taxonID	type	description	language	source
03D97C79011AFFD6F8EFFFAEFB847EAF.taxon	description	http: // zoobank. org / urn: lsid: zoobank. org: act: BB 8 AA 288 - 640 C- 4 A 66 - A 32 F-B 1 BDEC 480 B 14	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C79011AFFD6F8EFFFAEFB847EAF.taxon	type_taxon	Type genus. Tibionema Solier, 1851.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C79011AFFD6F8EFFFAEFB847EAF.taxon	type_taxon	Type genus. Tibionema Solier, 1851.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C79011AFFD6F8EFFFAEFB847EAF.taxon	type_taxon	Type genus. Tibionema Solier, 1851.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C79011AFFD6F8EFFFAEFB847EAF.taxon	diagnosis	Diagnosis. Tibionemini is proposed in Pityobiinae sensu Motyka et al. (2023 b) as a monogeneric tribe. Currently, the nominotypical Tibionema is the only genus placed in it, and its diagnosis presented below is simultaneously the diagnosis of the tribe. Tibionema has a medium-to-large body, flat pronotum and elytra (Fig. 1 A), distant longitudinal bulges in the pronotum, and acutely prominent posterior angles of the pronotum (Fig. 1 A). The prosternum is narrow, and the prosternal process is long and slender (Fig. 1 B). The lateral edges of the pronotum are complete (Fig. 1 E, F). There is a sharp keel between mesocoxae (Fig. 1 B). Antennae are 11 - segmented and serrate; the scapus is parallelsided, antennomeres 2 and 3 are short and ball-like, and antennomeres 1 – 3 are almost bare and shining (Fig. 1 D). The abdomen has five visible segments, a short intercoxal process, a straight posterior margin of the penultimate segment, and a triangular, apically rounded last visible abdominal segment. Internal abdominal segments consist of long, narrow penultimate, and small ultimate sternites (Fig. 1 H). Tibionema and Pityobius differ in the number of antennomeres (twelve in Pityobius versus eleven in Tibionema); the posterior pronotal angles bent toward the body in Tibionema (also in Hapatesus and Oxynopterus), but not in Pityobius. Tibionema has a prominent intercoxal keel in the mesosternum (Fig. 1 B) that has not been observed in other Pityobiinae. Campyloxenus differs in much longer antennomere 3. The larva and pupa of T. abdominalis were described by Angulo (Angulo, 1970).	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C79011AFFD6F8EFFFAEFB847EAF.taxon	discussion	Remark. Here, Tibionemini trib. nov. is proposed due to the unavailability of the same name published earlier by Motyka et al. (2023 b). As we cannot get a response from the journal and rectify this omission in another way, we have to re-publish the description to make it valid in the sense of the Code (ICZN 2012).	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD6F8EFFECFFE887995.taxon	type_taxon	Type species: Hiekeolycus berendti Winkler, 1987 (original designation).	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD6F8EFFECFFE887995.taxon	diagnosis	Diagnosis. Hiekeolycus resembles related Helcophorus Fairmaire in general appearance. The genera differ in the structure of the elytral humeri. The humeral bulge is built only by costa 3 in Hiekeolycus (Fig. 2 A, C) and by costae 3 and 4 in Helcophorus.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD6F8EFFECFFE887995.taxon	discussion	Justification of the name resurrection. Hiekeolycus Winkler, 1987 has been considered a synonym of Helcophorus Fairmaire, 1881 (Kazantsev 2004, 2013) as these genera are similar, and the elytral humeri of H. berendti and H. ekaterinae were poorly seen (Kazantsev & Perkovsky 2022; Winkler 1987). We compared all Eocene species with extant Helcophorus and noted differently arranged costae. Therefore, we resurrect Hiekeolycus as a separate genus.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD7F8EFF8FEFD387E3B.taxon	description	http: / zoobank. org / urn: lsid: zoobank. org: act: 812 CA 068 - 62 DB- 4 EFC-B 5 AD- 5 B 1 DB 5 E 15 EE 8 (Fig. 2 A – E).	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD7F8EFF8FEFD387E3B.taxon	materials_examined	Material examined. Holotype: ♂, specimen No. AMNH Ba-SY 2, deposited in the American Museum of Natural History, New York, NY, USA. Baltic amber, Eocene.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD7F8EFF8FEFD387E3B.taxon	diagnosis	Diagnosis. H. winkleri sp. nov. differs from contemporary H. berendti in the less conspicuous anterior and more acute posterior pronotal angles and distally widened hind trochanters (Fig. 2 D, E). Furthermore, H. berendti has denser and longer elytral vestiture and fewer elytral cells in each interstice than H. winkleri (Fig. 2 A – E).	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD7F8EFF8FEFD387E3B.taxon	description	Description. Male. Body elongated; dark brown (putatively red in the live specimen; Fig. 2). Head narrower than pronotum. Eyes prominent, interocular dorsal distance 1.4 times eye diameter (Fig. 2 B); terminal palpomeres long, wide distally; antennae reaching over elytral mid-length; antennomere 3 longer than antennomere 2 (Fig. 2 C), combined subequal in length to antennomere 4 (Fig. 2 C). Pronotum 1.6 times wider than long, anteriorly rounded, with widely rounded anterior and sharply prominent posterior angles; median areole rhomboidal; lateral carinae almost straight; noticeable fronto-lateral carina ventrally (Fig. 2 A). Elytra flat, elongated, 3.3 times longer than width at humeri, four primary costae, costa 3 stoutest and basally forming humeral bulge; interstices with transverse cells (Fig. 2 A, B). Femurs widened distally, tibiae with pair of small spurs; frontal and middle trochanters elongate, distally rounded, hind trochanters distally widened (Fig. 2 D); tarsomeres with plantar pads in tarsomeres 1 – 4 (Fig. 2 A, B, D). Abdomen with eight visible ventrites, penultimate ventrite with broad triangular incision (Fig. 2 E). Measurements. Body length 11.2 mm; pronotal length 1.4 mm; pronotal width 2.2 mm.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD7F8EFF8FEFD387E3B.taxon	etymology	Etymology. H. winkleri sp. nov. is named in honor of our late colleague Josef R. Winkler (1930 – 1993), who described the first erotine taxa from Baltic amber in the late 1980 s.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790119FFD7F8EFF8FEFD387E3B.taxon	discussion	Locality and horizon. Baltic amber, Eocene.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
03D97C790118FFD7F8EFF805FEFA7A03.taxon	discussion	Remark. We transfer Helcophorus ekaterinae to Hiekeolycus based on the updated genus diagnosis. The new combination with Hiekeolycus proposed in 2023 is invalid due to incomplete data in the work’s registration in ZooBank.	en	Bocak, Ladislav, Motyka, Michal, Kusy, Dominik, Arias-Bohart, Elizabeth T., Bybee, Seth M., Kazantsev, Sergey V., Yamamoto, Shuhei (2025): Validation of earlier proposed taxa Tibionemini new tribe and Hiekeolycus winkleri new species. Zootaxa 5609 (2): 293-297, DOI: 10.11646/zootaxa.5609.2.10, URL: https://doi.org/10.11646/zootaxa.5609.2.10
