identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03E5B46DCE770772FC7AFB45FEA9FD0A.text	03E5B46DCE770772FC7AFB45FEA9FD0A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Aethalops aequalis G. M. Allen 1938	<div><p>Aethalops aequalis G. M. Allen 1938</p><p>Borneo Fruit Bat (Cecadu Bukit)</p><p>Specimens examined (5).— Mt. Penrisen: TTU 108348 ( GenBank EU521595), TTU 108352, TTU 108357, TTU 108409, TTU 108381 .</p><p>Type locality.— Malaysia (Borneo), Sabah, Mount Kinabalu .</p><p>Malaysian distribution.—Recorded from highlands in Sabah and Sarawak. Specimens were collected from Mt. Penrisen in this study.</p><p>Remarks.— Aethalops aequalis is commonly found at higher elevations in hill and mountain forests (Payne et al. 1985). This species was caught using mist nets across small streams. Compared to other fruit bats, A. aequalis is fragile and requires careful handling and quick release if it is not to be prepared as a voucher specimen. This species was previously recognized as Aethalops alecto (Payne et al. 1985) . Studies (Kitchener et al. 1993) suggest that only Peninsular Malaysia populations, as well as Sumatra, Java, Bali, and Lombok populations, should be recognized as alecto (Pygmy Fruit Bat) whereas Borneo populations are referable to aequalis . Although morphological characters differentiate aequalis from alecto, allozymic variation between them is low (see Kitchener et al. 1993). Cytochrome- b gene sequences within our sample from Borneo averaged 0.2%. Pending genetic data from Peninsular Malaysia populations as well as specimens from Sabah, we follow Kitchener et al. (1993) for geographic boundaries of A. alecto and A. aequalis .</p></div>	https://treatment.plazi.org/id/03E5B46DCE770772FC7AFB45FEA9FD0A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE760772FED6FC9BFE15F89B.text	03E5B46DCE760772FED6FC9BFE15F89B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Balionycteris maculata (Thomas 1893)	<div><p>Balionycteris maculata (Thomas 1893)</p><p>Spotted-winged Fruit Bat (Cecadu Sayap Bertitik)</p><p>Specimens examined (7).— Bako NP: TTU 108265 (GenBank EU521596), TTU 108266, TTU 108299; Kubah NP: TTU 108312, TTU 108317, TTU 108321; Mt. Penrisen: TTU 108364 .</p><p>Type locality.— Malaysia (Borneo), Sarawak .</p><p>Malaysian distribution.—Recorded from Borneo.</p><p>Remarks.— Balionycteris maculata is the smallest fruit bat in Borneo (FA: 40-45 mm; weight: 10-15 g). This species was caught using mist nets near rock cliffs in mangrove forests, over ponds, and in open areas near hill edges. A diagnostic feature of this bat is the obvious white to pinkish spots on its wings. These frugivorous bats are commonly found in lowland dipterocarp forest and occasionally at higher elevations. Two subspecies, B. maculata seimundi and B. maculata maculata, have been recognized in Peninsular Malaysia and Borneo, respectively (Kloss 1921), a taxonomic arrangement followed by many authors (e.g., Payne et al. 1985; Hodgkison and Kunz 2006; Francis 2008). We found a genetic distance of 12% in the cyt- b gene sequences between populations in Borneo and Peninsular Malaysia. Following application of the 5% species level separation proposed by Baker and Bradley (2006), and the morphological variation between Kalimantan (Indonesian Borneo) and Sumatra (Indonesia) populations (Maryanto 2003), we recognize these populations as separate species.</p></div>	https://treatment.plazi.org/id/03E5B46DCE760772FED6FC9BFE15F89B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE760772FC78FE90FC16FC54.text	03E5B46DCE760772FC78FE90FC16FC54.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Balionycteris seimundi (Kloss 1921)	<div><p>Balionycteris seimundi (Kloss 1921)</p><p>Malayan Spotted-winged Fruit Bat (Cecadu Sayap Bertitik Malaya)</p><p>Specimens examined (2).— Krau: TK 152011 (GenBank EU521597), TTU 108175 .</p><p>Type locality.— Malaysia (Peninsular Malaysia), Pahang, Gunung Tahan .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states).</p><p>Remarks.— Balionycteris seimundi represents the smallest fruit bat of Peninsular Malaysia (FA: 39-43 mm; weight: 13-14.1 g). We propose that the common name for this species be the Malayan Spotted-winged Fruit Bat in English and Cecadu Sayap Bertitik Malaya in Malay. See B. maculata species account for details.</p></div>	https://treatment.plazi.org/id/03E5B46DCE760772FC78FE90FC16FC54	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE760772FC72FC57FC09F892.text	03E5B46DCE760772FC72FC57FC09F892.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cynopterus brachyotis Muller 1838	<div><p>Cynopterus brachyotis Müller 1838</p><p>Lesser Short-nosed Fruit Bat (Cecadu Pisang)</p><p>Specimens examined (18).— Monggis: TTU 108245 (GenBank EU521598), TTU 108246, TK 152094, TTU 108247, TTU 108248, TTU 108257, TTU 108258, TTU 108259; Bako NP: TTU 108280; Kubah NP: TTU 108320, TTU 108323, TTU 108324, TTU 108330, TTU 108338; Mt. Penrisen:TTU 108411, TTU 108401, TTU 108402, TTU 108382 .</p><p>Type locality.— Dewei River in Borneo, also known as Tewei River in Naga Cave .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo from sea level to 1600 m.</p><p>Remarks.— Cynopterus brachyotis was netted mainly in primary forest, secondary forest, and agricultural areas. We identified two forms (large FA: 60-68 mm; small FA: 54-59 mm) of C. brachyotis (reported in Abdullah 2003; Campbell et al. 2004), but these size groups were not congruent with mitochondrial DNA data. The level of cyt- b genetic distances within Peninsular Malaysia (10.3%) and Borneo (9.5%) suggested two independent lineages; however, both genetic lineages include individuals that overlap in forearm sizes.</p></div>	https://treatment.plazi.org/id/03E5B46DCE760772FC72FC57FC09F892	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE79077DFEF6FE91FEF1FAA0.text	03E5B46DCE79077DFEF6FE91FEF1FAA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Cynopterus horsfieldii Gray 1843	<div><p>Cynopterus horsfieldii Gray 1843</p><p>Horsfield’s Fruit Bat (Cecadu Pisang Besar)</p><p>Specimens examined (9).— Krau: TK 152002 (GenBank EU521599), TK 152003, TTU 108170, TTU 108212, TTU 108214, TTU 108215, TK 152058, TTU 108223, TTU 108224 .</p><p>Type locality.— Indonesia, Java .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states) and Borneo from sea level to 1400 m.</p><p>Remarks.— This species was caught using mist nets at Krau across forest trails. In Borneo, C. horsfieldii was captured at high elevations (900 m) in primary forest (Jayaraj et al. 2005; 2006). Forearm length (68- 76 mm) is a good character to separate this species from C. brachyotis in the field (Payne et al. 1985). Unlike C. brachyotis, there is low cyt- b genetic distance (2%) observed within and between Peninsular Malaysian (samples from this study) and Bornean (samples from UNIMAS museum collections) individuals. Habitats where C. horsfieldii occurs in Borneo should be given high conservation priority, as this species is rare or at least not commonly reported when compared to Peninsular Malaysia samples. Two of four adult female specimens were pregnant: TK 152002 (3 August 2006) and TK 152058 (6 August 2006); and two were carrying young: TK 152003 (6 August 2006) and TTU 108223 (7 August 2006).</p></div>	https://treatment.plazi.org/id/03E5B46DCE79077DFEF6FE91FEF1FAA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE79077DFEFBFAE2FA95FCB6.text	03E5B46DCE79077DFEFBFAE2FA95FCB6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eonycteris major K. Andersen 1910	<div><p>Eonycteris major K. Andersen 1910</p><p>Greater Dawn Bat (Cecadu Gua Besar)</p><p>S p e c i m e n s e x a m i n e d (4).— Kubah NP: TTU108327 (GenBank EU521600); Mt. Penrisen: TTU 108371, TTU 108372, TTU 108410 .</p><p>Type locality.— Malaysia (Borneo), Sarawak, Mount Dulit.</p><p>Malaysian distribution.— Eonycteris major is endemic to Borneo.</p><p>Remarks.—The genus Eonycteris is distinguished from all other fruit bats by the lack of a claw on the second digit of the wing. We caught Eonycteris major at Kubah NP in a net placed in front of a banana tree, and in Mt. Penrisen this species was caught at the edge of a steep cliff where bats were flying near stands of banana trees. This species is the largest nectar-feeding bat in Malaysia, with a long muzzle that is slender and curved. Eonycteris major also has an enlarged gland near the anus. Forearm length distinguishes E. major (71-80 mm) from the smaller E. spelaea (62-70 mm; Payne et al. 1985). Both of these species roost in limestone caves (SAMD database). Low cyt- b genetic distance (0.1%) was observed among Bornean individuals. A specimen, TTU 108410 (18 August 2006; Mt. Penrisen), was carrying earwigs (Scizochelisoches sp.; Fig. 2) on its dorsum near the base of the tail. This is the first record for earwigs on this bat species. Specimens of these earwigs are housed in the NSRL.</p></div>	https://treatment.plazi.org/id/03E5B46DCE79077DFEFBFAE2FA95FCB6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE79077DFC6BFCF5FAA6F8D9.text	03E5B46DCE79077DFC6BFCF5FAA6F8D9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Eonycteris spelaea (Dobson 1871)	<div><p>Eonycteris spelaea (Dobson 1871)</p><p>Lesser Dawn Bat (Cecadu Gua Kecil)</p><p>Specimens examined (7).—Krau: TK 152009 (GenBank EU521601), TTU 108211; Monggis: TTU 108240, TK 152091; Mt. Penrisen: TTU 108373, TTU 108378, TTU 108387.</p><p>Type locality.— Burma, Tenasserim, Moulmein, Farm Caves .</p><p>Malaysian distribution.—Recorded from the lowlands of Peninsular Malaysia (all states) and Borneo.</p><p>Remarks.—We netted this species near banana trees. Eonycteris spelaea can be distinguished from their larger counterpart E. major by forearm length and shorter muzzle size (see E. major species account). Like E. major, this species also has an enlarged external gland lateral to the anus. Eonycteris spelaea is distributed throughout most of Southeast Asia and often found in colonies of thousands in caves within agricultural areas (McClure et al. 1967; Heaney et al. 1989). Genetic distance value of &lt;0.8% was observed within Peninsular Malaysia and Borneo individuals, whereas genetic distance value of 1% was observed between Peninsular Malaysia and Borneo individuals. A pregnant female, TK 152085 (8 August 2006), was recorded with an embryo length of 31 mm.</p></div>	https://treatment.plazi.org/id/03E5B46DCE79077DFC6BFCF5FAA6F8D9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE78077FFF2CFAB2FD75FDA2.text	03E5B46DCE78077FFF2CFAB2FD75FDA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macroglossus minimus (E. Geoffroy 1810)	<div><p>Macroglossus minimus (E. Geoffroy 1810)</p><p>Dagger-toothed Long-nosed Fruit Bat (Cecadu Madu Bakau)</p><p>Specimens examined (11).—Monggis: TTU 108225 (GenBank EU521602), TTU 108226, TK 152070, TK 152071, TTU 108227, TTU 108231, TTU 108232; Mt. Penrisen:TTU 108350, TTU 108366, TTU 108374, TTU 108376.</p><p>Type locality.— Indonesia, Java .</p><p>Malaysian distribution.—Recorded from both Peninsular Malaysia (all states) and Borneo.</p><p>Remarks.— Macroglossus minimus and M. sobrinus were netted near banana trees and in most cases they were caught alongside C. brachyotis . Macroglossus minimus can be morphologically distinguished from M. sobrinus by the absence of a median groove in the upper lip, and its smaller FA length (FA &lt;42 mm vs. FA&gt; 42 mm; Payne et al. 1985; Francis 2008). Recent literature and SAMD database indicate M. minimus occurs throughout Southeast Asia, whereas M. sobrinus is restricted to Peninsular Malaysia, Sumatera, and Java. Low cyt- b genetic distance (1%) was observed within Bornean populations of M. minimus . A specimen, TTU 108366 (male), from Mt. Penrisen, Borneo, was identified as M. minimus based on the absence of a median groove in the upper lip and FA = 42 mm. This individual is genetically closer to M. sobrinus rather than M. minimus . Genetic distance between M. minimus (Borneo) and M. sobrinus (Peninsular Malaysia) averaged 2% and was the lowest for any sister species relationship within our data for the family Pteropodidae from Malaysia. Although they may represent two species, a genetic distance value of 2% is typical for conspecific bat populations (Bradley and Baker 2001; Baker and Bradley 2006) and we question the recognition of these geographically isolated populations as distinct species. Three of our five female specimens were pregnant; TK 152070 (7 August 2006), TTU 108231 (7 August 2006), and TTU 108350 (17 August 2006).</p></div>	https://treatment.plazi.org/id/03E5B46DCE78077FFF2CFAB2FD75FDA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7B077FFF25FDE3FE2FFB5B.text	03E5B46DCE7B077FFF25FDE3FE2FFB5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Macroglossus sobrinus K. Andersen 1911	<div><p>Macroglossus sobrinus K. Andersen 1911</p><p>Greater Long-nosed Fruit Bat (Cecadu Madu Bukit)</p><p>Specimens examined (4).—Krau: TTU 108174 (GenBank EU521603), TK 152047, TTU 108206, TTU 108207.</p><p>Type locality.— Malaysia (Peninsular Malaysia), Perak, Gunung Igari ( Mount Igari; 610 m) .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states).</p><p>Remarks.— Macroglosus sobrinus can be distinguished from their smaller counterpart M. minimus by larger forearm length, body size, and presence of median groove in the upper lip (see M. minimus species account for details).</p></div>	https://treatment.plazi.org/id/03E5B46DCE7B077FFF25FDE3FE2FFB5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7B077FFEDDFB6BFBA5FDE6.text	03E5B46DCE7B077FFEDDFB6BFBA5FDE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaerops ecaudatus (Temminck 1837)	<div><p>Megaerops ecaudatus (Temminck 1837)</p><p>Temminck’s Tailless Fruit Bat (Cecadu Tiada Berekor)</p><p>Specimens examined (4).—Mt. Penrisen: TK 152245 (GenBank EU521604), TTU 108349, TTU 108412, TTU 108413.</p><p>Type locality.— Indonesia, West Sumatra, Padang .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states) and Borneo.</p><p>Remarks.— Megaerops ecaudatus can be netted from lowland as well as mountain forests (Hodgkison 2004). The external morphology of this species is simlar to Cynopterus and Penthetor but with an absence of a tail in Megaerops . Other diagnostic characters include tubular nostrils and absence of white edged ears. Megaerops ecaudatus has low cyt- b genetic distance within our Borneo samples (&lt;2%). Comparison of DNA sequences with others from GenBank showed high genetic distances (7% and 11%) between Peninsular Malaysian (EF105533) and Bornean (EF105536; Mount Murud) cyt- b sequences, suggesting the presence of two species.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7B077FFEDDFB6BFBA5FDE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7B077FFC65FDA4FACAF9CB.text	03E5B46DCE7B077FFC65FDA4FACAF9CB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Penthetor lucasi (Dobson 1880)	<div><p>Penthetor lucasi (Dobson 1880)</p><p>Lucas’s Short-nosed Fruit Bat (Cecadu Hitam Pudar)</p><p>Specimens examined (10).—Bako NP: TTU 108284 (GenBank EU521605), TTU 108287; Kubah NP: TTU 108313, TTU 108318, TTU 108319, TTU 108322, TTU 108329, TTU 108346; Mt. Penrisen: TTU 108363, TTU 108377.</p><p>Type locality.— Malaysia (North Borneo), Sarawak .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.— Penthetor lucasi was netted near rock crevices (Bako NP and Mt. Penrisen) and ponds (Kubah NP). We verified the occurrence of this species at an elevation (857 m) higher than the previously reported upper limit of 600 m (Payne et al. 1985; Jayaraj et al. 2006). Within our sample there appears to be two genetically distinct clusters with about 5% genetic divergence, indicating separate lineages. Sequences in GenBank (EF105541 - EF105544) referred as Penthetor lucasi from Peninsular Malaysia and Borneo do not cluster with our sequences (25-27% genetic divergence), suggesting misidentification as they are closer to our Macroglossus sequences (2-4% genetic divergence). One of 10 adult female specimens, TTU 108313 (14 August 2006), was post lactating.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7B077FFC65FDA4FACAF9CB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7B077EFC71F9F9FD65FC52.text	03E5B46DCE7B077EFC71F9F9FD65FC52.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinolophus affinis Horsfield 1823	<div><p>Rhinolophus affinis Horsfield 1823</p><p>Intermediate Horseshoe Bat (Kelawar Ladam Hutan)</p><p>Specimens examined (4).—Mt. Penrisen: TTU 108353 (GenBank EU521607), TTU 108360, TTU 108361, TTU 108362.</p><p>Type locality.— Indonesia, Java.</p><p>Malaysian distribution.— Recorded from Peninsular Malaysia (all states) and Borneo .</p><p>Remarks.— Megaphyllus species group (Csorba et al. 2003). Rhinolophus affinis is distinguished from other similar species ( R. ferrumequinum, R. rouxi, and R. sinicus) by the broadly rounded connecting process, as well as the tall and triangular noseleaf with a straight-sided lancet (Csorba et al. 2003; Francis 2008). All of our specimens were caught using harp traps along a forest trail. Coloration for this cave dwelling species ranges from bright orange to brown. Bright orange facial color on the rostrum was thought to be a diagnostic character for this species but after examination of several other specimens with similar noseleaf characters and measurements, we concluded that the orange pelage represented intraspecific variation. Approximately 4% in cyt- b genetic distance was observed between Bornean individuals from this study and a single individual from Thailand (TK 2116).</p></div>	https://treatment.plazi.org/id/03E5B46DCE7B077EFC71F9F9FD65FC52	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7A077EFEC1FC53FAAAFE7B.text	03E5B46DCE7A077EFEC1FC53FAAAFE7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinolophus borneensis Peters 1861	<div><p>Rhinolophus borneensis Peters 1861</p><p>Bornean Horseshoe Bat (Local name not available)</p><p>Specimens examined (5).—Monggis: TTU 108228 (GenBank EU521608); Kubah NP: TTU 108308; Mt. Penrisen: TTU 108385, TTU 108388, TTU 108393.</p><p>Type locality.— Malaysia (Borneo), Sabah, Labuan Island.</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (Perlis, Kampung Juara, and Tioman Island) and Borneo.</p><p>Remarks.— Megaphyllus species group (Csorba et al. 2003). Rhinolophus borneensis is distinguished from other similar species ( R. stheno, R. celebensis, and R. malayanus) by its larger forearm size, narrower anterior swellings of upper skull, and the median septa of its lancet being wider than the middle pocket (Csorba et al. 2003; Francis 2008). In Kubah NP, we netted these bats near a small frog pond and also found them roosting inside a culvert. Orange and brown color forms were present within our specimens. Rhinolophus borneensis can be distinguished easily in Borneo as they do not occur in sympatry with species of similar external morphology such as R. stheno or R. malayanus . Despite a wide distribution, R. borneensis is locally rare in Peninsular Malaysia when compared to Borneo (SAMD). Low cyt- b genetic distance (0.3%) was observed within Bornean populations.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7A077EFEC1FC53FAAAFE7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7A077EFC7FFE4BFB02FB9F.text	03E5B46DCE7A077EFC7FFE4BFB02FB9F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinolophus luctus Temminck 1834	<div><p>Rhinolophus luctus Temminck 1834</p><p>Woolly Horseshoe Bat (Kelawar Ladam Terbesar)</p><p>Specimens examined (6).—Kubah NP: TTU 108342 (GenBank EU521609), TTU 108343, TTU 108345; Mt. Penrisen: TTU 108407, TTU 108408, TTU 108395.</p><p>Type locality.— Indonesia, Java, Tapos .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states) and Borneo. Our record is the second record for Mt. Penrisen after Medway (1977).</p><p>Remarks.— Trifoliatus species group (Csorba et al. 2003). In Kubah NP, this species was found roosting inside a culvert, alongside individuals of R. sedulus; whereas in Mt. Penrisen they were caught within a forest along a cliff edge. Rhinolophus luctus is the largest species of horseshoe bat in Malaysia with forearm size of 63-67 mm (Francis 2008).</p></div>	https://treatment.plazi.org/id/03E5B46DCE7A077EFC7FFE4BFB02FB9F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7A0779FCABFB2FFD4FFE25.text	03E5B46DCE7A0779FCABFB2FFD4FFE25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinolophus philippinensis Waterhouse 1843	<div><p>Rhinolophus philippinensis Waterhouse 1843</p><p>Large-eared Horseshoe Bat (Local name not available)</p><p>Specimens examined (2).—Mt. Penrisen: TTU 108359 (GenBank EU521610), TTU 108380.</p><p>Type locality.— Philippines, Luzon .</p><p>Malaysian distribution.—Recorded from Borneo (Sabah: Gomantong Cave, Madai Caves, Sapulut, and Tapadong Cave; Sarawak: Bintulu, Lobang, Long Lama, Niah Caves, and Puteh Caves). Our record is new for Mt. Penrisen.</p><p>Remarks.— Philippinensis species group (Csorba et al. 2003). Rhinolophus philippinensis was netted near a pond in Mt. Penrisen. Scarcity of records in Borneo suggests this species may be rare. Rhinolophus philippinensis is distinguished from other similar species by a FA length of 48-53 mm and by a peculiar noseleaf structure (i.e., wide internarial region and a horseshoe shape). There are morphological, genetic, and echolocation call variations within this species that are associated with different islands in Indonesia (Kingston and Rossiter 2004). One specimen, TTU 108380 (17 August 2006), was post-lactating.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7A0779FCABFB2FFD4FFE25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7D0779FED4FE69FDC4FA80.text	03E5B46DCE7D0779FED4FE69FDC4FA80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinolophus robinsoni Andersen 1918	<div><p>Rhinolophus robinsoni Andersen 1918</p><p>Peninsular Horseshoe Bat (Kelawar Ladam Semenanjung)</p><p>Specimen examined (1).— Krau: TTU 108185 ( GenBank EU521611) .</p><p>Type locality.— Thailand, Surat Thani, Bandon, Kaho Nawng.</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia.</p><p>Remarks.— Megaphyllus species group (Csorba et al. 2003). Rhinolophus robinsoni was caught using a harp trap across a walking trail. The noseleaf of R. robinsoni lacks lateral lappets and has a rounded connecting process. A diagnostic step-wise constriction at the mid length of sella can be observed (Kingston 2006). Csorba et al. (2003) classified robinsoni as a subspecies of R. megaphyllus because it had intermediate features with respect to other Asian mainland species. We recognize R. robinsoni as a valid species following Simmons (2005) and Francis (2008) because R. robinsoni has a restricted geographic distribution and does not overlap with other R. megaphyllus subspecies recognized by Csorba et al. (2003).</p></div>	https://treatment.plazi.org/id/03E5B46DCE7D0779FED4FE69FDC4FA80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7D0779FED2FA02FB2BFE07.text	03E5B46DCE7D0779FED2FA02FB2BFE07.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinolophus sedulus K. Andersen 1905	<div><p>Rhinolophus sedulus K. Andersen 1905</p><p>Lesser Woolly Horseshoe Bat (Kelawar Ladam Bulu Halus)</p><p>Specimens examined (5).—Kubah NP: TTU 108325 (GenBank EU521612), TTU 108341, TTU 108344; Mt. Penrisen: TTU 108394, TTU 108396.</p><p>Type locality.— Malaysia (Borneo), Sarawak .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states) and Borneo.</p><p>Remarks.— Trifoliatus species group (Csorba et al. 2003). Rhinolophus sedulus were found roosting with two R. luctus in a culvert at Kubah NP and were caught using a hand net. Due to scarcity of specimens, little is known about this species. Rhinolophus sedulus has darkly colored trilobated sella and the smallest forearm length (40-44 mm) compared to other species in the trifoliatus species group.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7D0779FED2FA02FB2BFE07	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7D0779FC4CFD87FADCFA80.text	03E5B46DCE7D0779FC4CFD87FADCFA80.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinolophus stheno K. Andersen 1905	<div><p>Rhinolophus stheno K. Andersen 1905</p><p>Lesser Brown Horseshoe Bat (Kelawar Ladam Bukit)</p><p>Specimens examined (2).— Krau: TTU 108171 (GenBank EU521613), TK 152014.</p><p>Type locality.— Malaysia (Peninsular Malaysia), Selangor.</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states).</p><p>Remarks.— Megaphyllus species group (Csorba et al. 2003). Rhinolophus stheno was caught using harp traps in Krau within lowland dipterocarp forests. This species has a unique, straight-sided lancet. The median septum is much wider than the middle pockets and the rounded connecting process does not originate from the tip of the sella. Two different subspecies have been recognized within this species, R. s. stheno (in Java, Sumatra, Peninsular Malaysia, and Thailand south of the Isthmus of Kra) and R. s. microglobosus (in Lao PDR, Vietnam, and Thailand north of the Isthmus of Kra; Csorba et al. 2003). Both of the specimens recorded in this study have the same cyt- b gene haplotype.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7D0779FC4CFD87FADCFA80	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7D0778FC55FA02FD19FDE6.text	03E5B46DCE7D0778FC55FA02FD19FDE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhinolophus trifoliatus Temminck 1834	<div><p>Rhinolophus trifoliatus Temminck 1834</p><p>Trefoil Horseshoe Bat (Kelawar Ladam Muka Kuning)</p><p>Specimens examined (9).— Krau: TTU 108173 (GenBank EU521614), TK 152045, TTU 108205; Monggis: TTU 108238, TK 152098, TTU 108250; Bako NP: TTU 108273, TTU 108275; Kubah NP: TTU 108315.</p><p>Type locality.— Indonesia, West Java, Bantam .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.— Trifoliatus species group (Csorba et al. 2003). This species was mainly caught using harp traps across trails. Rhinolophus trifoliatus can be distinguished easily from other members of the family as it has a distinct yellow noseleaf color. A genetic distance of 0.7% in cyt- b gene sequences was observed between Peninsular Malaysia and Borneo samples.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7D0778FC55FA02FD19FDE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7C0778FEFCFDC2FDA2F8DE.text	03E5B46DCE7C0778FEFCFDC2FDA2F8DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros ater Templeton 1848	<div><p>Hipposideros ater Templeton 1848</p><p>Dusky Leaf-nosed Bat (Local name not available)</p><p>Specimens examined (2).— Mt. Penrisen: TTU 108379 (GenBank EU521615), TTU 108391.</p><p>Type locality.— Sri Lanka, Western Province of Colombo.</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (no specific localities) and Borneo (Sabah: Gomantong, Kinabalu, Madai, Segarong, and Sukau; Sarawak: Bako NP, Bau, Jambusan Cave, Kubah NP, Mt. Murud, and Niah NP). Our record is new for Mt. Penrisen.</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). This bat was caught in harp traps set across walking trails beside a stream. Hipposideros ater has a pale pinkish noseleaf with no lateral leaflets. Important characters used for identification are forearm size and the internarial septum which is slightly swollen at the base and very narrow in the middle (Hill 1963; Hill and Francis 1984). There is morphological overlap (forearm, ear, and tail length) between H. ater, H. bicolor, H. cineraceus, and H. dyacorum . A specimen, TTU 108391 (19 August 2006), from Mt. Penrisen had a facial gland behind the posterior noseleaf (similar to the gland in H. galeritus). Low cyt- b genetic distance was observed between Borneo samples (&lt;1%). However, a GenBank sequence of H. ater (DQ054807) from Australia has a genetic distance value&gt;5% with respect to our Borneo specimen. The significance of this genetic distance value is limited as this is the only available cyt- b gene sequence of H. ater for comparison. In light of current findings, it is probable that H. ater in Borneo might represent a different taxon as proposed by Suyanto and Struebig (2007).</p></div>	https://treatment.plazi.org/id/03E5B46DCE7C0778FEFCFDC2FDA2F8DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7C077BFC4FFE91FD4FFE7A.text	03E5B46DCE7C077BFC4FFE91FD4FFE7A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros bicolor (Temminck 1834)	<div><p>Hipposideros bicolor (Temminck 1834)</p><p>Bicolored Leaf-nosed Bat (Kelawar Ladam Bulat Biasa)</p><p>Specimens examined (20).— Krau: TTU 108417, TK 152015, TTU 108178, TTU 108186, TTU 108189, TTU 108190, TTU 108191, TTU 108192, TTU 108196, TTU 108198, TTU 108199, TTU 108201, TTU 108209, TTU 108222; Bako NP: TTU 108267, TTU 108289, TTU 108294, TTU 108296, TTU 108297, TTU 108301 (GenBank EU521616).</p><p>Type locality.— Indonesia, Java, Anjer coast .</p><p>Malaysian distribution.—Recorded throughout Southeast Asia including Peninsular Malaysia (all states), and Borneo (Sabah: Gomantong, and Sukau; Sarawak: Bako NP, Fairy Cave, and Jambusan Cave).</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). This species was caught in harp traps in small flyways. Hipposideros bicolor has a simple noseleaf, lacking leaflets, and a straight internarial septum (Hill 1963; Hill et al. 1986). More than 20 individuals per night were captured in Krau and released after identification, representing the highest total number of specimens of a single species captured at any site. This species was classified into two different phonic types, 131 kHz and 142 kHz, in Peninsular Malaysia (Kingston et al. 2001). A third phonic type of 136 kHz has been recorded in Sarawak (Les Hall, pers. comm.). Although several morphological features have been described to differentiate the ‘131’ and ‘142’ types, some overlap exists among these features and the frequency of echolocation calls seems to be the best discriminator (Kingston et al. 2001). Variation in echolocation call frequency is reflected in cyt- b sequences with a genetic distance of 7% between the two phonic types in Peninsular Malaysian populations (Kingston et al. 2001). When these two types were compared to Borneo individuals, genetic distances of 5% (to the 131 type) and 11% (to the 142 type) were observed. The significance of this variation remains to be defined. The only available subspecies name in Borneo is H. b. bicolor that is similar to those from Java (type locality), whereas in Peninsular Malaysia it is H. b. atrox (Hill et al. 1986). The two phonic types that are found in Peninsular Malaysia occur sympatrically, suggesting there is at least one unrecognized species within currently designated H. bicolor from Malaysia. Specimen TTU 108296 (13 August 2006) was pregnant.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7C077BFC4FFE91FD4FFE7A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7F077BFEC7FE4BFEC6F950.text	03E5B46DCE7F077BFEC7FE4BFEC6F950.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros cervinus (Gould 1854)	<div><p>Hipposideros cervinus (Gould 1854)</p><p>Fawn-colored Leaf-nosed Bat (Kelawar Ladam Bulat Gua)</p><p>Specimens examined (28).—Krau: TTU 108172 (GenBank EU521617), TK 152007, TTU 108193, TTU 108194, TTU 108195, TTU 108197, TTU 108208, TTU 108220, TTU 108221; Monggis: TTU 108233; Bako NP: TTU 108268, TTU 108269, TTU 108270, TTU 108271, TTU 108288, TTU 108290, TTU 108291, TTU 108292, TTU 108300; Kubah NP: TTU 108316, TTU 108328, TTU 108331, TTU 108332; Mt. Penrisen: TTU 108365, TTU 108367, TTU 108414, TTU 108389, TTU 108392.</p><p>Type locality.— Australia, Queensland, Cape York and Albany Island.</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). Hipposideros cervinus is known to forage on small insects in the forest understory and was collected at all study sites using harp traps. Variation in color of dorsal fur was observed among individuals. In the field, H. cervinus is easily distinguished from most other Hipposiderids, with the exception of H. galeritus . Hipposideros galeritus and H. cervinus are very similar morphologically, but can usually be distinguished from each other by the longer tail length (30-43 mm) and broader intermediate noseleaf compared to the posterior noseleaf (Hill 1963; Hill 1983; Payne et al. 1985) in H. galeritus . Despite differences in color forms there were no substantial morphological differences or cyt- b genetic distance values (&lt;1%) within and between individuals of H. cervinus from Peninsular Malaysia and Borneo.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7F077BFEC7FE4BFEC6F950	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7F077BFC7FFE90FAADF9EB.text	03E5B46DCE7F077BFC7FFE90FAADF9EB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros cineraceus Blyth 1853	<div><p>Hipposideros cineraceus Blyth 1853</p><p>Ashy Leaf-nosed Bat (Kelawar Ladam Bulat Terkecil)</p><p>Specimens examined (1).—Krau: TTU 108176 ( GenBank EU521618) .</p><p>Type locality.— Pakistan, Punjab, Salt Range, near Pind Dadan Khan .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all the states) and Borneo (Sabah: Banturong, Kinabalu, Madai, Marudu Bay, Segarong, and Upper Sungai Kuamut; Sarawak: Bako NP, and Jambusan Cave).</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). In Krau and Mt. Penrisen, this species was caught in a harp trap along with H. bicolor . Hipposideros cineraceus has a pinkish noseleaf without lateral leaflets. Bright pinkish pigments can be observed on rostrum and noseleaf. Hipposideros cineraceus is difficult to distinguish using morphology alone as it closely resembles H. ater and H. dyacorum (Hill 1963; Philips 1967; Hill and Francis 1984). Characteristics that differentiate H. cineraceus from H. bicolor are a thickened internarial septum in the middle of the noseleaf that forms a slightly raised bump, and a smaller body size (Francis 2008). Hipposideros cineraceus is also distinguished from H. doriae by the presence of three vertical septa (Francis 2008). Cytochrome- b genetic distances of 4.3% and 8.7% were observed between our specimen from Krau (Peninsular Malaysia) to the sequence of specimens from Thailand (TK 21261; specimen in NSRL) and Krau (DQ054809; sequence in GenBank), respectively. These data are congruent with Francis (2008) in that there are at least two potential species within the H. cineraceus complex.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7F077BFC7FFE90FAADF9EB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7F077AFC62F9BBFEC9FCB4.text	03E5B46DCE7F077AFC62F9BBFEC9FCB4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros coxi Shelford 1901	<div><p>Hipposideros coxi Shelford 1901</p><p>Cox’s Leaf-nosed Bat (Local name not available)</p><p>Specimens examined (2).—Bako NP: TTU 108272 (GenBank EU521619); Mt. Penrisen: TTU 108358.</p><p>Type locality.— Malaysia (Borneo), Sarawak, Mt .</p><p>Penrisen (1,280 m).</p><p>Malaysian distribution.—Recorded from four localities in Borneo (Sarawak: Bako NP, Bungoh Cave, Jambusan Cave and Mt. Penrisen). This study is the third published record of this species in Sarawak since Payne et al. (1985) and Jub et al. (2003).</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). Hipposideros coxi was caught in a harp trap near rock crevices in Bako NP and in an open area near a clump of trees at Mt. Penrisen. This species has distinctly dark wings and ears covered with hair. A unique feature in this species is the enlarged noseleaf that covers the muzzle with two lateral leaflets (Payne et al. 1985).</p></div>	https://treatment.plazi.org/id/03E5B46DCE7F077AFC62F9BBFEC9FCB4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7E077AFEF2FCF9FE3CF8BC.text	03E5B46DCE7E077AFEF2FCF9FE3CF8BC.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros doriae (Peters 1871)	<div><p>Hipposideros doriae (Peters 1871)</p><p>Bornean Leaf-nosed Bat (Kelawar Ladam Bulat Lawas)</p><p>Specimens examined (2).—Mt. Penrisen: TK 152239 (GenBank FJ460489), TTU 108390.</p><p>Type locality.— Malaysia (Borneo), Sarawak .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (Pahang: Gunung Benom, Krau, Tasik Bera; Perak: Maxwell Hill; and Selangor: Hulu Gombak) and Borneo (Sabah: Gunung Emas-Crocker Range; and Sarawak: Lawas). Our record is new for Mt. Penrisen and is the second record for Sarawak. The first record was described as H. sabanus Thomas 1898, from Lawas, but today sabanus is recognized as a junior synonym of H. doriae Peters 1871 .</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). Hipposideros doriae was caught in a harp trap near a small stream and in a net placed across a small pond. This species is distinguished from other similar species by a posterior noseleaf that lacks vertical septa (Benda 2000; Fig. 3a), a slightly concave horseshoe that is swollen in the middle of an internarial septum, and nostrils covered with small lappets. Gray or dark brown pigments can be observed on rostrum and noseleaf. Both Bornean specimens have the same cyt- b gene haplotype.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7E077AFEF2FCF9FE3CF8BC	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7E077AFC40FE90FB29FB5A.text	03E5B46DCE7E077AFC40FE90FB29FB5A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros dyacorum Thomas 1902	<div><p>Hipposideros dyacorum Thomas 1902</p><p>Dayak Leaf-nosed Bat (Local name not available)</p><p>Specimens examined (3).—Kubah NP: TTU 108340 (GenBank EU521620); Mt. Penrisen: TTU 108355, TTU 108384.</p><p>Type locality.— Malaysia (Borneo), Sarawak, Baram, Mt. Mulu.</p><p>Malaysian distribution.— Recorded from Peninsular Malaysia (Perlis: Kaki Bukit; Pahang: Krau; and restricted areas near Peninsular Thailand) and Borneo (Sabah: Mt. Mulu, Tawau Hill; Sarawak: Bako NP, Bukit Kana, Fairy Cave, Jambusan cave, Kubah NP, Lambir NP, Mt. Hose, and Similajau NP). The specimens from Mt. Penrisen represent a new geographic record.</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). All individuals were collected using harp traps set across small forest trails. Hipposideros dyacorum is distinguished from similar species H. ater, H. cineraceus, and H. bicolor by their internarial septum, which is expanded in the middle (Philips 1967; Payne et al. 1985). Despite the overlap in size with other similar species noted above, this species has the shortest tail (19-24 mm; Payne et al. 1985).</p></div>	https://treatment.plazi.org/id/03E5B46DCE7E077AFC40FE90FB29FB5A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE7E0764FC72FB6BFD92FD4E.text	03E5B46DCE7E0764FC72FB6BFD92FD4E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros galeritus Cantor 1846	<div><p>Hipposideros galeritus Cantor 1846</p><p>Cantor’s Leaf-nosed Bat (Kelawar Ladam Bulat Gua)</p><p>Specimens examined (9).—Bako NP: TTU 108279 (GenBank EU521621), TTU 108283, TTU 108293, TTU 108302, TTU 108303; Mt. Penrisen:TTU 108354, TTU 108356, TTU 108375, TTU 108416.</p><p>Type locality.— Malaysia (Peninsular Malaysia), Penang Island.</p><p>Malaysia distribution.—Recorded from Peninsular Malaysia (Kedah: Pulau Singa, Ulu Muda; Negeri Sembilan: Broga; Pahang: Krau, Taman Negara; Perak: Maxwell Hill; Pulau Pinang: Balik Pulau; and Selangor: Bukit Kutu, Hulu Gombak, Hulu Langat) and throughout Borneo. Our specimens constitute a new geographic record for Mt. Penrisen .</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). Hipposideros galeritus was caught in harp traps along with H. cervinus . Field characters that distinguish H. galeritus from H. cervinus include tail length (30–43 mm in H. galeritus; 21-28 mm in H. cervinus) and broader median than posterior noseleaf; whereas broader posterior noseleaf exists in H. cervinus (Hill 1963) . A male H. galeritus (TTU 108375; 18 August 2006) from Mt. Penrisen had a facial gland behind the posterior noseleaf (Fig. 4). When pressure was applied to the side of this gland, hair-like structures with green secretions emerged.</p></div>	https://treatment.plazi.org/id/03E5B46DCE7E0764FC72FB6BFD92FD4E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE600764FED6FD5CFBDFFC91.text	03E5B46DCE600764FED6FD5CFBDFFC91.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros larvatus (Horsfield 1823)	<div><p>Hipposideros larvatus (Horsfield 1823)</p><p>Intermediate Leaf-nosed Bat (Kelawar Ladam Bulat Besar)</p><p>Specimens examined (4).—Bako NP: TTU 108274 (GenBank EU521622), TTU 108276, TTU 108281, TTU 108282.</p><p>Type locality.— Indonesia, Java .</p><p>Malaysia distribution.—Recorded from Peninsular Malaysia (all states) and Borneo (Sarawak: Bako NP, Fairy Cave, Jambusan, Kubah NP, Kuching, Lubok Simpon, Mt. Pueh, and Tanjung Datu NP).</p><p>Remarks.— Larvatus species group (Corbet and Hill 1992). Hipposideros larvatus was caught in harp traps placed across trails and rock crevices. This is the only species collected in our study that has three lateral leaflets (Payne et al. 1985). Taxonomic reviews from morphological and genetic datasets indicate that more than one genetic phylogroup occurs within this species (Kitchener and Maryanto 1993; Thabah et al. 2006). A genetic distance of 3.5% in the cyt- b gene was observed between a specimen from China (DQ888672) and our specimen from Borneo.</p></div>	https://treatment.plazi.org/id/03E5B46DCE600764FED6FD5CFBDFFC91	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE630767FF3FFE90FD8BFC11.text	03E5B46DCE630767FF3FFE90FD8BFC11.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hipposideros ridleyi Robinson and Kloss 1911	<div><p>Hipposideros ridleyi Robinson and Kloss 1911</p><p>Ridley’s Leaf-nosed Bat (Kelawar Ladam Bulat Singapura)</p><p>Specimens examined (3).—Kubah NP: TK 152188 (GenBank EU521623), TK 152189, TK 152191.</p><p>Type locality.— Singapore, Botanic Gardens .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo</p><p>Remarks.— Bicolor species group (Corbet and Hill 1992). Hipposideros ridleyi was caught inside a drainage tunnel in Kubah NP using hand nets. This species has a large noseleaf that covers the muzzle without lateral leaflets, and has a disc shaped structure on the nostril (Payne et al. 1985). Comparison with a GenBank sequence (DQ054811) indicated a low cyt- b genetic distance (&lt;2%) between Bornean and Peninsular Malaysian populations.</p></div>	https://treatment.plazi.org/id/03E5B46DCE630767FF3FFE90FD8BFC11	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE630767FEC2FBB3FACCFCD4.text	03E5B46DCE630767FEC2FBB3FACCFCD4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Megaderma spasma (Linnaeus 1758)	<div><p>Megaderma spasma (Linnaeus 1758)</p><p>Lesser False Vampire Bat (Kelawar Telinga Lebar)</p><p>Specimens examined (2).—Bako NP: TTU 108285 (GenBank EU521606); Kubah NP: TTU 108347.</p><p>Type locality.— Indonesia, Molucca Isls, Ternate .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.— Megaderma spasma was collected with harp traps across trails in the forest. Unlike Nycteris, Megaderma has long ears that are connected at the base (Payne et al. 1985), a long noseleaf with convex sides, and no tail. A comparison to a GenBank sequence from Java, Indonesia (AY057942), were about 0.4 to 2% cyt- b genetic distance, suggesting individuals we captured in Borneo probably represent M. spasma whereas others specimens compared here need to be reevaluated.</p><p>There are two species in this genus recognized within Malaysia: M. lyra and M. spasma . Morphologically, M. lyra (FA: 60-71 mm) differs in noseleaf shape and narial emargination of skull compared to M. spasma (FA: 52-63 mm) (Corbet and Hill 1992). Genetically, we have compared the cyt- b sequences of specimens identified as M. spasma from Thailand (TK 21322 and TK 21323) to our specimens from Borneo and found a genetic distance of 10-12% separates the two. Additionally, when we compared our specimens to individuals identified as M. lyra in GenBank (DQ888678: China and DQ680822: India), a genetic distance of 20-21% separated lyra from spasma . Specimens from Thailand or those in GenBank may actually represent M. lyra, but this hypothesis remains to be tested, as the voucher specimens were unavailable for examination.</p></div>	https://treatment.plazi.org/id/03E5B46DCE630767FEC2FBB3FACCFCD4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE630767FCA2FC12FAC1F933.text	03E5B46DCE630767FCA2FC12FAC1F933.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Saccolaimus saccolaimus (Temminck 1838)	<div><p>Saccolaimus saccolaimus (Temminck 1838)</p><p>Naked-rumped Pouched Bat (Kelawar Dada Putih)</p><p>Specimens examined (2).—Bako NP: TTU 108286 (GenBank EU521626), TTU 108295.</p><p>Type locality.— Indonesia, Java .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states) and Borneo (Sabah: Keningau, Kota Kinabalu, and Tenom; Sarawak: Bako NP, Kampung Tanah Putih, Kuching, Lundu, Samunsam, and Taman Sukma).</p><p>Remarks.—This species was collected using mist nets in front of a dead mangrove tree where the bats were found roosting. Individuals of Saccolaimus saccolaimus usually fly above the canopy and roost in tree branches or rock crevices. Diagnostic characters distinguishing this species include: absent to poorly developed metacarpal pouch, glandular pouch under the chin, and narrow whitish (almost translucent) wings (Francis 2008). Both specimens collected in this study have the same cyt- b gene sequence haplotype.</p></div>	https://treatment.plazi.org/id/03E5B46DCE630767FCA2FC12FAC1F933	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE620766FC5BFDA4FBD6FA0E.text	03E5B46DCE620766FC5BFDA4FBD6FA0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Arielulus cuprosus (Hill and Francis 1984)	<div><p>Arielulus cuprosus (Hill and Francis 1984)</p><p>Coppery Sprite (Local name not available)</p><p>Specimens examined (2).—Mt. Penrisen: TTU 108383 (GenBank EU521631), TTU 108397.</p><p>Type locality.— Malaysia (Borneo), Sabah, Sepilok (05°52'N, 117°56'E) .</p><p>Malaysian distribution.—Recorded from Sabah, Sepilok. A new geographic record was added for Mt. Penrisen and for the state of Sarawak.</p><p>Remarks.—Tribe Nycticeiini (Hoofer and Van Den Bussche 2003) . Arielulus cuprosus was caught using harp traps across a walking trail at Kubah NP and in mist nets along the edge of a steep cliff at Mt. Penrisen. This species has long, and thick, upper fur with black bases and red orange (coppery) tips (Fig. 3b). Most of the published literature and museum databases have classified A. cuprosus in the genus Pipistrellus or Eptesicus . Four specimens referred as “ Pipistrellus cuprosus ” from Kubah NP (four individuals) and Similajau NP (one individual) are in the UNIMAS museum database. Although not examined by us, these are likely to be Arielulus cuprosus and therefore we have included Kubah NP and Similajau NP as part of the distribution of A. cuprosus . This is a new distribution record for Sarawak.</p></div>	https://treatment.plazi.org/id/03E5B46DCE620766FC5BFDA4FBD6FA0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE620766FF00FECFFED1FAE4.text	03E5B46DCE620766FF00FECFFED1FAE4.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Emballonura alecto (Eydoux and Gervais 1836)	<div><p>Emballonura alecto (Eydoux and Gervais 1836)</p><p>Small Asian Sheath-tailed Bat (Kelawar Teng-Teng Ceteng Kecil)</p><p>Specimens examined (4).—Kubah NP: TTU 108335 (GenBank EU521625), TTU 108336, TTU 108337; Mt. Penrisen: TTU 108405.</p><p>Type locality.— Philippines, Luzon, Manila .</p><p>Malaysian distribution.— This species is known from three localities (Sabah: Poring, and Tawau Hill; Sarawak: Bako NP). Our specimen from Mt. Penrisen is a new geographic record.</p><p>Remarks.— Emballonura alecto was caught using hand nets in a culvert at Kubah NP and with a harp trap near the forest edge at Mt. Penrisen. Specimens are uniformly dark brown to black with long grayish fur. There is morphological overlap between E. alecto and E. monticola, however a short gap between the first and second upper premolars is present in E. alecto and absent in E. monticola (Corbet and Hill 1992) . There is less than 1% genetic divergence within our E. alecto samples from Borneo. Comparison of our E. alecto cyt- b gene sequences from Borneo to E. monticola from Thailand (EF584233) resulted in a genetic distance of 9.2%. Specimen TTU 108405 (19 August 2006) was post lactating.</p></div>	https://treatment.plazi.org/id/03E5B46DCE620766FF00FECFFED1FAE4	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE620766FEC6FAC5FAD3FE25.text	03E5B46DCE620766FEC6FAC5FAD3FE25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Nycteris tragata (K. Andersen 1912)	<div><p>Nycteris tragata (K. Andersen 1912)</p><p>Malayan Slit-faced Bat (Kelawar Muka Lekok)</p><p>Specimen examined (1).—Krau: TTU 108180 (GenBank EU521624).</p><p>Type locality.— Malaysia (Borneo), Sarawak, Bidi caves.</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states) and Borneo.</p><p>Remarks.— Javanica species group (Van Cakenberghe and De Vree 1930). A single individual was collected using harp traps near a walking trail. This is the only species from this family occurring in Malaysia. Nycteris tragata has a long tail and ears that are separated from each other at the base (distinguishing characters between Megadermatidae and Nycteridae); and a slit is present in the middle of the rostrum. Our Peninsular Malaysian specimen was separated by a genetic distance of 2.3% when compared to Borneo sequences obtained from UNIMAS collections.</p></div>	https://treatment.plazi.org/id/03E5B46DCE620766FEC6FAC5FAD3FE25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE620761FC7EF99DFD05FD4F.text	03E5B46DCE620761FC7EF99DFD05FD4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hesperoptenus tomesi Thomas 1905	<div><p>Hesperoptenus tomesi Thomas 1905</p><p>Large False Serotine (Kelawar Petang Palsu)</p><p>Specimen examined (1).—Mt. Penrisen: TTU 108400 (GenBank EU521633).</p><p>Type locality.— Malaysia, Malacca .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (Malacca: Asahan; Selangor: Cameron Highlands, Genting Highlands, Mt. Brinchang; and Perak: Maxwell Hill) and Borneo (Sabah: Sandakan Bay, and Tabin). A new geographic record was added for Mt. Penrisen and for the state of Sarawak.</p><p>Remarks.—Tribe Nycticeiini (Hoofer and Van Den Bussche 2003) . Hesperoptenus tomesi was caught along the edge of a steep cliff using mist nets. This specimen was identified by forearm size of 50-53 mm, a rounded head, and sloping forehead (Payne et al. 1985). Other external characters for this genus include rounded, short ears and a slightly hatchet-shaped tragus. The dorsal and ventral sides of our specimen are uniformly dark blackish brown (Fig. 3c).</p></div>	https://treatment.plazi.org/id/03E5B46DCE620761FC7EF99DFD05FD4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE650761FEFAFD5FFD4EF9E9.text	03E5B46DCE650761FEFAFD5FFD4EF9E9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Glischropus tylopus (Dobson 1875)	<div><p>Glischropus tylopus (Dobson 1875)</p><p>Common Thick-thumbed Bat (Kelawar Tapak Tangan Putih)</p><p>Specimens examined (14).—Monggis: TTU 108229 (GenBank EU521632), TTU 108239, TTU 108243, TK 152089, TTU 108244, TTU 108253, TTU 108255, TTU 108256, TTU 108260, TTU 108264; Kubah NP: TTU 108311; Mt. Penrisen: TTU 108368, TTU 108370, TTU 108398.</p><p>Type locality.— Malaysia (Borneo), Sabah .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.—Tribe Pipistrellini (Hoofer and Van Den Bussche 2003) . This species was caught mainly in harp traps set near bamboo trees. Their unique character is the unpigmented white to pink pad on the thumb and sole of foot that may facilitate their movement inside broken bamboo trees. Specimens have dark brown upperparts and pale buffy brown underparts. Low cyt- b genetic distance (&lt;1%) was observed between G. tylopus from different localities in Borneo.</p></div>	https://treatment.plazi.org/id/03E5B46DCE650761FEFAFD5FFD4EF9E9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE650761FED6F9BCFC49FC76.text	03E5B46DCE650761FED6F9BCFC49FC76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Pipistrellus stenopterus (Dobson 1875)	<div><p>Pipistrellus stenopterus (Dobson 1875)</p><p>Narrow-winged Pipistrelle (Kelawar Malam Kecil)</p><p>Specimens examined (2).—Monggis: TTU 108236 (GenBank EU521634); Mt. Penrisen: TTU 108399.</p><p>Type locality.— Malaysia (Borneo), Sarawak .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (Johore: Endau Rompin; Kedah: Pulau Singa; Negeri Sembilan: Serting; Pahang: Krau, Lakum, Sungai Relau, Taman Negara; Perak: Grik; and Selangor: Bukit Kutu, Bukit Lagong, Hulu Gombak) and Borneo (Sabah: Kota Kinabalu, Mt. Trus Madi, Poring, and Sandakan; Sarawak: Kelabit uplands, and Kuching).</p><p>Remarks.—Tribe Pipistrellini (Hoofer and Van Den Bussche 2003) . The specimen from Monggis was captured using a hand held mist net while it was hawking insects near a lamp in front of the Monggis station headquarters. The specimen from Mt. Penrisen was collected in a mist net along the edge of a steep cliff. These bats have brown to orange upper and underparts.</p></div>	https://treatment.plazi.org/id/03E5B46DCE650761FED6F9BCFC49FC76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE650761FC78FC34FACEF8DD.text	03E5B46DCE650761FC78FC34FACEF8DD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tylonycteris robustula Thomas 1915	<div><p>Tylonycteris robustula Thomas 1915</p><p>Greater Bamboo Bat (Kelawar Buluh Besar)</p><p>Specimens examined (3).—Monggis: TTU 108237; Kubah NP: TTU 108309; Mt. Penrisen: TTU 108386 (GenBank EU521635).</p><p>Type locality.— Malaysia (Borneo), Upper Sarawak .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.—Tribe Vespertilionini (Hoofer and Van Den Bussche 2003) . This species was caught mainly in harp traps near bamboo trees. The unique character of this species is the large, dark brown, flattened, disc-like pad on the thumb and sole of foot (different from those in G. tylopus; see above). Along with a flattened head and body, these pads might facilitate their movement inside hollow bamboo stems. They have dark brown upperparts and slightly paler underparts. Tylonycteris robustula is morphologically similar to T. pachypus which has a smaller greatest skull length of 10.4-10.6 mm compared to 12.3-12.7 mm in T. robustula . Low cyt- b genetic distance (&lt;1%) was observed between T. robustula from different localities in Borneo.</p></div>	https://treatment.plazi.org/id/03E5B46DCE650761FC78FC34FACEF8DD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE640760FEF4FECFFED0FA82.text	03E5B46DCE640760FEF4FECFFED0FA82.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myotis adversus (Horsfield 1824)	<div><p>Myotis adversus (Horsfield 1824)</p><p>Large-footed Myotis (Kelawar Kaki Panjang)</p><p>Specimens examined (3).—Bako NP: TTU 108277 (GenBank EU521628), TTU 108278, TTU 108307.</p><p>Type locality.— Indonesia, Java .</p><p>Malaysian distribution.—Records from Peninsular Malaysia and other parts of the Asian mainland remain uncertain (Francis 2008). In Sabah, this species is known only from Sandakan Bay (Payne et al. 1985). A new geographic record was added for Bako NP and for the state of Sarawak.</p><p>Remarks.—This species was captured in a harp trap and a mist net across a small stream between swamp forest and the beach. As the common name implies, M. adversus has a large foot with a wing membrane inserted at the ankle (Hill 1983; Kitchener et al. 1995; Fig. 3d). Myotis adversus also has a unique translucent wing membrane. A high genetic distance of 15.2% was documented when the Sarawak specimens were compared with a GenBank cyt- b gene sequence (AB106587; see Kawai et al. 2003) from Taiwan. Based on geographic distribution, our specimen would represent M. adversus carimatae and that from Taiwan should correspond to M. adversus taiwanensis (see Kitchener et al. 1995). However, the level of genetic variation suggests that these should be elevated to species status.</p></div>	https://treatment.plazi.org/id/03E5B46DCE640760FEF4FECFFED0FA82	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE640760FE9EFA00FBA7FDA3.text	03E5B46DCE640760FE9EFA00FBA7FDA3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myotis muricola (Gray 1846)	<div><p>Myotis muricola (Gray 1846)</p><p>Nepalese Whiskered Myotis (Kelawar Daun Pisang)</p><p>Specimens examined (5).— Kubah NP: TTU 108310 (GenBank EU521629), TTU 108314; Mt. Penrisen: TTU 108369, TTU 108403, TTU 108404.</p><p>Type locality.— Nepal .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.—This species was collected over small ponds in Kubah NP and in Mt. Penrisen using mist nets. The upperparts are brown to gray with dark bases and the underparts are broadly buffy white with pale gray tips. The wing membrane is attached to the toe (Hill 1983). In comparison to our specimens, high cyt- b genetic distances, from 6.8% (AY665144, locality unknown) to 18.2% (AJ841957 from Laos) were observed, indicating more than one species may be present under this name (see Hendrichsen et al. 2001; Francis et al. 2008).</p></div>	https://treatment.plazi.org/id/03E5B46DCE640760FE9EFA00FBA7FDA3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE640760FC0EFDE0FABDFAE5.text	03E5B46DCE640760FC0EFDE0FABDFAE5.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myotis ridleyi Thomas 1898	<div><p>Myotis ridleyi Thomas 1898</p><p>Ridley’s Myotis (Kelawar Kecil)</p><p>Specimens examined (8).—Krau: TTU 108179 (GenBank EU521630), TTU 108184, TTU 108187, TTU 108188, TTU 108200, TTU 108202, TTU 108203, TTU 108204.</p><p>Type locality.— Malaysia, Selangor, Kepong .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.— Myotis ridleyi was found roosting in small groups. All the specimens were captured using hand nets inside a culvert near Krau headquarters. Myotis ridleyi has small feet with the wing membrane attached to the side of the foot (Medway 1978). This species can be distinguished from similar species using a forearm size of 27-32 mm (Francis 2008).</p></div>	https://treatment.plazi.org/id/03E5B46DCE640760FC0EFDE0FABDFAE5	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE640763FC51FAC7FDCDFDA2.text	03E5B46DCE640763FC51FAC7FDCDFDA2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Murina rozendaali Hill and Francis 1984	<div><p>Murina rozendaali Hill and Francis 1984</p><p>Gilded Tube-nosed Bat (Kelawar Hidung Bulu Merah)</p><p>Specimens examined (2).—Krau: TTU 108216; Monggis: TTU 108241 (GenBank EU521627).</p><p>Type locality.— Malaysia (Borneo), Sabah, Gomantong .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (Pasoh, Negeri Sembilan; Krau, Pahang) and Borneo (Sabah: Gomantong Cave, Poring, and Tepadong; Sarawak: Bukit Kana, Kubah NP, and Mt. Murud).</p><p>Remarks.— Murina rozendaali, caught using harp traps, is difficult to distinguish from M. aenea and M. suilla (Francis 1997) . Our specimens have upperparts with dark bases and golden tips whereas underparts are white to the bases with a buffy tinge. We conclude that our specimen represents M. rozendaali as it was collected in similar habitat (disturbed lowlands over a small stream) and falls within described external measurements (Payne et al. 1985).</p></div>	https://treatment.plazi.org/id/03E5B46DCE640763FC51FAC7FDCDFDA2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE670763FEC0FD01FDFAFAA1.text	03E5B46DCE670763FEC0FD01FDFAFAA1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kerivoula hardwickii Horsfield 1824	<div><p>Kerivoula hardwickii Horsfield 1824</p><p>Hardwicke’s Woolly Bat (Kelawar Hutan)</p><p>Specimens examined (4).—Monggis: TTU 108235 (GenBank EU188768), TTU 108249, TTU 108263; Bako NP: TTU 108298.</p><p>Type locality.— Indonesia, Java .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.— Kerivoula hardwickii was collected using harp traps near streams and fragmented forest. Historically, this species has been confused with K. pellucida (Tomes 1858), however the wings of K. hardwickii are not as translucent as those of K. pellucida . Kerivoula hardwickii also has a less globular braincase, with a wider posterior palate extension. Our specimens have dark gray fur and long ears, whereas K. pellucida has orange-brown fur and smaller ears.</p></div>	https://treatment.plazi.org/id/03E5B46DCE670763FEC0FD01FDFAFAA1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE670763FF2FFAE4FA1EFD28.text	03E5B46DCE670763FF2FFAE4FA1EFD28.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kerivoula intermedia Hill and Francis 1984	<div><p>Kerivoula intermedia Hill and Francis 1984</p><p>Small Woolly Bat (Local name not available)</p><p>Specimens examined (3).—Krau: TK 152019 (GenBank EU188789), TTU 108181, TTU 108217.</p><p>Type locality.— Malaysia (Borneo), Sabah, Lumerao.</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (Pahang: Krau) and Borneo.</p><p>Remarks.—This species was captured using harp traps along with K. papillosa and K. minuta in Krau. It is slightly larger, with wider ears, and lacks the conspicuous banding (dark bases) of dorsal fur when compared to K. minuta . We originally assigned our specimens to K. whiteheadi based on the presence of an elongate P2 (following Corbet and Hill 1992). Subsequently, we learned that K. whiteheadi has a more elongate muzzle, with premolars that are oval in cross section and nearly twice as long as they are wide (C. M. Francis, pers. comm.). These characteristics were not obvious in our specimens. Kerivoula intermedia is difficult to distinguish from K. minuta as the unique character separating them is size. Genetic distance value of 17.26% in cyt- b gene between K. intermedia and K. minuta suggests that genetic data is crucial to provide better discrimination between these species.</p></div>	https://treatment.plazi.org/id/03E5B46DCE670763FF2FFAE4FA1EFD28	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE670763FC16FD7AFACAFA0E.text	03E5B46DCE670763FC16FD7AFACAFA0E.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kerivoula lenis Thomas 1916	<div><p>Kerivoula lenis Thomas 1916</p><p>Lenis Woolly Bat (Local name not available)</p><p>Specimens examined (2).—Krau: TTU 108210 (GenBank EU188772); Kubah NP: TTU 108326.</p><p>Type locality.— India, Calcutta .</p><p>Malaysian distribution.—The current study provides the first confirmed record of this species in both Peninsular Malaysia and Borneo. The distribution of K. lenis partially overlaps that of K. papillosa, a larger species of Kerivoula .</p><p>Remarks.— Kerivoula lenis was caught using harp traps in habitat similar to where K. papillosa and K. intermedia were captured. This species was recently recognized as a valid species from the K. papillosa complex through morphological evaluations (Vanitharani et al. 2003; Fig. 3e). Kerivoula lenis has a genetic distance of 3.92% in the cyt- b gene sequence data between individuals from Peninsular Malaysia and Borneo. See K. papillosa account for details.</p></div>	https://treatment.plazi.org/id/03E5B46DCE670763FC16FD7AFACAFA0E	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE670762FC13F99CFDFCFB47.text	03E5B46DCE670762FC13F99CFDFCFB47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kerivoula minuta Miller 1898	<div><p>Kerivoula minuta Miller 1898</p><p>Least Woolly Bat (Kelawar Hutan Terkecil)</p><p>Specimens examined (6).—Krau: TTU 108182 (GenBank EU188774); Monggis: TTU 108230, TTU 108234; Kubah NP: TTU 108339; Mt. Penrisen: TTU 108351, TTU 108415.</p><p>Type locality.— Thailand, Trang Province, Lay Song Hong .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (Kedah: Ulu Muda; Pahang: Lakum, Taman Negara; Perak: Maxwell Hill; and Selangor: Hulu Gombak, Hulu Langat) and Borneo (Sabah: Gomantong, Madai, Pulau Balembangan, Tabin, Tawau, and Witti Range; Sarawak: Lambir NP, and Mt. Penrisen).</p><p>Remarks.— Kerivoula minuta was caught in harp traps across walking trails and less effectively in mist nets across small closed ponds. This species has small ears (9-11.5 mm), dorsal hairs with bases dark-brown for almost one-third of their length, and light brown tips. Kerivoula minuta has a short forearm (FA &lt;30.5 mm), dark wing membranes and uropatagium, and the proximal dorsal half of the uropatagium has sparse, long, golden brown hairs. A new species, K. krauensis (Francis et al. 2007) described from Peninsular Malaysia that occurs in sympatry with K. minuta, can be distinguished by fur coloration (dorsal hairs with dark-brown bases for almost one-third of their length with tips light brown in K. minuta). Kerivoula minuta also occurs with K. intermedia and is easily misidentified as K. intermedia because the only character to differentiate between them is size. A genetic distance of 4.48% in the cyt- b gene was observed between Peninsular Malaysian and Bornean populations of K. minuta .</p></div>	https://treatment.plazi.org/id/03E5B46DCE670762FC13F99CFDFCFB47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE660762FECFFB47FAAEFE25.text	03E5B46DCE660762FECFFB47FAAEFE25.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kerivoula papillosa (Temminck 1840)	<div><p>Kerivoula papillosa (Temminck 1840)</p><p>Papillose Woolly Bat (Kelawar Hutan Besar)</p><p>Specimens examined (4).—Krau: TK 152020 (GenBank EU188782), TTU 108218, TTU 108219, TTU 108183.</p><p>Type locality.— Indonesia, Java, Bantam (restricted by Tate, 1940) .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (all states) and Borneo.</p><p>Remarks.— Kerivoula papillosa was captured using harp traps set along a walking trail. This species is the largest Kerivoula species (FA: 38-49 mm) recorded in this study and this measurement distinguishes it from other species in the genus. Previous literature suggested that K. papillosa might have included K. lenis, which occurs in broad sympatry with K. papillosa (Vanitharani et al. 2003) . However, K. lenis has smaller body size than K. papillosa (Vanitharani et al. 2003) and our study documented a genetic distance of 10% between these species in the cyt- b gene. Specimens of K. papillosa have a geographic variation of 3.24% in the cyt- b gene between Peninsular Malaysia and Borneo.</p></div>	https://treatment.plazi.org/id/03E5B46DCE660762FECFFB47FAAEFE25	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE660762FC4EFE69FB33FB79.text	03E5B46DCE660762FC4EFE69FB33FB79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Kerivoula pellucida (Waterhouse 1845)	<div><p>Kerivoula pellucida (Waterhouse 1845)</p><p>Clear-winged Woolly Bat (Kelawar Kepak Jernih)</p><p>Specimen examined (1).— Krau: TTU 108213 (GenBank EU188788) .</p><p>Type locality.— Philippines .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia and Borneo.</p><p>Remarks.— Kerivoula pellucida was caught using a harp trap set across a forest trail. Several unique characters were observed in this species: orange to pinkish facial coloration, glandular swelling on the tail of male specimens, and translucent wings. Caution needs to be taken when determining the degree of translucency, as it is best done by comparison between specimens of each species. Kerivoula pellucida has a genetic distance of 1.29% in the cyt- b gene between individuals from Peninsular Malaysia and Borneo.</p></div>	https://treatment.plazi.org/id/03E5B46DCE660762FC4EFE69FB33FB79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
03E5B46DCE66076DFC7AFB6AFBCBFE7B.text	03E5B46DCE66076DFC7AFB6AFBCBFE7B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Miniopterus magnater Sanborn 1931	<div><p>Miniopterus magnater Sanborn 1931</p><p>Western Long-fingered Bat (Kelawar Jari Panjang Besar)</p><p>Specimen examined (1).— Monggis: TTU 108242 (GenBank EU521636) .</p><p>Type locality.— Papua New Guinea, E Sepik, Marienberg .</p><p>Malaysian distribution.—Recorded from Peninsular Malaysia (uncertain locality) and Borneo (Sabah: Gomantong Cave, Madai, Mt. Kinabalu, Poring,Tawau Hills, and upper Sungai Kuamut).</p><p>Remarks.— Miniopterus magnater was caught using a hand held mist net when it was hawking insects around lights near the Monggis station headquarters. Cranial measurements were used to identify this species (Hill 1983; Corbet and Hill 1992). Characters used to identify this species in the field are subtle and this species is usually confused with Miniopterus schrebersii (see Hendrichsen et al. 2001). Comparison with available GenBank sequences showed that this species aligned closely with M. schrebersii and M. magnater . A genetic distance of 0.7% in the cyt- b gene sequence variation was observed between our specimen and an individual from China (EF517308) available on Genbank.</p></div>	https://treatment.plazi.org/id/03E5B46DCE66076DFC7AFB6AFBCBFE7B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	FAISAL ALI ANWARALI KHAN;VICKI J. SWIER;SERGIO SOLARI;PETER A. LARSEN;BESAR KETOL;WAHAP MARNI;SIVANATHAN ELLAGUPILLAY;MAKLARIN LAKIM;MOHAMMAD TAJUDDIN ABDULLAH;ROBERT J. BAKER	FAISAL ALI ANWARALI KHAN, VICKI J. SWIER, SERGIO SOLARI, PETER A. LARSEN, BESAR KETOL, WAHAP MARNI, SIVANATHAN ELLAGUPILLAY, MAKLARIN LAKIM, MOHAMMAD TAJUDDIN ABDULLAH, ROBERT J. BAKER (2008): USING GENETICS AND MORPHOLOGY TO EXAMINE SPECIES DIVERSITY OF OLD WORLD BATS: REPORT OF A RECENT COLLECTION FROM MALAYSIA. Occasional Papers of the Museum 281: 1-30, DOI: 10.5281/zenodo.15757593
