taxonID	type	format	identifier	references	title	description	created	creator	contributor	publisher	audience	source	license	rightsHolder	datasetID
03DBCE50FFF43B1CFF5FFA03FEE6FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262930/files/figure.png	https://doi.org/10.5281/zenodo.15262930	FIGURE 1. Map of Madagascar showing sampling localities of Lygodactylus ornatus (near Mandritsara) and L. namoroka sp. nov. (Tsingy de Namoroka National Park). The base map shows vegetation across Madagascar from the Madagascar Vegetation Mapping Project, available at https://web.archive.org/web/20180419112513/http://www.vegmad.org/). Vegetation is colored as follows: green, humid forest (rainforest); red, western dry deciduous forest; pink, mangroves; orange, spiny forest-thicket; greenish blue, western subhumid forest; yellow, Tapia forest.	FIGURE 1. Map of Madagascar showing sampling localities of Lygodactylus ornatus (near Mandritsara) and L. namoroka sp. nov. (Tsingy de Namoroka National Park). The base map shows vegetation across Madagascar from the Madagascar Vegetation Mapping Project, available at https://web.archive.org/web/20180419112513/http://www.vegmad.org/). Vegetation is colored as follows: green, humid forest (rainforest); red, western dry deciduous forest; pink, mangroves; orange, spiny forest-thicket; greenish blue, western subhumid forest; yellow, Tapia forest.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFF43B1CFF5FFA03FEE6FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262933/files/figure.png	https://doi.org/10.5281/zenodo.15262933	FIGURE 2. Habitat of Lygodactylus ornatus in the Mandritsara area, photographed on 23–24 October 2023.	FIGURE 2. Habitat of Lygodactylus ornatus in the Mandritsara area, photographed on 23–24 October 2023.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFF43B1CFF5FFA03FEE6FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262935/files/figure.png	https://doi.org/10.5281/zenodo.15262935	FIGURE 3. Individuals of Lygodactylus ornatus from the Mandritsara area in dorsolateral view. A, ZSM 79/2023 (ZCMV 13743); B, ZSM 80/2023 (ZCMV 13745); C, UADBA-ZCMV 13746; D, ZSM 81/2023 (ZCMV 13747). Not to scale.	FIGURE 3. Individuals of Lygodactylus ornatus from the Mandritsara area in dorsolateral view. A, ZSM 79/2023 (ZCMV 13743); B, ZSM 80/2023 (ZCMV 13745); C, UADBA-ZCMV 13746; D, ZSM 81/2023 (ZCMV 13747). Not to scale.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFF43B1CFF5FFA03FEE6FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262937/files/figure.png	https://doi.org/10.5281/zenodo.15262937	FIGURE 4. Individuals of Lygodactylus ornatus from the Mandritsara area in ventral view. A, ZSM 79/2023 (ZCMV 13743); B, UADBA-ZCMV 13744; C, ZSM 81/2023 (ZCMV 13747); D, ZSM 80/2023 (ZCMV 13745); E, UADBA-ZCMV 13746. Not to scale.	FIGURE 4. Individuals of Lygodactylus ornatus from the Mandritsara area in ventral view. A, ZSM 79/2023 (ZCMV 13743); B, UADBA-ZCMV 13744; C, ZSM 81/2023 (ZCMV 13747); D, ZSM 80/2023 (ZCMV 13745); E, UADBA-ZCMV 13746. Not to scale.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFF43B1CFF5FFA03FEE6FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262939/files/figure.png	https://doi.org/10.5281/zenodo.15262939	FIGURE 5. Close up photographs of the head of Lygodactylus ornatus from the Mandritsara area in ventral view in life.A, ZSM 79/2023 (ZCMV 13743); B, UADBA-ZCMV 13744; C, ZSM 80/2023 (ZCMV 13745); D, UADBA-ZCMV 13746; E, ZSM 81/2023 (ZCMV 13747). Not to scale.	FIGURE 5. Close up photographs of the head of Lygodactylus ornatus from the Mandritsara area in ventral view in life.A, ZSM 79/2023 (ZCMV 13743); B, UADBA-ZCMV 13744; C, ZSM 80/2023 (ZCMV 13745); D, UADBA-ZCMV 13746; E, ZSM 81/2023 (ZCMV 13747). Not to scale.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFF43B1CFF5FFA03FEE6FDB1.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262941/files/figure.png	https://doi.org/10.5281/zenodo.15262941	FIGURE 6. Maximum Likelihood tree of the Lygodactylus verticillatus group based on a concatenated alignment of DNA sequences (11,241 bp) of five mitochondrial and ten nuclear gene fragments. Inset pictures show representative individuals of the two target lineages, L. ornatus and L. namoroka sp. nov. Values at nodes are bootstrap proportions in percent (1,000 replicates). A sequence of L. miops was used as the outgroup (graphically excluded to better illustrate branch lengths within the L. verticillatus group). Included individuals are listed after the species name; sequences of the two paratypes of L. namoroka were also obtained and deposited in Genbank.	FIGURE 6. Maximum Likelihood tree of the Lygodactylus verticillatus group based on a concatenated alignment of DNA sequences (11,241 bp) of five mitochondrial and ten nuclear gene fragments. Inset pictures show representative individuals of the two target lineages, L. ornatus and L. namoroka sp. nov. Values at nodes are bootstrap proportions in percent (1,000 replicates). A sequence of L. miops was used as the outgroup (graphically excluded to better illustrate branch lengths within the L. verticillatus group). Included individuals are listed after the species name; sequences of the two paratypes of L. namoroka were also obtained and deposited in Genbank.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFFA3B19FF5FF955FB28FEB5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262944/files/figure.png	https://doi.org/10.5281/zenodo.15262944	FIGURE 7. Individuals of Lygodactylus namoroka sp. nov. from Tsingy de Namoroka National Park.A, Holotype ZSM 70/2023 (ZCMV 15820); B, paratype UADBA-ZCMV 15837. Not to scale.	FIGURE 7. Individuals of Lygodactylus namoroka sp. nov. from Tsingy de Namoroka National Park.A, Holotype ZSM 70/2023 (ZCMV 15820); B, paratype UADBA-ZCMV 15837. Not to scale.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFFA3B19FF5FF955FB28FEB5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262946/files/figure.png	https://doi.org/10.5281/zenodo.15262946	FIGURE 8. Holotype specimen ZSM 70/2023 (ZCMV 15820) of Lygodactylus namoroka sp. nov. from Namoroka National Park: A, dorsal view; B, ventral view; C, close up of the ventral view of the throat.	FIGURE 8. Holotype specimen ZSM 70/2023 (ZCMV 15820) of Lygodactylus namoroka sp. nov. from Namoroka National Park: A, dorsal view; B, ventral view; C, close up of the ventral view of the throat.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFFA3B19FF5FF955FB28FEB5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262950/files/figure.png	https://doi.org/10.5281/zenodo.15262950	FIGURE 9. Ventral views of throats of representative specimens in preservative of the L. verticillatus group (not including species of the L. tolampyae complex), illustrating differences in the mental and postmental scale arrangement and in throat color pattern. Specimens shown are ZSM 70/2023 (L. namoroka sp. nov.), ZSM 81/2023 (L. ornatus), ZSM 308/2004 (L. heterurus), ZSM 46/2006 (L. klemmeri), ZSM 190/2004 (L. verticillatus), ZSM 319/2019 (L. blancae), ZSM 584/2001 (L. arnoulti). Scale bars represent 1 mm.	FIGURE 9. Ventral views of throats of representative specimens in preservative of the L. verticillatus group (not including species of the L. tolampyae complex), illustrating differences in the mental and postmental scale arrangement and in throat color pattern. Specimens shown are ZSM 70/2023 (L. namoroka sp. nov.), ZSM 81/2023 (L. ornatus), ZSM 308/2004 (L. heterurus), ZSM 46/2006 (L. klemmeri), ZSM 190/2004 (L. verticillatus), ZSM 319/2019 (L. blancae), ZSM 584/2001 (L. arnoulti). Scale bars represent 1 mm.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFFA3B19FF5FF955FB28FEB5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262954/files/figure.png	https://doi.org/10.5281/zenodo.15262954	FIGURE 10. Schematic representation of the arrangement of throat scales in species of the Lygodactylus verticillatus group, drawn after photos in Fig. 9 and in Vences et al. (2024). M (light green), mental scale; PM (light blue), postmental scales; IL1 (dark green), first infralabial. In all species, the mental scale is semi-divided by two incomplete sutures, and there are three postmental scales. In L. namoroka sp. nov. the two lateral postmentals have a distinct and rather broad contact with the lateral parts of the mental (distal from the sutures) which is missing in all other species. The four species of the L. tolampyae complex are characterized by a broad contact between the first infralabial and the central part of the mental scale. Not to scale.	FIGURE 10. Schematic representation of the arrangement of throat scales in species of the Lygodactylus verticillatus group, drawn after photos in Fig. 9 and in Vences et al. (2024). M (light green), mental scale; PM (light blue), postmental scales; IL1 (dark green), first infralabial. In all species, the mental scale is semi-divided by two incomplete sutures, and there are three postmental scales. In L. namoroka sp. nov. the two lateral postmentals have a distinct and rather broad contact with the lateral parts of the mental (distal from the sutures) which is missing in all other species. The four species of the L. tolampyae complex are characterized by a broad contact between the first infralabial and the central part of the mental scale. Not to scale.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
03DBCE50FFFA3B19FF5FF955FB28FEB5.taxon	http://purl.org/dc/dcmitype/StillImage	image/png	https://zenodo.org/record/15262956/files/figure.png	https://doi.org/10.5281/zenodo.15262956	FIGURE 11. Habitat of Lygodactylus namoroka sp. nov. at Namoroka National Park. A and C, Grand Tsingy; B, Petit Tsingy.	FIGURE 11. Habitat of Lygodactylus namoroka sp. nov. at Namoroka National Park. A and C, Grand Tsingy; B, Petit Tsingy.	2025-04-17	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel		Zenodo	biologists	Rakotoarison, Andolalao;Gippner, Sven;Multzsch, Malte;Miralles, Aurélien;Razafimanafo, Delina;Hasiniaina, Alida;Glaw, Frank;Vences, Miguel			
