taxonID	type	description	language	source
03DB879AFFFCFFE8B189FF509E6EFC42.taxon	diagnosis	Diagnosis (adapted and slightly modified from Williams 2004). Body of adult female usually broadly oval; anal lobe bars absent. Antennae each with 8 (rarely 7) segments. Legs well developed, each claw without denticle; translucent pores frequently present on hind legs. Circulus present or absent. Anterior and posterior ostioles present. Cerarii numbering 12 – 17 pairs; preocular pair (C 2) always absent. Most cerarii each bearing 2 conical setae, except for those on head and sometimes on thorax, where some with 3 – 5 conical setae each. Anal lobe cerarii usually set on sclerotized cuticle; most species with all cerarii containing auxiliary setae, but these sometimes absent anterior to penultimate cerarii (C 17). Oral rim tubular ducts usually present on dorsum, if absent, then some present on venter. Oral collar tubular ducts present on venter, often of different sizes. Discoidal pores usually present, sometimes 1 or 2 situated adjacent to rim of each oral rim tubular duct, and sometimes a few present next to each eye. Quinquelocular pores always absent.	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFFCFFE8B189FF509E6EFC42.taxon	discussion	Remarks. Molecular phylogenetic analyses of mealybugs by Choi & Lee (2022) and Tanaka et al. (2022) showed that the genus Pseudococcus is non-monophyletic and includes species that belong to several other genera, such as Dysmicoccus Ferris, 1950; Palmicultor Williams, 1963; Paraputo Laing, 1929, etc. The current taxonomic definition of Pseudococcus is therefore probably arbitrary. Further molecular and morphological studies on species currently placed in Pseudococcus and related genera are greatly needed, to clarify the genus boundaries.	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFFCFFECB189FBC79FD6FC11.taxon	description	(Figs 1 and 2) [Japanese common name: Mikan-hime-kona-kaigaramushi]	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFFCFFECB189FBC79FD6FC11.taxon	materials_examined	Material examined. JAPAN: Okinawa I., Nago, around Teniya River, on Citrus sp., 3. vii. 2021, coll. J. Choi, 6 adult females mounted singly (6 OIST); Okinawa I., Nago, around Teniya River, on Citrus sp., 3. vii. 2021, coll. J. Choi, 1 adult female mounted singly (ELKU, DNA-extracted voucher specimen, 210703 JP 18 A). Material examined for comparison. SOUTH KOREA: Jeju Island, on Citrus sp., 13. ix. 2014, coll. J. Choi, 5 adult females mounted singly (5 OIST, from a same sample of DNA-extracted voucher specimen of Pseudococcus cryptus in Choi & Lee 2022).	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFFCFFECB189FBC79FD6FC11.taxon	description	Description (n = 7). Appearance in life. Live adult female found feeding on leaves and stems of host-plant; secreting white powdery wax over almost all body surfaces. Slide-mounted adult female (Figs 1 and 2): Body elongate oval, 1.8 – 2.2 mm long and 0.9 – 1.4 mm wide; derm membranous; segmentation recognizable but not well developed. Anal lobes clearly observable, dorsal surface of each lobe with well-sclerotized plate and ventral surface with long apical seta, 112 – 181 µm long; anal lobe bar absent. Antenna 430 – 470 µm long, with 8 segments and many flagellate setae; subapical segment with 1 fleshy seta and apical segment with 3 fleshy setae. Eyes situated on margins, not associated with discoidal pores. Legs well developed, with many flagellate setae; hind trochanter + femur 325 – 344 µm long; hind tibia + tarsus 343 – 370 µm long; claw 27 – 33 µm long without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur about 1.0 – 1.1: 1; ratio of lengths of hind tibia to tarsus 2.5 – 3.5: 1. Paired setose tarsal digitules present, subequal in length, knobbed; claw digitules minutely knobbed. Hind coxae, femora, and tibiae with translucent pores on hind (= dorsal) surfaces; hind trochanters and tarsi without translucent pores (Fig. 2). Labium 114 – 117 µm long, shorter than clypeus. Circulus present, well developed, divided by intersegmental line, located between abdominal segments III and IV, 81 – 160 µm long and 128 – 164 µm wide. Ostioles present, each with inner edges of lips weakly sclerotized; anterior ostioles each with a total for both lips of 10 – 22 trilocular pores and 1 – 5 setae; each posterior ostiole with a total for both lips of 12 – 19 trilocular pores and 0 – 4 setae. Anal ring 80 – 86 µm wide, bearing 6 setae, each seta 108 – 160 µm long. Cerarii numbering 17 pairs. Anal lobe cerarii (C 18) each present on well sclerotized area, usually containing 2 conical cerarian setae, each seta 20 – 34 µm long and about 8 – 13 µm wide at base; 2 – 5 auxiliary setae and a concentration of trilocular pores. Penultimate cerarii (C 17) each containing 2 conical cerarian setae, 2 – 5 auxiliary setae and a concentration of trilocular pores. Cerarii situated further forward generally each with 1 – 4 conical cerarian setae, a few auxiliary setae and a concentration of trilocular pores. Dorsum. Setae flagellate, each 14 – 120 µm long, distributed segmentally; longest setae present on head or posterior abdominal segments. Trilocular pores, each 2 – 4 µm wide, evenly distributed. Oral rim tubular ducts each about 10 – 12 µm in diameter, numbering 0 – 3 on marginal areas of thoracic segments and head. Oral collar tubular ducts each about 3 – 5 µm in diameter, mostly each wider than a trilocular pore, present on marginal areas of thoracic segments and head. Discoidal pores, each 1 – 2 µm wide, sparsely distributed, frequently associated with oral rim tubular ducts, oral collar ducts and some cerarii. Venter. Setae relatively long and flagellate, each 16 – 120 µm long; longest setae situated on medial area of head. Multilocular disc-pores, each 7 – 9 µm wide, present on medial areas of abdominal segments IV ‒ IX. Trilocular pores, same width as those on dorsum, sparsely distributed, frequently associated with oral rim tubular ducts and some cerarii. Oral rim tubular ducts, each about 7 – 10 µm in diameter, present in groups of 1 – 5 on margins of thoracic segments and on abdominal segments. Oral collar tubular ducts of 3 sizes: (i) large-type ducts, each about 4 – 5 µm wide, mostly wider than trilocular pores, present on marginal areas of whole body; (ii) small-type ducts, each about 2 – 3 μm in diameter, present on submarginal to submedial areas of abdominal and thoracic segments; and (iii) minute-type ducts, each about 1 – 2 um in diameter, mostly narrower than a trilocular pore, present on medial areas of posterior abdominal segments. Discoidal pores, same width as those on dorsum, sparsely distributed, frequently associated with oral rim tubular ducts and some cerarii. Host-plants in Japan. Citrus sp. (Rutaceae) (Kawai 1980), Fatsia japonica (Araliaceae) (Kawai 1980).	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFFCFFECB189FBC79FD6FC11.taxon	discussion	Remarks. The specimens of P. cryptus from Okinawa I., described above, differ slightly from those described by Williams (2004) in lacking dorsal oral collar tubular ducts associated with the abdominal cerarii, whereas Williams (2004) says that P. cryptus has narrower and longer or wide-type dorsal oral collar tubular ducts present next to most abdominal cerarii. This morphological discrepancy is probably due to individual and / or regional intraspecific variation in P. cryptus. The molecular phylogenetic analysis showed that the examined specimens above are closely related to P. cryptus collected on Jeju Island, Korea (Fig. 7). Five South Korean specimens belonging to the same sample as the individual used for our molecular phylogenetic analysis (Fig. 7) were examined in this study and they all have 3 to 5 wide-type oral collar ducts next to the abdominal cerarii (from C 11 to C 18, data not shown). This may indicate that, at least in this and the related species, the presence or absence of oral collar ducts next to the abdominal cerarii may be of limited taxonomic value. However, further morphological and molecular phylogenetic studies are probably needed to confirm the taxonomic significance of this morphological characteristic.	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFF8FFE3B189FC799A96FE41.taxon	description	(Plate 1 A, Figs 3 and 4) [Japanese common name: Matsuura-kona-kaigaramushi]	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFF8FFE3B189FC799A96FE41.taxon	materials_examined	Material examined. JAPAN: Okinawa I., Urasoe, Iso, Urasoe Dai Park on Murraya paniculata, 14. xi. 2020, coll. H. Tanaka, 3 adult females mounted on separate slides (2 ELKU, 1 EUMJ); Okinawa I., Onna-son, on Gymnosporia diversifolia, 29. x. 2022, coll. J. Choi, 1 adult female mounted singly (ELKU, DNA extracted voucher specimen, 221029 JP 01); Okinawa I., Nago, Kyoda, on Gymnosporia diversifolia, 17. v. 2023, coll. J. Choi, 8 adult females mounted on separate slides (4 ELKU, 4 EUMJ).	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFF8FFE3B189FC799A96FE41.taxon	description	Description (n = 12). Appearance in life (Plate 1 A): Live adult female found feeding on leaves and stems of the host-plant; secreting white powdery wax over almost all body surfaces (Plate 1 A). Slide-mounted adult female (Figs 3 and 4): Body elongate oval, 2.7 – 4.0 mm long and 1.5 – 2.6 mm wide; derm membranous; segmentation recognizable, but not well developed. Anal lobes clearly observable, dorsal surface of each lobe with well-sclerotized plate and ventral surface with long apical seta, 94 – 183 µm long; anal lobe bar absent. Antenna 415 – 560 µm long, with 8 segments and many flagellate setae; subapical segment with 1 fleshy seta, apical segment with 3 fleshy setae. Eyes situated on margin, not associated with discoidal pores. Legs well developed, with many flagellate setae; hind trochanter + femur 327 – 467 µm long; hind tibia + tarsus 352 – 480 µm long; each claw 24 – 40 µm long without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur about 0.97 – 1.16: 1; ratio of lengths of hind tibia to tarsus 2.64 – 3.90: 1. Paired setose tarsal digitules present, subequal in length, knobbed; claw digitules minutely knobbed. Translucent pores absent from hind coxae, present on hind (= dorsal) surfaces of femora and tibiae, but absent from tarsi; hind trochanters without translucent pores (Fig. 3). Labium 114 – 160 µm long, shorter than clypeus. Circulus present, well developed, divided by intersegmental line, located between posterior abdominal segments III and IV, 60 – 222 µm long and 150 – 257 µm wide. Ostioles present, each with inner edges of lips weakly sclerotized; anterior ostioles each with a total for both lips of 16 – 35 trilocular pores and 1 – 8 setae; each posterior ostiole with a total for both lips of 19 – 34 trilocular pores and 0 – 6 setae. Anal ring 92 – 122 µm wide, bearing 6 setae, each seta 130 – 187 µm long. Cerarii numbering 17 pairs. Anal lobe cerarii (C 18) each situated on well-sclerotized area, usually containing 2 conical cerarian setae (each seta 18 – 34 µm long and about 9 – 13 µm wide at base), 2 – 7 auxiliary setae and concentration of trilocular pores. Penultimate cerarii (C 17) each containing 2 (rarely 1) conical cerarian setae, 1 – 8 auxiliary setae and concentration of trilocular pores. Cerarii situated further forward generally each with 1 – 4 conical cerarian setae, few auxiliary setae and concentration of trilocular pores. Dorsum. Setae flagellate, each 12 – 64 µm long, distributed segmentally. Trilocular pores, each 2 – 4 µm wide, evenly distributed. Oral rim tubular ducts, each about 10 – 13 µm in diameter, numbering 13 – 49 in marginal and submarginal areas, on midline of posterior abdominal segments and on submedial areas of abdominal segments, occasionally on medial and submedial areas of thoracic segments and head. Oral collar tubular ducts, each about 3 – 4 µm in diameter, mostly each wider than a trilocular pore, sparsely distributed but mostly concentrated in marginal to submarginal areas. Discoidal pores, each approximately 1 µm wide, sparsely distributed, few present in cerarii and frequently associated with oral rim tubular duct and oral collar tubular duct orifices. Venter. Setae relatively long and flagellate, each 16 – 176 µm long; longest setae situated on medial area of head. Multilocular disc-pores, each 7 – 9 µm wide, present on medial areas of abdominal segments IV ‒ IX. Trilocular pores, same width as those on dorsum, distributed evenly. Discoidal pores present or absent, if present each approximately 1 µm in diameter, sparsely distributed, frequently associated with orifices of oral rim tubular ducts and largest type of oral collar tubular ducts. Oral rim tubular ducts, each about 7 – 9 µm in diameter, present in groups of 1 ‒ 3 on margins of thoracic and abdominal segments. Oral collar tubular ducts of 3 sizes: (i) large-type ducts, each about 3 – 4 µm wide, mostly wider than a trilocular pore, present on marginal areas of whole body; (ii) small-type ducts, each about 2 – 3 μm in diameter, present in transverse row across each of posterior abdominal segments; and (iii) minute-type ducts, each approximately 1 – 2 um in diameter, mostly narrower than a trilocular pore, present on medial areas of posterior abdominal segments. Host-plants in Japan. Gymnosporia diversifolia (Celastraceae), Citrus spp. (Tokihiro 2004), Murraya paniculata (Rutaceae).	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFF8FFE3B189FC799A96FE41.taxon	discussion	Remarks. Pseudococcus odermatti is similar to P. comstocki (Kuwana, 1902) in having dorsal oral collar tubular ducts across some segments (Williams 2004). However, P. odermatti differs from P. comstocki (contrasting character states in P. comstocki in parentheses) as follows: (i) hind coxae lacking translucent pores (hind coxae with translucent pores); and (ii) abdominal segment II lacking multilocular disc-pores (abdominal segment II with multilocular disc-pores). The molecular phylogenetic analysis also indicated that P. odermatti is closely related to P. comstocki (Fig. 7); however, further detailed morphological and molecular phylogenetic studies are needed to define the taxonomic status of P. odermatti further.	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFF7FFE6B189FDC99F70FD44.taxon	description	(Plate 1 B, Figs 5 and 6) [Japanese common name: Monpanoki-kona-kaigaramushi]	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFF7FFE6B189FDC99F70FD44.taxon	materials_examined	Material examined. Holotype: Japan: / Okinawa Is., Itomon, / Nishizaki, / on Heliotropium arboreum, / 10. iii. 2023, / coll. H. Tanaka and T. Uesato, / adult female mounted singly on a slide (ELKU, HT 179). Paratypes: same data as for holotype, 7 adult females mounted on separate slides (3 ELKU, 4 EUMJ, HT 179); 1 adult female mounted singly (ELKU, DNA-extracted voucher specimen, HT 179); Japan: / Okinawa Is., Itomon, / Nishizaki, / on Heliotropium arboreum, / 13. iii. 2023, / coll. H. Tanaka, / 7 adult females mounted on separate slides (4 ELKU, 3 EUMJ, HT 190); Japan: / Okinawa Is, Onna-son, / Tancha, / on Pandanus odoratissimus / 13. v. 2021, / coll. J. Choi, / 4 adult females mounted on separate slides (2 ELKU, 2 EUMJ, 210513 JP 09). Materials examined for comparison. Japan: Bonin Is. (Ogasawara Is.), Chichi-jima I., Komagari, 3 adult females on Celtis boninensis, 26. xii. 1969, coll. S. Kawai, mounted on single slide with other 2 adult females and 4 immature stage females, a part of syntype of Pseudococcus ogasawaraensis Kawai, 1973 (3 KTUA).	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFF7FFE6B189FDC99F70FD44.taxon	description	Description (n = 20) Appearance in life (Plate 1 B): Live adult females found feeding on leaves and stems of the host-plant; secreting white to pinkish powdery wax on almost all body surfaces. Slide-mounted adult female (Figs 5 and 6): Body elongate oval, 2.9 (2.0 – 3.9) mm long and 1.6 (0.9 – 2.2) mm wide; derm membranous; segmentation recognizable, but not well developed. Anal lobes clearly observable, dorsal surface of each lobe with well-sclerotized plate and ventral surface with long apical seta, 182 (120 – 201) µm long; anal lobe bar absent. Antenna 531 – 549 (453 – 551) µm long, with 8 segments and many flagellate setae; subapical segment with 1 fleshy seta, apical segment with 3 fleshy setae. Eyes situated on margins, not associated with discoidal pores. Legs well developed, with many flagellate setae; hind trochanter + femur 389 – 390 (334 – 415) µm long; hind tibia + tarsus 425 – 426 (369 – 447) µm long; claw 34 (28 – 37) µm long, without a denticle. Ratio of lengths of hind tibia + tarsus to trochanter + femur about 1.10 – 1.09 (1.05 – 1.17): 1; ratio of lengths of hind tibia to tarsus 2.94 – 3.05 (2.45 – 4.00): 1. Paired setose tarsal digitules present, subequal in length, clubbed; claw digitules minutely knobbed. Hind leg coxae usually with translucent pores on both surfaces; femora and tibiae usually with translucent pores on hind (= dorsal) surface; rarely, tarsi with translucent pores on front (= ventral) surface; and hind trochanters without translucent pores (Fig. 6). Labium 130 (128 – 156) µm long, shorter than clypeus. Circulus present, well developed, divided by intersegmental line, located between posterior abdominal segments III and IV, 150 (57 – 152) µm long and 129 (54 – 190) µm wide. Ostioles present, each with inner edges of lips weakly sclerotized; anterior ostioles each with a total for both lips of 22 – 23 (14 – 32) trilocular pores and 4 – 5 (0 – 9) setae; each posterior ostiole with a total for both lips of 19 – 26 (14 – 31) trilocular pores and 5 – 7 (2 – 8) setae. Anal ring 100 (97 – 110) µm wide, bearing 6 setae, each seta 90 – 153 (90 – 180) µm long. Cerarii numbering 17 pairs. Anal lobe cerarii (C 18) present on well-sclerotized area, usually each containing 2 conical cerarian setae, each seta 28 – 30 (18 – 38) µm long and about 10 – 11 (8 – 12) µm wide at base; 7 (3 – 10) auxiliary setae and a concentration of trilocular pores. Penultimate cerarii (C 17) each containing 2 conical cerarian setae, 5 – 6 (3 – 7) auxiliary setae and a concentration of trilocular pores. Cerarii situated further forward generally each with 1 – 4 conical cerarian setae, a few auxiliary setae and a concentration of trilocular pores. Dorsum. Setae flagellate, each 21 – 51 (14 – 55) µm long, distributed segmentally; longest setae usually present on head or posterior abdominal segments. Trilocular pores, each 2 – 4 µm wide, evenly distributed. Oral rim tubular ducts, each about 10 – 11 (8 – 12) µm in diameter, 19 (13 – 21) duct present marginal area, in longitudinal line of posterior abdominal segments and occasionally in longitudinal line of thoracic segments and rarely in submedial area of abdominal segments. Oral collar tubular ducts each about 3 – 4 µm in diameter, mostly wider than trilocular pores, present on marginal area of thoracic segments and head. Discoidal pores, approximately 1 µm wide, sparsely distributed, frequently associated with oral rim tubular ducts, oral collar tubular ducts and in some cerarii. Venter. Setae relatively long and flagellate, each 20 – 109 (12 – 168) µm long; longest setae on medial area of head. Multilocular disc-pores, each 6 – 8 (6 – 10) µm wide, present on medial area of abdominal segments IV to IX. Trilocular pores, same width as those on dorsum, evenly distributed. Oral rim tubular ducts, each about 6 – 7 (6 – 8) µm in diameter, present in groups of 1 – 3 on margins of thoracic and abdominal segments. Oral collar tubular ducts of 3 sizes: (i) large-type ducts, each about 3 – 4 (3 – 5) µm wide, mostly wider than a trilocular pore, present on marginal and submarginal areas of entire body; (ii) small-type ducts, each about 2 – 3 μm in diameter, present on submarginal to medial areas of abdominal segments; and (iii) minute-type ducts, each about 1 um in diameter, mostly narrower than a trilocular pore, present on medial areas of posterior abdominal segments. Discoidal pores, same width as those on dorsum, sparsely distributed, frequently associated with orifices of oral rim tubular ducts and largest type of oral collar ducts. Host-plants. Heliotropium arboreum (Boraginaceae), Pandanus odoratissimus (Pandanaceae).	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
03DB879AFFF7FFE6B189FDC99F70FD44.taxon	discussion	Remarks. Pseudococcus okinawaensis Tanaka & Choi sp. nov. is similar to P. ogasawarensis Kawai, 1973 in having translucent pores on both surfaces (anterior and posterior surfaces) of hind coxae, and in sharing other general morphological features. However, the new species differs from P. ogasawarensis (contrasting character states in P. ogasawarensis in parentheses) as follows: (i) total number of dorsal discoidal pores associated with the orifices of oral collar tubular ducts and rims of oral rim tubular ducts 9 – 16 (total number of dorsal discoidal pores associated with the orifices of oral collar tubular ducts and rims of oral rim tubular ducts 0 – 4); (ii) penultimate cerarii (C 17) situated on non-sclerotized cuticle (penultimate cerarii (C 17) situated on slightly sclerotized cuticle). The species is also similar to P. gilbertensis Beardsley, 1966 in having translucent pores on both surfaces (anterior and posterior surfaces) of hind coxae, and in sharing other general morphological features. However, the new species differs from P. gilbertensis (contrasting character states in P. gilbertensis in parentheses) as follows: (i) dorsal oral collar tubular ducts mostly wider than trilocular pores (dorsal oral collar tubular ducts narrower than trilocular pores); (ii) penultimate cerarii (C 17) situated on non-sclerotized cuticle (penultimate cerarii (C 17) situated on sclerotized cuticle). In the molecular phylogenetic tree, the three populations of P. okinawaensis clustered together into a single clade that was sister to the clade of P. cryptus (Fig. 7). The host-plants of P. okinawaensis are very common and endemic to Okinawa, suggesting that the species is probably endemic to the island.	en	Tanaka, Hirotaka, Choi, Jinyeong, Kamitani, Satoshi (2025): Taxonomic review of three species of the genus Pseudococcus Westwood, 1840 (Hemiptera: Coccomorpha: Pseudococcidae) found in Okinawa Island, Japan, with description of a new species. Zootaxa 5637 (3): 579-593, DOI: 10.11646/zootaxa.5637.3.6, URL: https://doi.org/10.11646/zootaxa.5637.3.6
