one pair of medial costae. Each costae one to four rosette pores wide. Eleven panels between two raised costae. Gland pores small, one pair immediately lateral to costae at about the level of insertion of leg IV.</p>
            <p>AE with seven long setae (40–50) and a pair of epimeral pores. Epimeral II has two setae instead of one. Epimeral processes I and II are large and lamellar. Posterior margin of AE straight. PE anteriad extending to the level of gland pores, posteriad almost to the level of middle of GO. PE with one dorsolateral and three ventral setae. PE with canaliculi arranged in groups. Genitoanal plate with 20 pgs surrounding genital opening. Each genital sclerite bears four sgs (2:2 grouped) (Fig. 8B).</p>
            <p>Gnathosoma (Fig. 8C) 1.66 times longer than wide. Gnathosomal base with pair of distinct areolae. One pair of maxillary setae on gnathosomal base, other pair half-way along rostrum. Rostrum triangular. Tectum truncate, Rostrum as long as 0.54 of gnathosomal length. Palps are slender and consist of four segments. P1 to P4 lengths 20, 53, 15, 60 respectively. P1 and P3 without setae, P2 with single seta, P4 with three setae at basal whorl. P4 longer than P2.</p>
            <p>All legs (Figs. 8D–G) slender and long but shorter than idiosoma. Leg I to leg IV lengths 341, 316, 380, 336, respectively. Telofemora, genua and tibiae with articular membranes; those of tibiae distinctly triangular and pointed. Surface of telofemora, genua and tibiae I and II foveate. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 5, 4, 7(2), 65; leg II: 1, 2, 5, 4, 7(2), 3; leg III: 1, 2, 3, 3, 5(1), 3?; leg IV: 0, 2, 3, 3, 5(1), 4. Telofemura I to IV 3.33, 3.46, 3.33 and 3.4 times longer than high. Tibiae I to IV with 2, 2, 1, 1 bipectinate setae. Tarsi I to IV with 3, 3, 4?, 43 dorsal setae and 3, 0, 0, 0 ventral setae. All legs with two lateral claws with accessory process. All claws with pecten and number of pecten tines between 20-22 (oil immersion needed) and a small bidentate median claw.</p>
            <p>Protonymph – Dorsal and ventral plates and porose areolae on plates less developed than adults (Fig 11H, I). AD with middle areolae and glp-1 near to ds-1. OC longer than wide. Ds-1 on porose areaolae on AD. Pairs of ds-2 and ds-3 on striated integument. Ds-4 and ds-5 on PD. Ds-6 on anal cone. AE with three setae and pair of epimeral pore. GP rounded with a pair of genital acetabula and patterns as shown with arrow on Figure 11I. Gnathosoma longer than wide (Fig. 11J). Solenidion on tarsi I almost 20 long. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 3, 4, 5(1), 6; leg II: 1, 2, 3, 4, 4(1), 3; leg III: 1, 2, 2, 3, 5(1), 4; leg IV: 0, 1+2 (basifemur + telofemur), 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Larva – AD with pair of gland pores anterior to pair of ds-1; posterior margin truncate (Fig. 11L). Both pairs of OC more rounded than adults and protonymphs, PD and anal cone fused. Ventral side of the specimen was torned therefore not illustrated. Leg III long (195) five-segmented. Basifemura and telofemora III fused. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 4, 4, 5(1), 3?; leg II: 1, 4, 4, 5(1), 3; leg III: 1, 3, 3, 5, 4; Tibiae I to III with 1, 1, 0 bipectinate setae. Tarsi I to III with 3, 3, 4 dorsal setae and 3?, 0, 0 ventral setae. All legs with two long lateral claws with delicate accessory process and a bidentate median claw (Fig. 11M).</p>
            <p> Remarks –  Copidognathus humerosus was originally described by Trouessart (1896) from the French Mediterranean (Marseille, Bay of La Ciotat) at a depth of approximately 45 m depth, and from the French Atlantic (Brest) at 1–5 m depths. Later, Bartsch (1979) also reported the species from the French Atlantic (Roscoff). In this study, the species is recorded for the first time from Fethiye, Muğla (Türkiye, Levantine Sea). The external morphological characteristics of the specimens (male, protonymphs, and larva) are consistent with those from the French coasts. </p>
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	https://treatment.plazi.org/id/03DDE642FFC4EF1AFB6090B2DE04268F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC7EF1BFB15922FDF252137.text	03DDE642FFC7EF1BFB15922FDF252137.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus lamelloides Bartsch 2000	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus lamelloides Bartsch, 2000 (Figs. 12D–F, Table 4) </p>
            <p>Morphology and notes</p>
            <p>Protonymph – Idiosomal length 192 (Fig. 12D). Porose areolae and a pair of costae of dorsal plates with rosette pores. AD with three porose areolae and glp-1 near to porose areolae. OC longer than wide. PD anteriorly ovate. Ds-1 on anterior edge of porose areaolae on AD. Pairs of ds-2 and ds- 3 on striated integument. Glp lateral to costae of PD at the level of insertion of leg IV. Ds-4 and ds-5 on PD. AE with three long (33) setae and pair of epimeral pore (5). GP large, almost quadrangular, a pair of genital acetabula and pair of setae (Fig. 12E). Gnathosoma longer than wide (Fig. 12F). Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 3, 4, 5(1), 6; leg II: 1, 2, 3, 4, 5(1), 3; leg III: 1, 1, 2, 3, 5(1), 4; leg IV: 0, 1+2 (basifemur + telofemur), 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Remarks</p>
            <p> Copidognathus lamelloides  was previously reported from Antalya (  Yakamoz Beach ) in fine sand at a depth of 2 m, based on adult specimens (Durucan, 2019). In this study, the protonymph stage of the species is reported for the first time from Antalya, Türkiye  .   The protonymph stage of  C. lamelloides , as reported and illustrated by  Bartsch (2000) from the Northeastern Atlantic (Baie de Morlaix, French Atlantic), corresponds with the specimen from Türkiye  ."   C. lamelloIdes is reported previously from Antalya (  Yakamoz beach), fine sand at 2 m depth (Durucan 2019) based on adult specimens. In this study, protonymph stage of the species reported for the first time from Antalya, Türkiye   and protonymph stage of  C. lamelloIdes reported and illustrated by  Bartsch (2000) from Northeastern Atlantic (Bai de Morlaix, French Atlantic) accord with the specimen from Türkiye  . </p>
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	https://treatment.plazi.org/id/03DDE642FFC7EF1BFB15922FDF252137	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC6EF1BFB1996B6D872261E.text	03DDE642FFC6EF1BFB1996B6D872261E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus magnipalpus (Police 1909)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus magnipalpus (Police, 1909) (Figs. 12A–C, Table 4) </p>
            <p>Morphology and notes</p>
            <p>Protonymph – Idiosomal length 237–280. Porose areolae of dorsal plates with canaliculi arranged within polygons (Fig. 12A). AD with an anterior and two posterior porose areola. OC with two corneae. PD anteriorly ovate. Ds-1 on anterior edge of porose areaolae on AD. Pairs of ds-2 and ds-3 on striated integument. Ds-4 and ds-5 on PD. AE with three setae and pair of epimeral pore. GP small, ovate, a pair of genital acetabula. No setae on genital plate (Fig. 12B). Tibia I with one smooth and one dentate seta (Fig. 12C). Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 3, 4, 5(1), 6; leg II: 1, 2, 3, 4, 4(1), 3; leg III: 1, 1, 2, 3, 5(1), 4; leg IV: 0, 1+2 (basifemur + telofemur), 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Remarks</p>
            <p>  Adults of  C. magnipalpus were previously reported from several locations in Antalya (Yakamoz Beach, Bilem Beach, and Finike) in macroalgae habitats at various depths (1, 2, and 6 m) (Durucan 2019), as well as from Sinop in the  Black Sea (Bartsch 2001). In this study, the protonymph stage of the species is reported and illustrated for the first time from Antalya, Türkiye  . The protonymph stage, as described and illustrated by Bartsch (2001) from the Black Sea, corresponds with the specimen from the Levantine Sea, Türkiye. </p>
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	https://treatment.plazi.org/id/03DDE642FFC6EF1BFB1996B6D872261E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC6EF14FB29919FD9E3208A.text	03DDE642FFC6EF14FB29919FD9E3208A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus majusculus (Trouessart 1894)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus majusculus (Trouessart, 1894) (Figs. 9A–G, Table 4) </p>
            <p>Morphology and notes</p>
            <p>Female – All dorsal plates separated but very close to each other. Porose areolae both dorsal and ventral plates with rosette pores. Idiosoma anteriorly with pointed protuberance. AD has “A” shaped internal sclerite. AD rectangular posteriorly. PD with single costae and two rosette pores wide. Pair of ds-1 near lateral margin of AD. Setae ds-2 on OC. ds-3 to ds-5 on PD. Ds-6 on anal cone (Fig. 9A). Ventral plates with uniformly punctated. AE with three pairs setae and a pair of epimeral pores. PE with one dorsal and three ventral setae. GA truncate anteriorly with three pairs pgs and one pair of sgs. Ovipositor extending far beyond GO and pgs-1 (Fig. 9B).</p>
            <p>Gnathosoma (Fig. 9D) 1.60 times longer than wide. Gnathosomal base with pair of distinct areolae. One pair of maxillary setae on gnathosomal base, other pair half-way along rostrum. Rostrum triangular. Rostral sulcus long, reaching up to 0.6 of rostral length. Palps are slender and consist of four segments. P1 to P4 13, 33, 10, 54 long, respectively. P1 and P3 without setae, P2 with single seta, P4 with three setae at basal whorl. P4 longer than P2.</p>
            <p>Legs short (Figs. 9 E-G) and with lamellae. Some of lamellae with reticulate patterns as illustrated. All basifemura and telofemora with ventrolateral lamellae. Genua I and II with lamellae, genua III and IV has no lamellae. All tibiae with ventrolateral and ventromedial lamellae. Trochanter III (Fig. 9G) and IV with triangular projection. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 5, 4, 7(2), 6 5; leg II: 1, 2, 5, 4, 7(1), 3; leg III: 1, 2, 2, 3, 5(1), 4 3; leg IV: 0, 1, 2, 3, 5(1), 4 2. Tarsi I to IV with 3, 3, 4, 3 dorsal setae and 2, 0, 0, 0 ventral setae. All legs with two lateral claws with accessory process and a small bidentate median claw. Fossa membrane well-developed in tarsi I and II, while slender in tarsi III and IV.</p>
            <p>Male – Male is similar to female except for the genital plate (Fig. 9C). GO surrounded by 20-21 pgs. Spermatopositor large (110), extending beyond ring of pgs. Four pairs of sgs are present; anterior sgs hair-like, posterior ones spur-like. GO 50 long, 25 wide. Distance between edges of GA and GO 102, equalling 0.49 of GO length.</p>
            <p>Remarks</p>
            <p> Copidognathus majusculatus was originally described by Trouessart (1894) from the French Mediterranean (Marseille) (Bartsch, 2004), and has since been recorded from the North Sea (Helgoland, Germany) (Bartsch, 1977) and the Turkish Mediterranean (Antalya) in fine sand sediment at a depth of 9 m (Durucan, 2019). The specimens reported here were found at a station very close to the latter locality.  C. majusculatus belongs to the  C. gibbus species group, differing from  C. gibbus in the size of its rosette pores and the number of ostia. In  C. gibbus , each pore is large, with an ostium surrounded by up to 20 canaliculi, whereas in  C. majusculatus , the rosette pores and ostia are smaller and surrounded by 2–4 canaliculi (Bartsch, 2004). </p>
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	https://treatment.plazi.org/id/03DDE642FFC6EF14FB29919FD9E3208A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC9EF14FB1D9023D98A2471.text	03DDE642FFC9EF14FB1D9023D98A2471.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus remipes (Trouessart 1894)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus remipes (Trouessart, 1894) (Figs. 12G–J, Table 4) </p>
            <p>Morphology and notes</p>
            <p>Protonymph – Idiosoma 210 long. AD anteriorly has “A” shaped sclerite. OC longer than wide, with two cornea. Gland pores small. Pore canaliculus posterior to gland pore. PD anteriorly and posteriorly ovate with a pair of costae. Ds-1 on anterior edge of porose areaolae on AD. Pairs of ds-2 and ds-3 on striated integument. Glp lateral to costae of PD at the level of insertion of leg IV. Ds-4 and ds-5 on PD. Ds-6 on anal cone (Fig. 12G). AE with three long (33) setae and pair of epimeral pore (5). GP large, almost quadrangular, a pair of genital acetabula. No setae on genital plate (Fig. 12H). Gnathosoma longer than wide. Tectum triangular (Fig. 12I). Leg I short. Telofemura I, genua I and tibia I with lamellae (Fig. 12J).Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 3, 4, 5(1), 6; leg II: 1, 2, 3, 4, 4(1), 3; leg III: 1, 1, 2, 3, 5(1), 4; leg IV: 0, 1+2 (basifemur + telofemur), 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Remarks</p>
            <p> Copidognathus remipes is another species belonging to the  C. gibbus species group. It was previously reported from Antalya (Yakamoz Beach)   in fine sand sediment habitats at a depth of 6 m, based on adult specimens (Durucan 2019).  In this study, the protonymph stage of the species is reported and illustrated for the first time from Antalya, Türkiye  . </p>
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	https://treatment.plazi.org/id/03DDE642FFC9EF14FB1D9023D98A2471	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
03DDE642FFC9EF15FB1D9309D98A2572.text	03DDE642FFC9EF15FB1D9309D98A2572.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Copidognathus tabellio (Trouessart 1894)	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Copidognathus tabellio (Trouessart, 1894) (Figs. 10A–L, Table 4) </p>
            <p>Morphology and notes</p>
            <p>Integument of the specimens have brown pigmentation.</p>
            <p>Female – Porose areolae on dorsal and ventral plates with typical rosette pores. AD with three porose areolae. OC with two corneae. PD with a pair of costae two rosette pores wide. Glp-1 on AD near anterolateral margin of the plate. Ds-1 on AD almost as same level as glp-1. Ds-2 on OC. Ds- 3 in margin of PD (Fig. 10A). GA truncate anteriorly with three pairs pgs and one pair of sgs. Ovipositor extending tip levels with pgs-1 (Fig. 10B). AE with a pair of epimeral pores and three pairs of ventral setae. PE with one dorsal and three ventral setae (Fig. 10B).</p>
            <p>Gnathosoma (Fig. 10G) 1.7 times longer than wide. Rostrum triangular, extending beyond P2. Tectum triangular, Palps are slender and consist of four segments. P1 to P4 lengths 13, 34, 10, 40 respectively. P1 and P3 without setae, P2 with single seta, P4 with three setae at basal whorl. Leg I 185 long (Fig. 10H). Telofemur I laterally reticulated. Length: height ratio 2.0. Tibia IV has five smooth setae. Tarsi IV with 3 dorsal setae (Fig. 10I). All legs (Figs. 2 I-O) slender and long but shorter than idiosoma. On the surface of telofemora, genua and tibiae I and II foveate with rosette pores. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 5, 4, 7(2), 6 5; leg II: 1, 2, 5, 4, 6(2), 3; leg III: 1, 2, 3, 3, 5(1), 4; leg IV: 0, 2, 3, 3, 5(1), 3. Tibiae I to IV with 2, 2, 1, 1 bipectinate setae. Tarsi I to IV with 3, 3, 4, 3 dorsal setae and 3, 0, 0, 0 ventral setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw.</p>
            <p>Male – Male is similar to female except for the genital plate (Fig. 10C). GO surrounded by 24 pgs. Spermatopositor large (110), extending beyond ring of pgs. Four pairs of sgs are present; anterior pairs of sgs hair-like, posterior ones spur-like. GO 50 long, 30 wide. Distance between edges of GA and GO 82.</p>
            <p> Protonymph –  Idiosomal length of only one specimen 275. Porose areolae and a pair of costae of dorsal plates with rosette pores. AD with three porose areolae and glp-1 on lateral margin of the plate. OC longer than wide. PD anteriorly ovate. Ds-1 on anterior edge of porose areaolae on AD. Pairs of ds-2 and ds-3 on striated integument. Ds-4 and ds-5 on PD. AE with three setae and pair of epimeral pore. GP small, ovate, a genital acetabula. No setae on genital plate (Fig. 10J, K). Gnathosoma longer than wide (Fig. 10G). Solenidion on tarsi I 14 long (Fig. 10L). Legs with two lateral claws (Fig. 10L) with accessory process and a small bidentate median claw. Fossa membrane well-developed in tarsi I and II, while slender in tarsi III and IV. Leg chaetotaxy from trochanter to tarsus (bipectinated setae between parentheses, parambulacral setae and solenidia excluded): leg I: 1, 2, 3, 4, 5(1), 6; leg II: 1, 2, 3, 4, 4(1), 3; leg III: 1, 1, 2, 3, 5(1), 4; leg IV: 0, 1+2 (basifemur + telofemur), 3, 5(1), 3. Each of tibiae I to IV with one bipectinate setae. All legs with two lateral claws with accessory process but large pectines (oil immersion) and a small bidentate median claw . </p>
            <p>Remarks</p>
            <p> Copidognathus tabellio was previously reported from Antalya (Finike)   in fine sand at a depth of 6 m, based on adult specimens (Durucan 2019).  In this study, the protonymph stage of the species is reported and illustrated for the first time from Antalya, Türkiye  . </p>
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	https://treatment.plazi.org/id/03DDE642FFC9EF15FB1D9309D98A2572	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Durucan, Furkan	Durucan, Furkan (2024): Three new records and some juveniles of the genus Copidognathus (Acari, Halacaridae) from Türkiye. Persian Journal of Acarology 13: 771-796, DOI: 10.22073/pja.v13i4.85812
