identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03ED8780FF97FFFF74A9FD2B7BE1FC92.text	03ED8780FF97FFFF74A9FD2B7BE1FC92.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Belicinus Arce-Perez & Shepard 2012	<div><p>Genus Belicinus Arce-Pérez &amp; Shepard, 2012</p><p>Monotypic taxon characterized by slender and clearly arched tibiae (Fig. 2) with tiny teeth on the inner margin, as well as a pronotum with a longitudinal glabrous line and a weak carina on its distal half (Arce-Pérez et al. 2012).</p></div>	https://treatment.plazi.org/id/03ED8780FF97FFFF74A9FD2B7BE1FC92	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF97FFFF74A9FCC37FB6FA46.text	03ED8780FF97FFFF74A9FCC37FB6FA46.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Belicinus rhomboideus Arce- Perez & Shepard 2012	<div><p>Belicinus rhomboideus Arce-Pérez &amp; Shepard, 2012</p><p>Figs. 1–4</p><p>Diagnosis. Adult male (Figs. 1, 2): body oval and depressed, with widespread vestiture of long and short setae, longer and darker on venter. Head long with a longitudinal furrow on postoccipital region, concave between the eyes; eyes spherical, very prominent; clypeus subtrapezoidal; antennae short, filiform to subserrate, hardly reaching scutellum. Maxillary palpus short, with 4 palpomeres, three basal ones subconical, apical palpomere ovoid. Pronotum trapezoidal, anterior margin arcuate, posterior margin bisinuate and almost as wide as elytral base, lateral margins rounded and unexpanded; with a longitudinal glabrous line and a weak carina on distal half, with a slight depression on each side. Male genitalia: Trilobate, ventrally subrectangular; parameres subtrapezoidal, shorter than middle lobe, with external margin slightly sinuate, ending in a truncate and emarginate apex (Fig. 3); penis in ventral view with expanded rhomboidal shape, with small spines on surface (Fig. 4).</p><p>Material examined. MEXICO: Chiapas: Marqués de Comillas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-90.90984&amp;materialsCitation.latitude=16.125809" title="Search Plazi for locations around (long -90.90984/lat 16.125809)">Río Lagartos</a>, 16°07’32.91”N, 90°54’35.42”W, 150 m, 10.i.2017, Barba, López, Benítez, 2 males, D net (CNIN); same data except 30.i.2019, Barba, 1 male (CNIN) .</p><p>Geographical distribution. BELIZE, Belize, Cayo, Orange Walk, Stann Creek, and Toledo districts. MEXICO, Chiapas, first country record .</p></div>	https://treatment.plazi.org/id/03ED8780FF97FFFF74A9FCC37FB6FA46	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF97FFFF74A9FF7B7F30FDBA.text	03ED8780FF97FFFF74A9FF7B7F30FDBA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psepheninae Lacordaire 1854	<div><p>Subfamily Psepheninae Lacordaire, 1854</p><p>Adult diagnosis. Head hypognathous, concealed by or deeply inserted into prothorax; clypeus produced anteriorly and sharply recurved in front of antennal insertions (e.g., Psephenotarsis); corpotentorium narrow, without median process, or reduced; labrum partly exposed; antenna with 6–11 antennomeres, usually sexually dimorphic, in males filiform (e.g., Pheneps), moniliform (e.g., Psephenus, Psephenops, Psephenotarsis), or filiform to subserrate (e.g., Belicinus and Psephenopalpus); maxillary palps well developed, with 4 palpomeres, apex securiform; labium with trapezoidal mentum, labial palps with 2 or 3 palpomeres, ligula reduced (Arce-Pérez 2004; Arce-Pérez et al. 2012; Lee et al. 2016). Males with seven visible abdominal ventrites and females with only six visible abdominal ventrites (Lee et al. 2007).</p></div>	https://treatment.plazi.org/id/03ED8780FF97FFFF74A9FF7B7F30FDBA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF97FFFF74A9F9B77AE9F93A.text	03ED8780FF97FFFF74A9F9B77AE9F93A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psephenopalpus Arce-Perez 2004	<div><p>Genus Psephenopalpus Arce-Pérez, 2004</p><p>Monotypic taxon characterized by a pronotum with lateral margins rounded, not expanded on posterior half, with a feeble carina; maxillary palpus extremely long and second palpomere much longer than other palpomeres; tarsomeres with short ventral expansions that do not cover apical tarsomere (Arce-Pérez et al. 2012).</p></div>	https://treatment.plazi.org/id/03ED8780FF97FFFF74A9F9B77AE9F93A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF97FFF974A9F8AB7B33FE72.text	03ED8780FF97FFF974A9F8AB7B33FE72.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psephenopalpus browni Arce-Perez 2004	<div><p>Psephenopalpus browni Arce-Pérez, 2004</p><p>Figs. 5–8</p><p>Diagnosis. Adult male (Figs. 5, 6): body oval and tapered anteriorly. Head black, short, transversely with very prominent and spherical eyes.Antenna long, filiform subserrate. Maxillary palpus extremely long, second palpomere much longer than other palpomeres. Male genitalia. Trilobate, in dorsal view parameres broad, shorter than middle lobe with external margins sinuate distally, ending in rounded apex with lateral membrane (Fig. 7). In ventral view, penis longer than parameres, digitiform, widening at basal half with robust longitudinal, sublanceolate sclerite (Fig. 8).</p><p>Material examined. MEXICO: Chiapas: Cacahoatán, Ejido El Águila, cascada <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.1795&amp;materialsCitation.latitude=15.093611" title="Search Plazi for locations around (long -92.1795/lat 15.093611)">La Sirena</a>, 15°06’08.8”N, 92°11’01.3”W, 1115 m, 06.xii.2018, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.1795&amp;materialsCitation.latitude=15.093611" title="Search Plazi for locations around (long -92.1795/lat 15.093611)">Luna-Luna</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.1795&amp;materialsCitation.latitude=15.093611" title="Search Plazi for locations around (long -92.1795/lat 15.093611)">Cancino-López</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.1795&amp;materialsCitation.latitude=15.093611" title="Search Plazi for locations around (long -92.1795/lat 15.093611)">Almaraz-Hernández</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.1795&amp;materialsCitation.latitude=15.093611" title="Search Plazi for locations around (long -92.1795/lat 15.093611)">Marquez-López</a>, 1 male, D net (CNIN); Cacahoatán, Ejido El Águila, cascada La Sirena, 15°05’37.0”N, 92°10’46.2”W, 1250 m, 24.iii.2022, Chacón-Hartleven, Chamé-Vázquez, 2 males, Surber sampler (ECO-TAP-E); same data except 21.iv.2022, 15 males, Surber sampler, 4 males, collected manually (ECO-TAP-E) .</p><p>Geographical distribution. MEXICO, Chiapas (new state record) and Veracruz.</p><p>Habitat. The specimens from Ejido El Águila were collected from a wadable stream in the sub-basin Alto Cahoá, a tributary of the Cahoacán River, which drains areas with remnants of primary and secondary montane mesophyll forest. Most specimens were collected in riffles with Surber sampler, where water temperature was 18.73 °C (± 0.34 SD, standard deviation) and dissolved oxygen concentration was 13.41 mg /L (± 0.25 SD). In addition, a few specimens were collected manually on the periphyton of water-covered rocks. High habitat heterogeneity was observed, with an optimal condition of physical habitat (RBP = 0.90), including abundant riparian native vegetation with nearby coffee plantations (shade-grown coffee, Coffea sp.). The stream had turbulent water flow, the elements of the channel were dominated by riffles and rapids zones (60% of total reach sampled), and composition of the inorganic substrate was dominated by cobbles and boulders (45% and 35% of reach sampled, respectively). Also, abundant allochthonous elements (e.g., woody material and leaf packs) were observed in the stream channel.</p></div>	https://treatment.plazi.org/id/03ED8780FF97FFF974A9F8AB7B33FE72	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF91FFF974A9FDE27DD2FD76.text	03ED8780FF91FFF974A9FDE27DD2FD76.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psephenops Grouvelle 1898	<div><p>Genus Psephenops Grouvelle, 1898</p><p>This genus is recognized by a pronotum expanded laterally on posterior half, with a well-developed carina; maxillary palp is short with last palpomere the longest; first and second tarsomeres, and occasionally the third, with wide ventral expansion that covers apical tarsomere (Arce-Pérez et al. 2012).</p></div>	https://treatment.plazi.org/id/03ED8780FF91FFF974A9FDE27DD2FD76	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF91FFFB74A9FCE67FFCF9D6.text	03ED8780FF91FFFB74A9FCE67FFCF9D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psepehenops arcei Luna-Luna, Chacon-Hartleven & Contreras-Ramos 2025	<div><p>Psepehenops arcei Luna-Luna, Chacón-Hartleven &amp; Contreras-Ramos, sp. nov.</p><p>Figs. 9–15</p><p>Diagnosis. Adult male (Figs. 9–15): body oval and depressed; head short, transverse, antennae short, not reaching pronotal base, moniliform; pronotum subtrapezoidal, anterior margin arcuate, posterior margin bisinuate, straight in antescutellar area. Pronotal disc with well-developed, medial, longitudinal carina on distal half. Protibia with a tiny apicolateral denticle and long tarsal lobes. Male genitalia (Figs.13–15): short, wide, subrectangular, trilobate; parameres robust and elongate, close to each other with internal margins fused in dorsal view, apex rounded. Penis digitiform, acuminate apically in dorsal view; apically narrowed, directed ventrally. Psephenops arcei, sp. nov. is close to P. triangularis of Panama by sharing smooth elytra and elongated tarsal lobes (lobe of tarsomere 2 covers ventrally the full-length of tarsomeres 3-4 and reaching at least 80% the length of tarsomere 5). Characters of the genitalia may separate males of both species: in the new species (Figs. 13, 14), parameres in dorsal view are medially close to each other, with margins smooth in ventrolateral view, while in P. triangularis the parameres are medially broadly separate, with an evident tooth on inner margin near the apex. The penis in the new species is digitiform, apically acuminate (Figs. 13, 14), while in P. triangularis it is digitiform, with a basal, lateral, subtriangular projection from each side.</p><p>Description. Male: body length X̅ (mean) = 2.7 mm, width X̅ = 1.5 mm (n = 3). Body oval, depressed; integument yellowish-brown, head black; pronotum dark; elytra yellowish-brown, covered with short yellow setae with golden reflections (Figs. 9, 11); coxae, trochanters, and femora reddish-brown; tibiae and tarsi reddish brown with vestiture short, fine and dense, yellowish-gray (Fig. 10).</p><p>Head: cypeus subrectangular, declivous at less than 90 degrees from plane of frons, distal margin widely emarginate; fronto-clypeal surface and vertex with long yellowish-brown setae, vertex minutely punctate. Antennae short (i.e., when fully extended backwards, not reaching posterior margin of pronotum), moniliform, with 11 antennomeres; scape longest, pedicel subglobose, half as long as scape; third antennomere subtriangular, pale brown, shorter than pedicel; antennomeres 4–10 small, moniliform; apical antennomere acuminate and dark brown (Fig. 9). Eyes lateral, rounded, very prominent, reddish-brown, with dark brown postocular area (Fig. 11). Labrum subrectangular, similar to clypeus but shorter. Maxillary palp with 4 palpomeres, covered with long setae, basal palpomere smallest, apical palpomere longest, with wide sensory area, relative length of palpomeres 0.04, 0.08, 0.06, 0.14; palpomeres 1–3 subconical, palpomere 4 acuminate. Labial palp short, with 3 palpomeres, palpomeres 1–2 rounded and wide, palpomere 3 subconical, nipple-shaped and slightly longer than preceding ones.</p><p>Thorax: pronotum width X̅ = 1.14 mm (n = 3); lateral margin length X̅ = 0.60 mm (n = 3). Black, subtrapezoidal, basally almost as wide as elytral base; anterior margin arcuate, posterior margin bisinuate, straight in antescutellar area, expanded posterolaterally; posterolateral angles subacute. Pronotal disc with well-developed, dorsomedial, longitudinal carina on distal half, and wide, oval, deep depression at each side of carina (Fig. 9); pronotal surface finely punctate, with minute, yellowish-brown vestiture, and long, dark brown setae. Scutellum short, apex rounded. Elytra with short yellow setae producing golden shining and long yellowish-brown setae; lateral margins parallel on basal three-fourths, then gradually converging posteriorly; humeral width 1.10 mm; surface without striae; length (from base to apex of suture) 1.45 mm; elytral base dark brown. Prosternum dark brown, short; prosternal process lanceolate, deeply grooved longitudinally, apex concave(Fig. 10) reaching anterior half of mesoventrite; mesoventrite shorter than metaventrite, with wide groove receiving prosternal process; metaventrite large and bulky. Legs (Figs. 10, 12): femora robust; tibiae slender, metatibia longer than metafemur, protibia with tiny apicolateral denticle; tarsomeres 1 and 2 ventrally lobed, with thick sponge-like vestiture; ventral lobe of tarsomere 1 extending over basal 0.30 of tarsomere 2; tarsomere 2 the longest, twice as long as tarsomere 1, extending distally and completely covering tarsomeres 3, 4, and basal 0.80 of tarsomere 5; apical claws long, curved, slender, each with small basal tooth.</p><p>Abdomen: vestiture short, fine, dense, and grayish. Ventrite VI usually hidden under ventrite V; ventrites I and II wide, with posterior margin curved anteriorly in the middle; ventrites III and IV narrow, with posterior margin straight; ventrite V with posterior margin slightly emarginate; ventrite VI short, subtriangular, with posterior margin widely curved posteriorly; ventrite VII or anal plate, oval, subtriangular, narrow, covered with long whitish setae; pygidium oval, partly concealed, emarginate, with long brown setae.</p><p>Male genitalia: short, wide, subrectangular, trilobate (Figs. 13–15); total length 0.60 mm, total width 0.24 mm. Parameres robust and elongate, 0.40 mm long, shorter than penis, uniformly widened, apex rounded; with internal margins fused at basal 0.2 in dorsal view (Fig. 13). Penis length X̅ = 0.49 mm (n = 3), longer than parameres (Figs. 13–15); phallobase concave in dorsal view, bulky-subcylindrical in ventral view, length X̅ = 0.25 mm, width X̅ = 0.23 mm (n = 3) (Fig. 13); apical half, subtriangular, lanceolate in ventral view (Fig. 14); apically narrowed, directed ventrally (Fig. 15).</p><p>Variation. One specimen does not present a dark brown coloration at distal portion of the elytra, body length range was 2.6–2.8 mm (n = 3).</p><p>Holotype. MEXICO: Chiapas: Ocosingo, Reserva de la Biosfera Montes Azules, Estación de Biología Chajul, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-90.93989&amp;materialsCitation.latitude=16.110666" title="Search Plazi for locations around (long -90.93989/lat 16.110666)">Río Lacantún</a> (embarcadero), 16°06’38.4”N, 90°56’23.6”W, 154 m, 12.i.2017, Barba, 1 male, D net (CNIN).</p><p>Paratypes. MEXICO: Chiapas: Marqués de Comillas, río Ixcán (embarcadero), Rta.307 Palenque-La <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.0917&amp;materialsCitation.latitude=16.092903" title="Search Plazi for locations around (long -91.0917/lat 16.092903)">Trinitaria</a>, 16°05’22.43”N, 91°05’19.42”W, 150 m, 22.ii.2016, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.0917&amp;materialsCitation.latitude=16.092903" title="Search Plazi for locations around (long -91.0917/lat 16.092903)">Barba</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.0917&amp;materialsCitation.latitude=16.092903" title="Search Plazi for locations around (long -91.0917/lat 16.092903)">Mayorga</a>, 1 male (CNIN); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.0917&amp;materialsCitation.latitude=16.092903" title="Search Plazi for locations around (long -91.0917/lat 16.092903)">Marqués de Comillas</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-91.0917&amp;materialsCitation.latitude=16.092903" title="Search Plazi for locations around (long -91.0917/lat 16.092903)">río Ixcán</a> (embarcadero), Rta. 307 Palenque-La Trinitaria, 16°05’34.45”N, 91°05’30.11”W, 166 m, 8.i.2017, Barba, 1 male (CNIN) .</p><p>Geographical distribution. MEXICO, Chiapas, Ocosingo and Marqués de Comillas municipalities.</p><p>Etymology. This species is named after entomologist Roberto Arce-Pérez, for his contribution to the taxonomy of aquatic Coleoptera of Mexico, including Psephenidae .</p><p>Habitat. Specimens were collected in the riparian zones of large, unwadable rivers. The holotype was collected in the Lacantún River, a tributary of Usumacinta river, draining remnants of tropical rainforest, and characterized by a highly turbulent water flow, with channel elements dominated by fast-flowing habitats (e.g., rapids and riffles), and inorganic substrate dominated by cobbles. Specimens from Marqués de Comillas (paratypes) were collected at Ixcán River, from the sub-basin Lacantún River. The river had scarce riparian vegetation and a highly turbulent water flow, with channel elements dominated by fast-flowing habitats (e.g., rapids and riffles) and inorganic substrate dominated by pebbles.</p></div>	https://treatment.plazi.org/id/03ED8780FF91FFFB74A9FCE67FFCF9D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF93FFFA74A9F9337AADFB97.text	03ED8780FF93FFFA74A9F9337AADFB97.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psephenops shepardi Arce-Perez 2013	<div><p>Psephenops shepardi Arce-Pérez, 2013</p><p>Figs. 16–19</p><p>Diagnosis. Adult male (Figs. 16, 17): body oval and depressed, head short, transverse, antenna long and moniliform; pronotum subtrapezoidal, anterior margin arcuate, posterior margin bisinuate as wide as elytral base, lateral margins straight along anterior half and expanded laterally on posterior half. Pronotal disc with median longitudinal carina well developed on distal half. Protibia with a denticle and tarsal lobes short. Male genitalia (Figs. 18, 19): short, wide, subrectangular, trilobate with parameres subparallel in apical third, medial margin of each paramere separated by wide U-shaped cleft, with tips of parameres not truncate and not divergent, basal half of parameres fused, ventral sclerite of penis subtriangular with apex bifid.</p><p>Material examined. MEXICO: Chiapas: Pichucalco, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.11472&amp;materialsCitation.latitude=17.482779" title="Search Plazi for locations around (long -93.11472/lat 17.482779)">Tectuapan</a>, 17°28’58.0”N, 93°06’53.0”W, 46 m, [?]. v.2017, Castillo-Uzcanga, Espinoza-Toledo, 2 males, Surber sampler (ECO-TAP-E) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.05728&amp;materialsCitation.latitude=17.558582" title="Search Plazi for locations around (long -93.05728/lat 17.558582)">Pichucalco</a>, sitio experimental de INIFAP, 17°33’30.9”N, 93°03’26.2”W, 54 m, 29.v.24, Chacón-Hartleven, 2 males (ECO-TAP-E) ; <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.093056&amp;materialsCitation.latitude=17.427757" title="Search Plazi for locations around (long -93.093056/lat 17.427757)">Ixtacomitán</a> (17°25’39.93”N, 93°05’34.98”W), 193 m, 30.v.2024, 1 male (ECO-TAP-E) ; Veracruz: Los Tuxtlas, 18.vi.1985, Muñoz, 1 male (CNIN) ; same data except 18.v.2015, Contreras-Ramos, 1 male, D net (CNIN); Los Tuxtlas, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.09072&amp;materialsCitation.latitude=18.62111" title="Search Plazi for locations around (long -95.09072/lat 18.62111)">Río Máquinas</a>, 18°37’16.0”N, 95°05’26.6”W, 21 m, 29.iv.2016, Contreras-Ramos, 9 males, D net (CNIN) .</p><p>Geographical distribution. BELIZE, Cayo and Toledo districts; MEXICO, Chiapas and Veracruz states - first records for the country .</p><p>Habitat. Specimens from Tectuapan, Chiapas, were collected in fast-flowing zones of wadable streams in the Pichucalco River sub-basin, a tributary of the Grijalva River. Two specimens were collected in rapids of a stream that drains areas of cultivated pasture, where water temperature was 27.16 °C (± 2.14 SD) and dissolved oxygen concentration was 8.31 mg /L (± 1.04 SD). The stream had a good physical habitat condition (RBP = 0.67), with channel elements dominated by habitats with fast-flowing water, specifically rapid zones (40% of reach sampled) and inorganic substrates dominated by cobble and gravel (40% and 35% of reach sampled, respectively) (Espinoza-Toledo 2018). Two specimens were collected in riffles of a stream that drains areas of pristine primary forest with abundant riparian vegetation, where water temperature was 28.65 °C (± 0.35 SD) and dissolved oxygen concentration was 6.61 mg /L (± 0.29 SD). This stream had an excellent quality of physical habitat (RPB = 0.95), with channel elements dominated by riffle and rapids zones (&gt;65% of reach sampled) and inorganic substrates dominated by cobbles (70% of reach sampled). The specimen from Ixtacomitán, Chiapas, was collected in riffles of a stream from the Pichucalco River sub-basin, a tributary of the Grijalva River, running through human settlements. The stream presented a moderate physical habitat condition (RBP = 0.60), with direct wastewater discharge into the channel and abundant inorganic and organic waste materials; despite abundant riparian vegetation, anthropic contamination and alterations are evident. Channel elements were dominated by rapids (50 % of reach sampled), where water temperature was 26.71 °C (± 0.23 SD), dissolved oxygen concentration was 7.13 mg /L (± 0.34 SD), and inorganic substrates were dominated by pebbles and sand (50% and 30% of reach sampled, respectively).</p></div>	https://treatment.plazi.org/id/03ED8780FF93FFFA74A9F9337AADFB97	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF92FFFA74A9FBC67C83FAB7.text	03ED8780FF92FFFA74A9FBC67C83FAB7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psephenus Haldeman 1853	<div><p>Genus Psephenus Haldeman, 1853</p><p>Species of Psephenus are characterized by antennae short, scarcely or not extending beyond elytral humeri, subserrate; tibiae straight, without teeth; pronotum without a longitudinal glabrous line, without carina; tarsomeres 1–4 subconical, slender, slightly emarginate at apex, tarsomeres 1–2 usually with ventral setae or papillae that never cover contiguous tarsomere (Arce-Pérez et al. 2012).</p></div>	https://treatment.plazi.org/id/03ED8780FF92FFFA74A9FBC67C83FAB7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF92FFF774A9FA267DF3FDE6.text	03ED8780FF92FFF774A9FA267DF3FDE6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psephenus palpalis Champion 1913	<div><p>Psephenus palpalis Champion, 1913</p><p>Figs. 20–23</p><p>Diagnosis. Adult male (Figs. 20, 21): body oblong-ovate, rather broad, depressed, dorsum shining, densely pubescent. Head profusely rugulose, vertex foveate, dorsal concavity not divided. Elytra oblong, little wider than thorax; pronotal sides strongly convergent anteriorly, apex not more than half as wide as base. Maxillary palp threequarters length of antenna, second segment of maxillary palp usually longer than third and fourth combined. Male genitalia (Figs. 22, 23): apex of median penial spicule enlarged and bluntly rounded.</p><p>Material examined. MEXICO: Chiapas: Cacahoatán, Ejido Ahuacatlán (15°02’20.4”N, 92°10’15.6”W), 680 m, 22.iii.22, Chacón-Hartleven, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-92.171&amp;materialsCitation.latitude=15.0390005" title="Search Plazi for locations around (long -92.171/lat 15.0390005)">Chamé-Vázquez</a>, 1 male, D net (ECO-TAP-E) ; Pichucalco, Tectuapan, 17°28’47.74”N, 93°07’21.32”W, 66 m, 8.iii.2024, Chacón-Hartleven, 3 males, Surber sampler (ECO-TAP- E); same data except 01.iv.2024, 1 male (ECO-TAP-E); same data except 30.iv.2024, 2 males (ECO-TAP-E); Pichucalco, Tectuapan, 17°26’13.30”N, 93°09’31.76”W, 99 m, 29.iv.2024, 2 males (ECO-TAP-E); Pichucalco, sitio experimental del INIFAP, 17°33’30.9”N, 93°03’26.2”W, 54 m, 29.v.24, 1 male (ECO-TAP-E); Guerrero: Cacahuamilpa, Río Dos Bocas, 1.iv.1984, Vázquez, Villa, 1 male (CNIN) ; R[uta] 1[3]4 Altamirano-Zihuatanejo Km 102, 22-vi-1990, 1300 m, Bueno, Barrera, 1 male (CNIN); Morelos: Amacuzac, Río Chalma (ojo de agua), 25.iii.1984, Vázquez, 1 male (CNIN) ; Tlaltizapán, Balneario Ejidal Sta. Isabel, 150 m, 16.iv.1997, Bueno, Barba, Rojas, González, 1 male (CNIN); Oaxaca, Teotitlán, Río Xiquila, 5.v.1988, Cervantes, 2 males (CNIN) ; San Juan Bautista Cuicatlán, (Santiago) Dominguillo, río Las Huertas, 12.vi.1996, Barba, Rojas, 1 male (CNIN) ; Puebla: Telpatlán, 3.x.1988, Cervantes, 1 male (CNIN); Tabasco: Huimanguillo, Villa G [uadalu]pe 2da. Secc., arroyo Las Flores, Carr [etera] Malpasito-Carlos A. Madrazo 5.92 km, 17°22’30.0”N 93°36’15.0”W, 12.vi.1997, Bueno, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-93.604164&amp;materialsCitation.latitude=17.375" title="Search Plazi for locations around (long -93.604164/lat 17.375)">Barba</a>, 1 male (CNIN) ; Veracruz: Los Tuxtlas, Río Máquinas, 14.vii.1979, Bueno, 1 male (CNIN) ; Los Tuxtlas, Balzapote, 5.v.1981, Hernández, Ortega, 2 males (CNIN), Santiago, 1 male (CNIN); same data except 14.v.1984, Rodríguez, 2 males (CNIN); Los Tuxtlas, río La Palma, 18.vi.1996, Alatorre-Castillo, 1 male (CNIN) ; Los Tuxtlas, río Máquinas, 18°37’16”N, 95°05’26.6”W, 21 m, 29.iv.2016, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=-95.09072&amp;materialsCitation.latitude=18.62111" title="Search Plazi for locations around (long -95.09072/lat 18.62111)">Contreras-Ramos</a>, 2 males, D-net (CNIN) .</p><p>Geographical distribution. MEXICO, Chiapas (new state record), Estado de México, Guerrero (new state record), Hidalgo, Morelos, Oaxaca (new state record), Puebla (new state record), Tabasco (new state record), and Veracruz (new state record) (Fig. 25).</p><p>Habitat. The specimen from Ahuacatlán, Chiapas was collected from emergent macrophytes at the riparian zone of a wadable stream of the sub-basin Bajo Cahuá, a tributary of the Cahoacán River. The stream drains agricultural areas, with sun coffee plantations ( Coffea sp.) and exotic eucalypt trees ( Eucalyptus sp.) along the riparian zone. The specimen came from riffles, where water temperature was 21.93 °C (± 0.63 SD) and the dissolved oxygen concentration was 13.68 mg /L (± 0.78 SD). The stream had a good physical habitat condition (RBP = 0.66) with turbulent water flow, channel elements were dominated by riffles and rapids (60% of reach sampled), and inorganic substrate was dominated by pebbles and gravel (35% and 30% of reach sampled, respectively). The specimens from Tectuapan, Chiapas were collected in fast-flowing zones of wadable streams of the Pichucalco River sub-basin, tributaries of the Grijalva River. Six specimens came from riffles of a stream that drains areas with mixed land uses, including primary-secondary forests and cultivated pastures, where water temperature was 28.68 °C (± 3.75 SD) and dissolved oxygen concentration was 4.55 mg /L (± 0.27 SD). Two specimens were found in riffles of a stream that drains areas of secondary forest, where water temperature was 30.42 °C (± 0.62 SD) and dissolved oxygen concentration was 7.05 mg /L (± 0.51 SD). One specimen was found in riffles of a stream that drains areas of primary forest with abundant riparian vegetation and pristine conditions, where the water temperature was 28.65 °C (± 0.35 SD) and dissolved oxygen concentration was 6.61 mg /L (± 0.29 SD). The stream had a physical habitat condition ranging from good to excellent (RBP = 0.70, 0.75 and 0.95, respectively), channel elements were dominated by riffles and rapids (50-60% of reach sampled), and inorganic substrates were dominated by pebbles (50% and 60% of reach sampled) and cobbles (70% of reach sampled), respectively.</p></div>	https://treatment.plazi.org/id/03ED8780FF92FFF774A9FA267DF3FDE6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
03ED8780FF98FFF074A9FF337BF5FA86.text	03ED8780FF98FFF074A9FF337BF5FA86.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Psephenops Grouvelle 1898	<div><p>Key to adult males of New World species of Psephenops Grouvelle</p><p>(Based on Arce-Pérez &amp; Novelo-Gutiérrez 2017 and Barr &amp; Shepard 2024; the species P. robacki (Spangler, 1966), was not included in the key because only description of the female is available).</p><p>1. Pronotum with 3 small tuberculiform prominences; elytra with several longitudinal ridges on disc.............. grouvellei</p><p>- Pronotum without tuberculiform prominences; elytra without longitudinal ridges on disc............................. 2</p><p>2. Protibiae without apicolateral denticle; male genitalia as in figure 26...................................... prestone</p><p>- Protibiae with apicolateral denticle; male genitalia variously arranged............................................ 3</p><p>3. Tarsal lobes enlarged (lobe of tarsomere 2 covers ventrally the full-length of tarsomeres 3-4 and reaching at least 80% the length of tarsomere 5)....................................................................................... 4</p><p>- Tarsal lobes short (reaching the basal half or less of tarsomere 5)................................................ 5</p><p>4. Basal half of parameres, in dorsal view, separated by a deep cleft (Fig. 32)........................................ 7</p><p>- Basal half of parameres, in dorsal view, fused (Fig. 37)....................................................... 9</p><p>5. Parameres apically truncated (Fig. 27); 4th maxillary palpomere at least twice as long as preceding palpomeres together smithi</p><p>- Parameres not apically truncated; 4th maxillary palpomere not as above.......................................... 6</p><p>6. Parameres strongly convergent on apical third, medial margins of parameres separated from each other by a narrow, drop-like cleft (Fig. 28).................................................................................... lupita</p><p>- Parameres more or less parallel on apical third, medial margins of parameres separated from each other by a wide, U-shaped cleft (Fig. 29).................................................................................. shepardi</p><p>7. Apex of parameres, in dorsal view, spine-like, widely divergent (Fig. 30)............................... spiniparameri</p><p>- Apex of parameres, in dorsal view, not spine-like, convergent (Figs. 28, 29)....................................... 8</p><p>8. Prosternal process flat, with apex lanceolate. Penis with a small, lateral tooth (Fig. 31)....................... mexicanus</p><p>- Prosternal process subcylindrical, with apex bifid. Penis lacking lateral projections (Fig. 32)...................... bifidus</p><p>9. Small individuals, length less than 2.4 mm; antennae moniliform-serrate; parameres truncate, with apex straight (Fig. 33)............................................................................................. panamaensis</p><p>- Large individuals, length greater than 2.5 mm; antennae not serrate; parameres not truncate......................... 10</p><p>10. Antennae long, filiform; fore- and middle legs with segments 1 and 2 lobulate, hind legs only with segment 1 lobulate (Haiti)............................................................................................. haitianus</p><p>- Antennae short, moniliform; all legs with segments 1 and 2 lobulate............................................ 11</p><p>11. Elytra slightly striate.................................................................................. 12</p><p>- Elytra smooth....................................................................................... 14</p><p>12. Parameres in dorsal view widened on apical fifth, with apex rounded; penis slightly produced apically (Fig. 34). maculicollis</p><p>- Parameres in dorsal view uniformly broad; penis not produced apically......................................... 13</p><p>13. Parameres in dorsal view short, with apices broadly but unevenly rounded; penis with apex semi-truncate (Fig. 35)...... trini</p><p>- Parameres in dorsal view elongate, apices evenly rounded; penis with apex rounded (Fig. 36)............... argentinensis</p><p>14. Parameres in dorsal view broadly separate, apex subtriangular; penis digitiform, with a basal, lateral, subtriangular projection from each side (Fig. 37)........................................................................ triangularis</p><p>- Parameres in dorsal view close to each other, apex rounded (Fig. 38); penis acuminate apically (Figs. 13, 14).. arcei, sp. nov.</p></div>	https://treatment.plazi.org/id/03ED8780FF98FFF074A9FF337BF5FA86	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Chacón-Hartleven, Erick A.;Luna-Luna, Alba Magali;Chamé-Vázquez, Eduardo R.;Mejía-González, Gamaliel;Contreras-Ramos, Atilano	Chacón-Hartleven, Erick A., Luna-Luna, Alba Magali, Chamé-Vázquez, Eduardo R., Mejía-González, Gamaliel, Contreras-Ramos, Atilano (2025): A new species of Psephenops Grouvelle, 1898, with new distribution data and habitat notes of the psephenine water-penny beetles from Mexico (Coleoptera: Psephenidae: Psepheninae). Zootaxa 5722 (1): 79-94, DOI: 10.11646/zootaxa.5722.1.3, URL: https://doi.org/10.11646/zootaxa.5722.1.3
