taxonID	type	description	language	source
03D94977FFE5FFAC24A8FAEFFE0A8D76.taxon	description	(Fig. 14) urn: lsid: zoobank. org: act: 9 E 284050 - 50 CB- 4 D 22 - B 1 CE- 5718 B 3221202 Tope material: Holotype: ♂ (NWAFU), China: Ruili Co., Yunnan Province, 24 August 2014. Paratype: 1 ♂ (NWAFU), China: Mt Baihualing, Baoshan, Yunnan Province, 30 May 2018. Measurements of topes: (in millimetres; 2 ♂♂): body length, ♂ 42.8 – 43.2; forewing length, ♂ 46.7 – 47.1; forewing width, ♂ 16.5 – 17.0; width of head including eyes, ♂ 9.6 – 9.9; pronotum width (including pronotal collar), ♂ 11.1 – 11.4; and mesonotum width, ♂ 10.4 – 10.9. Etomologo: Te species name is a Latin adjective meaning ‘ short’, signifying the spines of lateral branches of the uncus being comparatively short. Description of male: Head (Fig. 14 A, B, D). Mostly ochraceous. Head including eyes slightly narrower than the base of the mesonotum. Eyes fuscous and ocelli yellow. Ventral part of head brown, with a broad pale yellowish transverse band across head and postclypeus. Postclypeus much swollen. Anteclypeus ochraceous, with dense golden hair. Rostrum ochraceous, with apical part blackish, not extending to the posterior margin of abdominal sternite II. Torax (Fig. 14 A, E). Pronotum longer than head, almost brown; inner area of pronotum with fuscous longitudinal fascia medially and fuzzy patches on each side. Pronotal collar ochraceous, with small spot medially and two pairs of blackish spots on posterolateral area; lateral part of pronotal collar obtusely dentate. Mesonotum fuscous, with the following ochraceous markings: pair of delicate fasciae along submedian sigilla reaching to half of mesonotum; and narrow fasciae along lateral margins of mesonotum. Cruciform elevation ochraceous, posterior angles of cruciform elevation with black markings. Legs (Fig. 14 C). Yellowish ochraceous, with some brown patches. Fore femora swollen and equipped with three blackish brown spines beneath: primary spine oblique to femur; secondary spine broader and erect; and subapical spine short. Wings (Fig. 14 A, B). Hyaline, heavily tinged with pale brown. Forewing with distinct spots on radial, radiomedial, and medial crossveins, obscure fuscous spot present on mediocubital crossvein, and spot on cubitus anterior vein; marginal series of faint and minute spots present on apex of longitudinal veins of apical cells. Abdomen (Fig. 14 A, B, E, F). Cylindrical, much longer than distance from head to cruciform elevation. Abdominal tergites brown, with posterior margins of each tergite narrowly black. Abdominal sternites translucent and ochraceous. Timbal covers semicircular and blackish brown; timbal concealed by timbal cover in dorsal view. Opercula ochraceous, separated, with posterior margins semicircular and extending to posterior margin of sternite II. Genitalia (Fig. 14 G – I). Pygofer elliptical in ventral view, ochraceous, with apex blackish brown. Distal shoulder broadly triangular, distally extended into pointed lobe. Upper lobe of pygofer relatively large and erect, substantially confluent with pygofer margin in ventral view (Fig. 14 G). Basal lobes large and paramedian, nearly straight, and parallel to each other. Uncus trapezoid, with a narrow and short median incision. Lateral branches of uncus protruding from below uncus and each with paired short spines, subequal in length in ventral view. Distribution: China (Yunnan). Diagnostic features: Pomponia brevisa can be distinguished from other species of the P. linearis species complex by the distinctly fuscous spots on radial, radiomedial, and medial crossveins of forewing, obscure fuscous spot on the mediocubital crossvein of forewing, spot on cubitus anterior vein of forewing, and the distinctly short spines of lateral branches of uncus. Tis new species is most similar to P. ponderosa within the P. linearis species complex, but can be distinguished from P. ponderosa by the upper lobe of the pygofer being erect, large, and long, and by the basal lobes of the pygofer being large and paramedian, nearly straight, and parallel to each other.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA727AEFB37FE2A8EC7.taxon	distribution	Distribution: Vietnam and Bangladesh.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFE9FFAE24ADF929FEE889B8.taxon	description	(Figs 11, 12) urn: lsid: zoobank. org: act: 2 F 4 D 3226 - 7 D 3 E- 47 AB-BF 2 E- 01 B 55 C 2 FFB 20 Tope material: Holotype: ♂ (NWAFU), China: Yinjiang Co., Yunnan Province, 14 August 2022, Wenzhe Zhang leg. Paratypes: 10 ♂ (NWAFU), China: Yinjiang Co., Yunnan Province, 14 August 2022, Wenzhe Zhang leg. Measurements of topes: (in millimetres; 15 ♂♂): body length, ♂ 38.0 – 39.2; forewing length, ♂ 41.5 – 42.4; forewing width, ♂ 13.8 – 14.3; width of head including eyes, ♂ 9.7 – 10.2; pronotum width (including pronotal collar), ♂ 12.8 – 13.3; and mesonotum width, ♂ 10.1 – 10.4. Etomologo: Te species name is a Latin adjective meaning ‘ eminent’, signifying the greatly swollen postclypeus. Description of male: Head (Fig. 11 A, B, D). Mostly green afer death. Head including eyes as wide as the base of the mesonotum. Eyes blackish brown and ocelli greenish yellow. A fuscous median longitudinal fascia extending from anterior margin of head to frontoclypeal suture and median blackish brown spot enclosing ocelli. Ventral part of head ochraceous, with a broad pale yellowish transverse band across head and postclypeus. Postclypeus much swollen. Anteclypeus blackish brown, with dense golden hair. Rostrum ochraceous, with apical part blackish brown, almost extending to posterior margin of abdominal sternite II. Torax (Fig. 11 A, E). Pronotum longer than head, almost greenish yellow; inner area of pronotum with fuscous longitudinal fascia medially and with brown irregular patch on each side; pronotal collar green, with small median spot and with two pairs of large brown spots on each lateral angle of pronotal collar. Mesonotum fundamentally blackish brown, with the following greenish markings: paired delicate fasciae along submedian sigilla; paired fasciae along parapsidal suture, reaching to half of mesonotum; paired lobate fasciae along lateral sigilla, reaching to anterior angles of cruciform elevation; and narrow fasciae along lateral margins of mesonotum, reaching to anterior angles of cruciform elevation. Cruciform elevation greenish brown. Legs (Fig. 11 C). Green, with blackish brown patches. Fore femora swollen and equipped with three blackish brown spines beneath: primary spine pointed and oblique to femur; secondary spine broader with acute tip, erect; and subapical spine short and small. Wings (Fig. 11 B). Hyaline. Forewing with fuscous spots on radial, radiomedial, medial, and mediocubital crossveins, small fuscous spots on both ends of cubitus anterior vein, and nearly uniform marginal series of faint and minute spots present on apex of longitudinal veins of apical cells. Abdomen (Fig. 11 B, E, F). Cylindrical balloon, noticeably longer than head and thorax together. Abdominal tergites brown to ochraceous, with posterior margins of each tergite narrowly black. Abdominal sternites translucent and mostly pale brown. Timbal cover semicircular and brown; timbal mostly concealed by timbal cover in dorsal view. Opercula ochraceous, separated, and broader than long, extending slightly beyond posterior margin of sternite II. Genitalia (Fig. 11 G – I). Pygofer elliptical in ventral view, ochraceous, with apex blackish brown. Distal shoulder broadly triangular, distally extended into pointed lobe. Upper lobes large, substantially confluent with pygofer margin in ventral view (Fig. 11 G). Basal lobes small, nearly straight, and parallel to each other. Uncus trapezoid, with a narrow and short median incision. Lateral branches of uncus protruding from below uncus and each with two long and acute spines: lateral spine slightly curved laterally, and medial spine nearly straight and longer than lateral spine in ventral view. Description of female (Fig. 12 A – E): Opercula small, posterior margin extending abdominal sternite II. Ovipositor sheath not protruding beyond apex of segment IX, posterior margin of sternite VII incised at middle. Other characteristics similar to male. Distribution: China (Yunnan). Diagnostic features: Pomponia eminula can be distinguished from other species of the P. linearis species group by the distinctly fuscous spots on radial, radiomedial, medial, and mediocubital crossveins of forewing, two small fuscous spots on both ends of cubitus anterior vein (except P. brevisa) of forewing, nearly uniform marginal series of faint and minute spots on apex of longitudinal veins of apical cells (except P. ramifera and P. urania), and the slightly laterally curved lateral spine of lateral branches of uncus shorter than nearly straight medial spine of lateral branches of uncus. Pomponia eminula can be distinguished from P. brevisa by the smaller body size, the ochraceous markings on the mesonotum, and the small basal lobes. Pomponia eminula can be distinguished from P. ramifera and P. urania by the distinctly fuscous spot on the mediocubital crossvein and by the semicircular timbal cover.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFE7FFAA267BF9B1FE438E2D.taxon	description	(Figs 15, 16) urn: lsid: zoobank. org: act: B 442 E 5 AE- 9239 - 409 A- 9 EAB- B 0654277 E 3 C 3 Tope material: Holotype: ♂ (NWAFU), China: Mt. Limu, Qiongzhong Co., Haikou, Hainan Province, 22 August 2010, 730 m, Mingsheng Yang leg. Paratypes: 1 ♂ (NWAFU), Mt. Jianfenglin, Ledong Co., Hainan Province, 6 August 2009, 325 m, Rui Meng and Manqiang Wang leg.; 1 ♂ (NWAFU), China: Mt. Wuzhi, Hainan Province, 12 August 2010, 738 m, Mingsheng Yang leg.; 4 ♂♂, 1 ♀ (NWAFU), China: Mt. Wuzhi, Hainan Province, 25 August 2020, Jiali Wang leg.; 3 ♂♂, 2 ♀♀ (NWAFU), China: Mt. Jianfenglin, Ledong Co., Hainan Province, 2 September 2020, Jiali Wang leg. Measurements of topes: (in millimetres; 8 ♂♂, 3 ♀♀): body length, ♂ 43.4 – 45.2, ♀ 32.8 – 33.5; forewing length, ♂ 46.4 – 49.2, ♀ 47.1 – 49.5; forewing width, ♂ 16.4 – 16.8, ♀ 16.5 – 16.8; width of head including eyes, ♂ 10.5 – 11.0, ♀ 10.1 – 10.5; pronotum width (including pronotal collar), ♂ 12.3 – 13.2, ♀ 13.1 – 14.1; and mesonotum width, ♂ 10.8 – 11.5, ♀ 10.9 – 11.5. Etomologo: Te species name is derived from the type locality. Description of male: Head (Fig. 15 A, B, D). Mostly greenish yellow afer death. Head including eyes as wide as the base of the mesonotum. Eyes blackish brown and ocelli yellow. Ventral part of head fuscous, with a broad yellow transverse band across head and postclypeus, which extends to anterior part of prothorax. Postclypeus moderately swollen. Anteclypeus green. Rostrum ochraceous, with apical part blackish, extending almost to posterior margin of abdominal sternite II. Torax (Fig. 15 A, E). Pronotum longer than head, almost greenish yellow; inner area of pronotum ochraceous, with brown markings along FIssures and on parts between posterior ends of paramedian and lateral FIssures. Pronotal collar yellowish green, with brown small spot medially and two paired spots on posterolateral area; lateral part of pronotal collar obtusely dentate. Mesonotum fundamentally brown, with the following yellow markings: paired delicate fasciae along submedian sigilla, and paired lobate fasciae along parapsidal suture. Cruciform elevation yellowish green, posterior angles of cruciform elevation with black markings. Legs (Fig. 15 C). Green, with some brown markings. Fore femora swollen and equipped with three blackish brown spines beneath: primary spine pointed and oblique to femur; secondary spine broader and erect; and subapical spine small and with acute tip. Wings (Fig. 15 A, B). Hyaline, slightly tinged with pale brown. Forewing with fuscous spots on radial and radiomedial crossveins; marginal series of faint and minute spots present on apex of longitudinal veins of apical cells. Abdomen (Fig. 15 A, B, E, F). Cylindrical, much longer than distance from head to cruciform elevation. Abdominal tergites brown, tergites VII and VIII darker than others, with posterior margins of each tergite narrowly black. Abdominal sternites translucent, sternites II – VI ochraceous, and sternites VII – VIII blackish ochraceous. Timbal covers semicircular and brown; timbal completely concealed by timbal cover in dorsal view. Opercula yellowish green, separated, with posterior margins semicircular and extending to posterior margin of sternite II. Genitalia (Fig. 15 G – I). Pygofer elliptical in ventral view, yellow, with apex blackish brown. Distal shoulder of pygofer acute and erect, distally extended into pointed lobe. Upper lobes large, substantially confluent with pygofer margin in ventral view (Fig. 15 G). Basal lobes of pygofer relatively long, large, and paramedian, nearly straight and parallel to each other. Uncus trapezoid, with a narrow and short median incision. Lateral branches of uncus protruding from below uncus and each with two spines; spines comparatively short, subequal in length in ventral view. Description of female (Fig. 16 A – E): Wingspan slightly larger. Operculum small, posterior margin just reaching abdominal sternite II. Ovipositor relatively long, narrowly indented, not protruding beyond segment IX, posterior margin of segment 7 incised at middle. Distribution: China (Hainan). Diagnosis features: Tis new species can be distinguished from other species of the P. linearis species complex by the obscure and fuscous spots only on radial and radiomedial crossveins of forewing and by the long lateral branches of uncus, which have comparatively short spines.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFEDFFA72640FB06FDDA8FF1.taxon	materials_examined	Materials examined: 5 ♂♂, 1 ♀ (NWAFU), China: Mt. Emei, Sichuan Province, 20 June 2009, Weixin Huo leg.; 10 ♂♂ (NWAFU), China: Mt. Qingcheng, Sichuan Province, 24 June 2009, Weixin Huo leg.; 10 ♂♂ (NWAFU), China: Donghui, Lancang Co., Yunnan Province, 25 May 2011, Xiao Chen leg.; 10 ♂♂ (NWAFU), China: Mt. Xi, Kunming, Yunnan Province, 17 May 2009, Libin Ma leg.; 10 ♂♂ (NWAFU), China: Xishuangbanna, Yunnan Province, 23 July 2007, Guoxi Xue leg.; 10 ♂♂ (NWAFU), China: Mt. Longtang Co., Zhejiang Province, 15 June 2016, Yunxiang Liu leg.; 25 ♂♂ (NWAFU), China: Mt. Tianmu, Zhejiang Province, 26 June 2016, Yunxiang Liu leg.; 11 ♂♂, 5 ♀♀ (NWAFU), China: Quanzhou, Fujian Province, 14 July 2016, Xu Wang leg.; 5 ♂♂, 3 ♀♀ (NWAFU), China: Dehua Co., Fujian Province, 2 June 2010, Xiangqun Yuan leg.; 5 ♂♂ (NWAFU), China: Changde, Hunan Province, 4 June 2015, Xu Wang leg.; 11 ♂♂ (NWAFU), Enshi, Hubei Province, 10 July 2019, Xiaodong Peng leg.; 9 ♂♂ (NWAFU), Mt. Bamian, Hunan Province, 29 July 2019, Siyue Wang leg.; 3 ♂♂ (NWAFU), China: Mt. Fanjing, Guizhou Province, 28 July 2001; 12 ♂♂ (NWAFU), China: Guilin, Guangxi Province, 18 July 2019, Siyue Wang leg.; 15 ♂♂ (NWAFU), China: Mt. Jianfengling, Hainan Province, 5 May 2011, Mingsheng Yang leg.; 15 ♂♂ (NWAFU), China: Mt. Limu, Hainan Province, 8 May 2011, Mingsheng Yang leg.; 1 ♂ (NWAFU), China: Baling, Taiwan Province, 12 June 1991; 10 ♂♂, 4 ♀♀ (NWAFU), China: Nanling National Park, Ruyuan Co., Guangdong Province, 11 June 2020, Jiali Wang leg.; 6 ♂♂, 2 ♀♀ (NWAFU), China: Mt. Wuzhi, Hainan Province, 25 August 2020, Jiali Wang leg.; 2 ♂♂, 2 ♀♀ (NWAFU), China: Chebaling National Park, Ruyuan Co., Guangdong Province, 23 July 2020, Jiali Wang leg.; 9 ♂♂, 5 ♀♀ (NWAFU), China: Dupangling National Park, Yongzhou Co., Hunan Province, 15 July 2020, Jiali Wang leg.; 10 ♂♂, 4 ♀♀ (NWAFU), China: Huaping Natural Reservation Area, Longsheng Co., Guangxi Province, 15 July 2020, Jiali Wang leg.; 12 ♂♂ (NWAFU), China: Mt. Tiantai, Qionglai Co., Sichuan Province, 20 July 2021, Jiali Wang leg.; 15 ♂♂ (NWAFU), China: Mt. Baihualing, Longyang Co., Yunnan Province, 10 June 2022, Jiali Wang leg.; 12 ♂♂ (NWAFU), China: Rongjiang Co., Guizhou Province, 24 June 2022, Jiali Wang leg.; 3 ♂♂ (NWAFU), Japan: Mt. Omotodake, Ishigaki Island, Ryukyu Islands, 1 July 2003, M. Hayashi leg.; 2 ♂♂ (NWAFU), Japan: Shirahama For. Road, Iriomote Island, Ryukyu Islands, 10 September 2003, M. Hayashi leg.; 2 ♂♂ (NWAFU), China: Taitung Co., Taiwan Province, 22 June 2002, M. Hayashi leg. Distribution: China (Anhui, Chongqin, Fujian, Guizhou, Guangdong, Guangxi, Hainan, Hunan, Hubei, Jiangxi, Sichuan, Taiwan, Yunnan, and Zhejiang); Bhutan, India, Japan, Nepal, Philippines, Tailand, and Vietnam. Remarks: Pomponia linearis has been proved to be different from the type species of the genus, P. picta (Supporting Information, Fig. S 1 A) (Duffels and Hayashi 2006). Pomponia linearis together with its allies probably represents an independent genus, and a phylogenetic assessment of phylogenetic affinities in Pomponia is expected in the future. Chou et al. (1997) identiFIed that the dominant species of Pomponia occurring in China was P. linearis. Duffels and Hayashi (2006) came to the conclusion that specimens from Assam of India and those from Bhutan, Nepal, Tailand, and Vietnam belong to P. linearis. However, Lee (2009) indicated that P. ponderosa was misidentiFIed as P. linearis by Chou et al. (1997), which led to the identity of the dominant species of Pomponia occurring in China remaining problematic. In this paper, a large number of individuals of ‘ P. linearis ’ collected from China are identiFIed to be conspeciFIc to the individuals collected from Vietnam and Ryukyu Islands based on both molecular phylogenetic analyses (Figs 2 B, 3) and analysis of genetic distances (Table 2), which conFIrm that the dominant species of Pomponia occurring in China is P. linearis, and P. oaoeoamana is recognized to be a new synonym of P. linearis. According to investigations of ~ 400 specimens, we revealed that this species exhibits remarkable morphological variation (Figs 5 – 7).	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: China (Guangxi); Vietnam.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Vietnam and Malaysia.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Vietnam, Malaysia, Tailand.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Vietnam and Tailand. Pomponia gemella Boulard, 2005	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Tailand. Pomponia namtokola Boulard, 2005	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Tailand.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Vietnam.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	materials_examined	Materials examined: 8 ♂♂ (NWAFU), China: Donghui, Lancang Co., Yunnan Province, 5 June 2015, Xu Wang leg.; 3 ♂♂ (NWAFU), China: Xishuangbanna, Jinghong, Yunnan Province, 29 May 2013, Mingsheng Yang leg.; 3 ♂♂ (NWAFU), China: Jingdong, Yunnan Province, 2 June 2013, Mingsheng Yang leg.; 2 ♂♂, 3 ♀♀ (NWAFU), China: Jingdong, Yunnan Province, 10 June 2011, Lin Lü leg.; 2 ♂♂ (NWAFU), China: Nanning, Guangxi Province, 11 July 2020; 2 ♂♂ (NWAFU), China: Mt. Huanglian, Yunnan Province, 29 June 2017, Yan Su leg.; 6 ♂♂ (NWAFU), China: Pu’er, Yunnan Province, 20 June 2022, Jiali Wang leg. Distribution: China (Yunnan, Guangxi), Cambodia, Tailand, and Vietnam. Remarks: Hill et al. (2021) investigated the molecular systematics and diversiFIcation of various Asian cicada tribes related to Cicadini, in which P. backanensis (accession numbers: MT 639290, MT 599237, and MT 599082) was included. Te specimen of P. backanensis in the study by Hill et al. (2021) was collected from the same locality as part of the type series (Pham and Yang 2009), which nested in the P. subtilita lineage in all the trees constructed in our present study (Figs 2 B, 4). Pomponia backanensis is very similar to P. subtilita in morphology, including the coloration, opercula, wing venation, wing cell morphology, timbal covers, and genitalia. In addition, the body length of P. subtilita males ranges from 37.3 to 41.5 mm (based on 10 ♂♂), slightly overlapping the body length of P. backanensis (36.8 – 37.5 mm) (Pham and Yang 2009). Pomponia subtilita was FIrstly described from Tailand (Lee 2009). Later, it was recorded from China and Cambodia (Sanborn 2013, Pham et al. 2015). Combined with the intraspeciFIc genetic distances between P. backanensis and P. subtilita (0.005 to 0.034) detected in our present study, we synonymize P. backanensis with P. subtilita. It should be stated that the images in FIg. 2 of the paper by Pham and Yang (2009) should be a male individual of another Pomponia species (with a body length of ≥ 45 mm), which was included erroneously in that plate.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: China (Guangxi); Vietnam. Remarks: Pomponia ponderosa is morphologically similar to P. linearis. An assessment of species boundary between P. ponderosa and P. linearis is expected when materials of P. ponderosa are available for molecular analysis.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Indonesia.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Tailand.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Tailand.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	materials_examined	Materials examined: 5 ♂♂, 3 ♀♀ (NWAFU), China: Mt. Jinzhong, Guilin, Guangxi Province, 9 August 2014; 1 ♂ (NWAFU), China: Mt. Huanglian, Yunnan Province, 29 June 2017, Yan Su leg.; 1 ♂ (NWAFU), China: Hekou Co., Yunnan Province, 16 August 2015; 2 ♂♂ (NWAFU), Vietnam: Cuc Phuong National Park, Ninh Binh Province, 7 July 2019, Cong Wei leg. Distribution: China (Guangxi and Yunnan); Vietnam (Cuc Phuong National Park). Remarks: Pomponia brevialata was recorded only from Vietnam, and it is recorded from China for the FIrst time. Pham et al. (2015) assigned P. brevialata within the P. linearis species complex, but we exclude P. brevialata from the P. linearis species complex owing to the basal lobes of the pygofer of P. brevialata being slender and small.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: Malaysia.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA2264EF8B5FBCA8DEE.taxon	distribution	Distribution: India. Remarks: Te collection locality of Pomponia sp. A [3] (Idukki, Kerala; accession numbers: MT 639321 and MT 599274) in the study by Hill et al. (2021) is the type locality of P. pseudolinearis. Te distribution of the remaining specimens of Pomponia sp. A (except Pomponia sp. A [4] distributed in West Bengal, Jalpaiguri) in the study by Hill et al. (2021) is along the Western Ghats, which suggests that these specimens (except Pomponia sp. A [4]) are individuals of P. pseudolinearis. Examination of the morphology of Pomponia sp. A [1 – 3, 5] (accession numbers: MT 639234, MT 599170, MT 599018, MT 639321, MT 639318, MT 599270, MT 599274, MT 639350, and MT 599307) in the study by Hill et al. (2021) further conFIrms that these specimens are individuals of P. pseudolinearis in this study. Our molecular phylogenetic analysis revealed that these four individuals diverged distinctly from the individual of Pomponia sp. A [4] (i. e. Pomponia sp. 1 in our present study), and their genetic distances are higher than the intraspeciFIc genetic distances of the P. linearis species group. Tis indicates that Pomponia sp. 1 might represent an independent species from P. pseudolinearis. Pomponia curvata sp. n. (Figs 9, 10) urn: lsid: zoobank. org: act: 0593456 A- 61 AA- 48 DB-BACC- ED 5 A 438 C 9 B 99 Tope material: Holotype: ♂ (NWAFU), China: Wanding Co., Ruili, Yunnan Province, 30 June 1977, Cheping Yang leg. Paratypes: 1 ♂ (NWAFU), China: Mt. Xishan, Kunming, Yunnan Province, 17 May 2009, Libin Ma leg.; 1 ♂ (NWAFU), China: Xishuangbanna Tropical Botanical Garden, Jinghong, Yunnan Province, 23 July 2007, Guoxi Xue leg. Measurements of topes: (in millimetres; 3 ♂♂): body length, ♂ 38.1 – 39.9; forewing length, ♂ 43.2 – 44.6; forewing width, ♂ 14.3 – 15.3; width of head including eyes, ♂ 9.1 – 9.7; pronotum width (including pronotal collar), ♂ 11.6 – 12.1; mesonotum width, ♂ 9.2 – 9.9. Etomologo: Te species name is a Latin adjective meaning ‘ curved’, signifying the curved spines of lateral branch of uncus. Description of male: Head (Fig. 9 A, B, D). Mostly ochraceous afer death. Head including eyes as wide as the base of mesonotum. Eyes yellow and ocelli greyish yellow. A fuscous median longitudinal fascia extending from anterior margin of head to frontoclypeal suture and median blackish brown spot enclosing ocelli. Ventral part of head ochraceous with a broad pale yellowish transverse band across head and postclypeus. Postclypeus moderately swollen. Anteclypeus ochraceous, with dense golden hair. Rostrum ochraceous with apical part blackish brown, almost extending to posterior margin of abdominal sternite II. Torax (Fig. 9 A, E). Pronotum longer than head, almost ochraceous; inner area of pronotum ochraceous, with fuzzy fuscous longitudinal fascia medially; pronotal collar ochraceous, with small median spot and with two pairs of large brown spots on each lateral angle of pronotal collar. Mesonotum fundamentally blackish brown, with the following ochraceous markings: paired delicate fasciae along submedian sigilla; paired lobate fasciae along parapsidal suture, reaching to anterior angles of cruciform elevation; narrow fasciae along lateral margins of mesonotum, reaching to anterior angles of cruciform elevation. Cruciform elevation ochraceous, with a fuzzy fuscous spot medially. Legs (Fig. 9 C). Ochraceous with brown patches. Fore femora swollen and equipped with three blackish brown spines beneath: primary spine pointed and slightly oblique to femur; secondary spine broader, erect; and subapical spine very small. Wings (Fig. 9 A, B). Hyaline. Forewing with smudged fuscous spots on radial, radiomedial, and medial crossveins, and marginal series of faint and minute spots present on apex of longitudinal veins of apical cells. Abdomen (Fig. 9 A, B, E, F). Cylindrical balloon, noticeably longer than head and thorax together. Abdominal tergites brown to ochraceous, with posterior margins of each tergite narrowly black, tergites VII and VIII darker than others. Abdominal sternites translucent and mostly pale brown, but sternites VII and VIII darker than others. Timbal cover semicircular and brown; timbal mostly concealed by timbal cover in dorsal view. Opercula ochraceous, broader than long, extending slightly beyond posterior margin of sternite II; opercula overlapped at inner margins. Genitalia (Fig. 9 G – I). Pygofer elliptical in ventral view, ochraceous, with apex blackish brown. Distal shoulder broadly triangular, distally extended into pointed lobe. Upper lobes small, substantially confluent with pygofer margin in ventral view (Fig. 9 G). Basal lobes small, slightly obliquely inward. Uncus trapezoid, with a narrow and short median incision. Lateral branch of uncus protruding from below uncus and each with two long and acute spines, curved laterally, with medial spine slightly shorter than lateral spine in ventral view. Description of female (Fig. 10 A – E): Opercula small, posterior margin extending to abdominal sternite II. Abdominal tergite IX blackish brown in dorsal view and ochraceous in ventral view; ovipositor sheath not protruding beyond apex of segment IX, posterior margin of sternite VII incised at middle. Other characteristics similar to male. Distribution: China (Yunnan). Diagnosis features: Pomponia curvata can be distinguished from other species of the P. linearis species group by the smudged spots on the radial and radiomedial crossveins of the forewing and the faintly smudged spot on the medial crossvein of the forewing, and by the distinctly laterally curved spines on the lateral branches of the uncus.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA7265DFA41FE878D2D.taxon	distribution	Distribution: Vietnam.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFF2FFA62679FCCCFEBE8F81.taxon	diagnosis	Diagnosis: Medium to large size. Head including eyes as wide as or wider than the base of mesonotum. Pronotum longer than head and distinctly shorter than mesonotum including cruciform elevation, lateral margin of pronotal collar narrow, with a small tooth. Forewing and hindwing hyaline, with eight and six apical cells, respectively. Abdomen gradually tapering to apex and usually longer than the distance from head to cruciform elevation in both sexes; timbals mostly concealed by timbal covers in dorsal view; opercula in male short, scale-like, and wider than long. Male abdominal sternites III and IV without tuberclelike projections. Pygofer with distal shoulders developed into pointed lobes; basal lobes of pygofer present; uncus with two lobes and paired lateral branches of uncus protruding below the uncus. Distribution: Bangladesh, Cambodia, China, India, Indonesia, Japan, Nepal, Malaysia, Philippines, Tailand, and Vietnam. Remarks: Although some species of Pomponia were recognized as new genera (Boulard 2005 a, 2008), Pomponia still comprises a very heterogeneous group of species, and the relationships within Pomponia remain unclariFIed. Currently, species of Pomponia could be separated easily and sorted into groups: (i) P. picta and its allies (e. g. Pomponia fugax Boulard, 2013, Pomponia mickwanae Boulard, 2009); (ii) P. linearis species group; and (iii) remaining species, which might be members of other genera (e. g. P. bullata, P. rajah) (for details, see the section on ‘ Te monophyly of Pomponia and phylogeny of Pomponia and allies’ in the Discussion).	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFF2FFA62679FCCCFEBE8F81.taxon	description	2. Male abdomen shorter than head and thorax together ................................................................................................... P. brevicaudata Male abdomen longer than head and thorax together ............................................................................................................................ 3 3. Tergite IV almost as wide as tergite III ....................................................................................................................................................... 4 Tergite IV wider than tergite III ........................................................................................................................................... P. xizangensis 4. Fasciae along parapsidal suture reaching to half of mesonotum ........................................................................................................... 6 Fasciae along parapsidal suture reaching to cruciform elevation .......................................................................................................... 5 5. Body length <44.0 mm ...................................................................................................................................................................... P. latiffi Body length> 50.0 mm .................................................................................................................................................................... P. dolosa 6. Median spines of lateral branch of uncus as long as lateral spines ........................................................................................................ 7 Median spines of lateral branch of uncus longer than lateral spines ....................................................................................... P. linearis 7. Basal lobes of pygofer erect ........................................................................................................................................................................... 8 Basal lobes of pygofer obliquely inward ................................................................................................................................. P. namtokola 8. Distal shoulder erect in ventral view ........................................................................................................................................................... 9 Distal shoulder curved laterally in ventral view ........................................................................................................................ P. brevisa 9. Upper lobes of pygofer obliquely inward ............................................................................................................................... P. ponderosa Upper lobes of pygofer obliquely lateral ........................................................................................................................... P. hainanensis 10. Basal lobes of pygofer paramedian ........................................................................................................................................................... 11 Basal lobes of pygofer placed laterally ..................................................................................................................................................... 15 11. Distal shoulder of pygofer lower than anal tube .................................................................................................................................... 12 Distal shoulder of pygofer higher than anal tube ............................................................................................................... P. minilinearis 12. Median spines of lateral branch of uncus about as long as lateral spines .......................................................................................... 13 Median spines of lateral branch of uncus longer than lateral spines ................................................................................... P. eminula 13. Mediocubital crossvein of forewing with spot ....................................................................................................................................... 14 Mediocubital crossvein of forewing without spot .................................................................................................................. P. curvata 14. Length of forewing ~ 1.1 times the body length .................................................................................................................... P. brevialata Length of forewing ~ 1.3 times the body length ............................................................................................................. P. pseudolinearis 15. Spines of lateral branch of uncus straight ............................................................................................................................................... 16 Spines of lateral branch of uncus curved ..................................................................................................................................... P. gemella 16. Spines of lateral branch of uncus equal in length ..................................................................................................................... P. subtilita Spines of lateral branch of uncus unequal in length ............................................................................................................................. 17 17. Median spines of lateral branch of uncus shorter than lateral spines ................................................................................................ 18 Median spines of lateral branch of uncus longer than lateral spines 20 18. Male operculum somewhat semicircular ................................................................................................................................................ 19 Male operculum obliquely triangular .................................................................................................................................... P. tricornisa 19. Larger body size (~ 42.0 mm) ......................................................................................................................................................... P. piceata Smaller body size (~ 37.5 mm) ............................................................................................................................................... P. daklakensis 20. Male operculum somewhat semicircular ................................................................................................................................................ 21 Male operculum obliquely triangular ..................................................................................................................................... P. pornnapae 21. Mediocubital crossvein of forewing without spot ......................................................................................................... P. langkawiensis Mediocubital crossvein of forewing with spot .............................................................................................................................. P. zakrii Note that P. cinctimanus, P. ramifera and P. urania are not included in the key because no materials / images are available for examination of the genitalia. Te identity of these three species needs to be investigated further.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFE5FFAE266FFCFFFB948FA8.taxon	description	(Fig. 13) urn: lsid: zoobank. org: act: 7552 FD 15 - E 83 E- 4762 - A 75 A- 13 B 8 B 47 CB 71 B Tope material: Holotype: ♂ (NWAFU), China: Mt. Jianshan, Jinhua, Zhejiang Province, 15 June 2009, Lei Zhang leg. Paratypes: 1 ♂ (NWAFU), China: Mt. Jianshan, Jinhua, Zhejiang Province, 15 June 2009, Lei Zhang leg.; 1 ♂ (NWAFU), China: Mt. Wuyi, Zhejiang Province, 20 July 2006. Measurements of topes: (in millimetres; 3 ♂♂): body length, ♂ 33.6 – 38.8; forewing length, ♂ 41.7 – 43.4; forewing width, ♂ 14.9 – 15.8; width of head including eyes, ♂ 9.8 – 10.3; pronotum width (including pronotal collar), ♂ 11.3 – 12.7; and mesonotum width, ♂ 9.9 – 10.5. Etomologo: Te species name is a Latin adjective meaning ‘ triangular’, signifying the triangular operculum. Description of male: Head (Fig. 13 A, B, D). Mostly ochraceous. Head including eyes as wide as the base of mesonotum. Eyes fuscous and ocelli red. Ventral part of head brown, with a broad pale yellowish transverse band across head and postclypeus. Postclypeus less swollen. Anteclypeus yellowish green. Rostrum yellow, with apical part blackish, extending to trochanters of hind legs. Torax (Fig. 13 A, E). Pronotum longer than head, almost brown; inner area of pronotum with fuzzy fuscous longitudinal fascia medially and fuzzy patches on each side. Pronotal collar yellowish green, with a small spot medially and two pairs of blackish spots on posterolateral area. Mesonotum fuscous, with the following ochraceous markings: pair of delicate fasciae along submedian sigilla reaching to half of mesonotum, and irregular markings around the fasciae; and narrow fasciae along lateral margins of mesonotum. Cruciform elevation yellowish green. Legs (Fig. 13 C). Yellowish green, with some brown patches. Fore femora swollen and equipped with three blackish brown spines beneath: primary spine pointed and slightly oblique to femur; secondary spine broader, erect; and subapical spine very small, with acute tip. Wings (Fig. 13 A, B). Hyaline, slightly tinged with pale brown. Forewing with distinct spots on radial, radiomedial, medial, and mediocubital crossveins, and marginal series of faint and minute spots present on apex of longitudinal veins of apical cells. Abdomen (Fig. 13 A, B, E, F). Cylindrical, much longer than distance from head to cruciform elevation. Abdominal tergites brown, with posterior margins of each tergite narrowly black. Abdominal sternites translucent and yellow. Timbal covers semicircular and brown; timbal concealed by timbal cover in dorsal view. Opercula yellow, triangular, separated from each other and broader than long, outer margins narrowly black reaching posterior margin of sternite II. Genitalia (Fig. 13 G – I). Pygofer elliptical in ventral view, ochraceous, with apex blackish brown. Distal shoulder broadly triangular, distally extended into pointed lobe. Upper lobe of pygofer large, long, and erect. Basal lobes minute, placed laterally adjacent to side of pygofer. Uncus trapezoid, with a narrow and short median incision. Lateral branches of uncus protruding from below uncus and each with two narrow, long, and straight spines, medial spine shorter than lateral spine in ventral view. Female: Unknown. Distribution: China (Zhejiang). Diagnosis features: Pomponia tricornisa can be distinguished from other species of the P. linearis species group by the long and straight spines of lateral branches of uncus, the medial spine which is shorter than the lateral spine, and the very small basal lobes of pygofer. Tis new species is most similar to P. subtilita within the P. linearis species group, but can be distinguished from P. subtilita by the triangular opercula and the large and erect upper lobe of the pygofer.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFECFFA72654F96BFE878C73.taxon	distribution	Distribution: Vietnam.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
03D94977FFE1FFA82679FA6BFCA98DA5.taxon	description	(Figs 17, 18) urn: lsid: zoobank. org: act: 92 C 300 D 0 - 0944 - 4284 - B 728 - E 8 B 32 C 574166 Tope material: Holotype: ♂ (NWAFU), China: Motuo Co., Xizang Autonomous Region, 27 July 2022, Qingquan Xue leg. Paratypes: 7 ♂♂, 5 ♀♀ (NWAFU), China: Motuo Co., Xizang Autonomous Region, 27 July 2022, Qingquan Xue leg. Measurements of topes: (in millimetres; 5 ♂♂, 5 ♀♀): body length, ♂ 47.8 – 49.4, ♀ 34.8 – 35.5; forewing length, ♂ 53.4 – 54.2, ♀ 51.1 – 52.5; forewing width, ♂ 17.4 – 17.8, ♀ 16.9 – 17.4; width of head including eyes, ♂ 10.6 – 11.0, ♀ 10.3 – 10.7; pronotum width (including pronotal collar), ♂ 13.8 – 14.2, ♀ 14.1 – 14.5; andmesonotumwidth, ♂ 11.8 – 12.5, ♀ 11.9 – 12.5. Etomologo: Te species name is derived from the type locality. Description of male: Head (Fig. 17 A, B, D). Mostly ochraceous. Head including eyes slightly wider than the base of the mesonotum. Eyes fuscous and ocelli yellow. Ventral part of head brown, with a broad pale yellowish transverse band across head and postclypeus. Postclypeus much swollen. Anteclypeus ochraceous. Rostrum yellow, with apical part blackish, extending to posterior margin of abdominal sternite II. Thorax (Fig. 17 A, E). Pronotum longer than head, almost brown; inner area of pronotum with fuzzy fuscous longitudinal fascia medially and fuzzy patches on each side. Pronotal collar ochraceous, with a small spot medially and two pairs of blackish spots on posterolateral area; lateral part of pronotal collar obtusely dentate. Mesonotum fuscous, with the following ochraceous markings: pair of delicate fasciae along submedian sigilla reaching to half of mesonotum; paired lobate fasciae along parapsidal suture; and narrow fasciae along lateral margins of mesonotum. Cruciform elevation ochraceous, posterior angles of cruciform elevation with black markings. Legs (Fig. 17 C). Yellowish ochraceous, with some brown patches. Fore femora swollen and equipped with three blackish brown spines beneath: primary spine pointed and oblique to femur; secondary spine broader and erect; and subapical spine very small. Wings (Fig. 17 A, B). Hyaline, slightly tinged with pale brown. Forewing with distinct spots on radial, radiomedial, and medial crossveins, and obscure fuscous spot present on the mediocubital crossvein; marginal series of faint and minute spots present on apex of longitudinal veins of apical cells. Abdomen (Fig. 17 A, B, E, F). Cylindrical, much longer than distance from head to cruciform elevation. Abdominal tergites brown, with posterior margins of each tergite narrowly black. Abdominal sternites translucent and yellow. Timbal covers semicircular and brown; timbal concealed by timbal cover in dorsal view. Opercula ochraceous, separated, with posterior margins semicircular and extending to posterior margin of sternite II. Genitalia (Fig. 17 G – I). Pygofer elliptical in ventral view, ochraceous, with apex blackish brown. Distal shoulder broadly triangular, distally extended into pointed lobe. Upper lobe of pygofer relatively large and erect, substantially confluent with pygofer margin in ventral view (Fig. 17 G). Basal lobes large and paramedian, obliquely inward, with apex widely separated. Uncus trapezoid, with a narrow and short median incision. Lateral branches of uncus protruding from below uncus and each with spines: medial spine long and straight; lateral spine very short, lateral spine shorter than medial spine in ventral view. Description of female (Fig. 18 A – E): Wingspan slightly larger. Operculum small, posterior margin just reaching abdominal sternite II. Ovipositor relatively long, narrowly indented, protruding beyond segment IX; posterior margin of segment 7 incised at middle. Distribution: China (Xizang). Diagnostic features: Tis new species can be distinguished from other species of the P. linearis species complex by the large and robust body and by the junction of the radiomedial crossvein of the hindwing and the FIrst median vein of the hindwing being located nearly at the base of the FIrst median vein. Remarks: Pomponia xizangensis was collected from Motuo County, Xizang Autonomous Region, China, and this species might also be distributed in adjacent areas in Bhutan and West Nepal.	en	Wang, Jiali, Hayashi, Masami, Wei, Cong (2025): Phylogeny, biogeography, and diversiFIcation of the cicada Pomponia linearis and its allies (Hemiptera: Cicadidae). Zoological Journal of the Linnean Society 203 (4), DOI: 10.1093/zoolinnean/zlaf019, URL: https://doi.org/10.1093/zoolinnean/zlaf019
