taxonID	type	description	language	source
03F6A046FF886252EEC2FB22FA6EF99B.taxon	materials_examined	Type: — CHINA. Yunnan: Pingbian County, Daweishan National Natural Reserve, tropical montane moist forests, elevation 1424 m, 23 ° 00 ' 07 " N, 103 ° 41 ' 53 " E, 19 May 2023, C. Y. Hao et al. Hao 0159 (holotype IBSC 0918555!, isotypes IBSC 0918556, IBSC 0918557!). Erect subshrub, 0.7 – 1 m high, aromatic, dioecious. Stem greenish, terete, 3 – 4 mm in diameter, young branchlet sparsely puberulent, glabrescent, finely striate when dry, swollen at node. Leaf dimorphism present, prophyll 1 – 2 cm long. Sympodial leaf membranous to chartaceous, finely glandular, oblong-lanceolate to lanceolate, 10 – 16 cm × 2.5 – 4 cm, base subequilaterally truncate to oblique, one side rounded, the other tapered and acute; apex acutely acuminate to long acuminate, adaxially green and glabrous, abaxially pale greenish and sparsely puberulent on veins; petioles glabrous, 5 – 7 mm long; venation plinerved, veins 5 – 9, reticulate, uppermost distal pair of secondary veins alternate, arising 1 – 3 cm above base, reaching leaf apex, others basal. Monopodial leaf exhibits morphological similarities to sympodial leaf, with the primary distinction being that the base of monopodial leaf is predominantly symmetrical. Inflorescence a pedunculate spike, leaf-opposed, solitary, pendulous, cylindrical, white. Male inflorescence delicate, 10 – 15 cm × 2 – 3 mm; peduncle slender, 3 – 3.5 cm long, glabrous; axis glabrous; bracts imbricate, white, subcircular, 1.5 – 2 mm wide, base often concave, apex slightly protruding, peltate, stalk 0.8 – 1 mm long; stamens 2; filaments marginally longer than anthers, anthers ovoid to globose, 2 - locular. Female inflorescences shorter than male ones, 4 – 8 cm × 3 – 5 mm; peduncle and bracts as in male spikes; rachis sparsely pubescent; ovary ovoid, distinct; stigmas 3 or 4. Infructescence pendulous, 5 – 10 × 0.5 – 0.8 cm, glabrous, cylindrical, white. Fruit single – seeded berry, ± 1.5 mm in diam., ovoid to globose, connate on rachis, glabrous, distinct; floral bracts persistent, white; seed pale brown, ovoid, ± 1 mm in diam., smooth. Phenology: — Flowering from March to June; fruiting from July to November.	en	Fan, Rui, Zhao, Wen-Jing, Su, Fan, Li, Yang, Ji, Xun-Zhi, Su, Bao-Fen, Xia, Nian-He, Hao, Chao-Yun (2025): Piper hongheense (Piperaceae), a new species from Yunnan, China. Phytotaxa 697 (1): 129-137, DOI: 10.11646/phytotaxa.697.1.9, URL: https://doi.org/10.11646/phytotaxa.697.1.9
03F6A046FF886252EEC2FB22FA6EF99B.taxon	etymology	Etymology: — The epithet refers to its distribution, Honghe Prefecture in Yunnan, China.	en	Fan, Rui, Zhao, Wen-Jing, Su, Fan, Li, Yang, Ji, Xun-Zhi, Su, Bao-Fen, Xia, Nian-He, Hao, Chao-Yun (2025): Piper hongheense (Piperaceae), a new species from Yunnan, China. Phytotaxa 697 (1): 129-137, DOI: 10.11646/phytotaxa.697.1.9, URL: https://doi.org/10.11646/phytotaxa.697.1.9
03F6A046FF886252EEC2FB22FA6EF99B.taxon	distribution	Distribution and habitat: — Piper hongheense is currently known only from its type locality in Pingbian county, Honghe Prefecture in Yunnan (Fig. 3), in wet tropical montane forest, at elevations of 1000 – 1600 m. Nevertheless, considering that this new species thrives in the shadow of wet tropical montane forests in Jinping, a region adjacent to Tinh Ha Giang and Tihn Lao Cai in Vietnam, it is highly likely that the species is also present in Vietnam. Taxonomic affinities: — The new species is morphologically similar to P. boehmeriifolium and P. boehmeriifolium var. glabricaule. Following a detailed comparison with specimens and the academic literature (De Candolle 1869, Gilbert & Xia 1999), we have determined that P. hongheense can be unequivocally distinguished by several key characteristics, as outlined in the diagnosis above and summarized in Table 1 and Fig. 4.	en	Fan, Rui, Zhao, Wen-Jing, Su, Fan, Li, Yang, Ji, Xun-Zhi, Su, Bao-Fen, Xia, Nian-He, Hao, Chao-Yun (2025): Piper hongheense (Piperaceae), a new species from Yunnan, China. Phytotaxa 697 (1): 129-137, DOI: 10.11646/phytotaxa.697.1.9, URL: https://doi.org/10.11646/phytotaxa.697.1.9
03F6A046FF886252EEC2FB22FA6EF99B.taxon	materials_examined	Additional specimens examined: — CHINA. Yunnan: Pingbian County of Jinping Prefecture, Shibanxiang, 3 October 1954, G. – M. Feng 04736 (PE!); Pingbian County of Jinping Prefecture, Xinnongxiang, 13 July 1953, P. - Y. Mao 02455 (KUN!); Pingbian County of Jinping Prefecture, Pujijing, 29 March 1958, G. - M. Feng 21778 (KUN!).	en	Fan, Rui, Zhao, Wen-Jing, Su, Fan, Li, Yang, Ji, Xun-Zhi, Su, Bao-Fen, Xia, Nian-He, Hao, Chao-Yun (2025): Piper hongheense (Piperaceae), a new species from Yunnan, China. Phytotaxa 697 (1): 129-137, DOI: 10.11646/phytotaxa.697.1.9, URL: https://doi.org/10.11646/phytotaxa.697.1.9
