identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03FBD87AFFA95877FF5AFC86FDB1FA79.text	03FBD87AFFA95877FF5AFC86FDB1FA79.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyopteris divaricata (Okamura) Okamura 1932	<div><p>* Dictyopteris divaricata (Okamura) Okamura, 1932</p><p>Japanese name: エゾヤハズ (ezoyahazu)</p><p>Basionym: Haliseris divaricata Okamura, 1907</p><p>Syntype localities: —Cape Iwai, Kesen’numa Bay, Miyagi, Prefecture and Hakodate, Japan</p><p>Description and remarks: —This specimen is upright and 10 cm tall. The stipe is about 1.5 mm in diameter but is thinner and less robust as it extends as a midrib to each branch. Branching is dichotomous with obtuse apices. Reproductive sori occur as oblong to slightly elongate black spots and arranged as 2–3 rows on both sides of the midrib.</p><p>D. divaricata grows at 1–2 m on the leeward side of Yokoura’s rocky subtidal. This species is uncommon but can be found growing together with geniculate coralline algae and among other species of Dictyopteris such as D. prolifera and D. undulata in early spring. Previous collection records indicate that the distribution of this species was only found in Korea, China and in Japan’s northern island of Hokkaido and central part of the main island of Honshu and Shikoku (Nagai 1940). However, this species may also be found in tropical regions (Lastimoso &amp; Santiañez 2021), suggesting a much broader temperature tolerance. This specimen may be a representative for its distribution in Japan’s southern Kyushu Island.</p><p>Material examined: —18ECS1-116-1</p></div>	https://treatment.plazi.org/id/03FBD87AFFA95877FF5AFC86FDB1FA79	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA95877FF5AFA0EFD71F8A9.text	03FBD87AFFA95877FF5AFA0EFD71F8A9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyopteris latiuscula (Okamura) Okamura 1932	<div><p>* Dictyopteris latiuscula (Okamura) Okamura, 1932</p><p>Japanese name: ヤハズグサ (yahazugusa)</p><p>Basionym: Haliseris latiuscula Okamura, 1907</p><p>Type locality: — Japan 0</p><p>Description and remarks: —Thalli are upright and measure 8–23 cm tall. The stipe is 2 mm in diameter and has a robust and leathery texture. The blades are 3–13 cm long, lanceolate and grow individually on the stipe. Branching may occur along the midrib giving rise to a new leaf-like blade. Blade margins are usually entire but some may be undulate to slightly crisped and ruffled. The reproductive sori occur as small brown dots and scattered along the blade on both sides of the midrib. This species was collected at Mushima old port and was common in the spring months.</p><p>Materials examined: —18ECS1-56 (1 to 2)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA95877FF5AFA0EFD71F8A9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA95876FF5AF8BEFD67FDC2.text	03FBD87AFFA95876FF5AF8BEFD67FDC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyopteris pacifica (Yendo) Hwang, Kim & Lee 2004	<div><p>* Dictyopteris pacifica (Yendo) Hwang, Kim &amp; Lee, 2004</p><p>Japanese name: コモングサ (komongusa)</p><p>Basionym: Spatoglossum pacificum Yendo, 1920</p><p>Type locality: —Mera, Chiba Prefecture, Japan</p><p>Description and remarks: —Thalli are upright, reaching up to 10–28 cm tall. The entire plant is attached to the substrate by its rhizomatous holdfasts. Arising from the holdfast is a thickened stipe, 3–5 mm in diameter and 1–2 cm in height. The distal parts of the stipe becomes the midrib where the blades arise. The midrib is only found in the lower parts of the plant. The blades are dichotomously branched, 2–5 mm in width and have smooth margins but sometimes have dentate margins near the apices. This species was only found in Aokata and was common during the early summer months.</p><p>Unlike other Dictyopteris spp., the midrib of this species is found only on the lower parts of the plant. This anatomical characteristic of the thallus, along with other evidence from molecular analyses prompted the synonymization of Spatoglossum pacificum to its current nomenclature (Hwang et al., 2004a).</p><p>Materials examined: —18ECS1-159 (1 to 5)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA95876FF5AF8BEFD67FDC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA85876FF5AFD88FD67FC16.text	03FBD87AFFA85876FF5AFD88FD67FC16.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyopteris prolifera (Okamura) Okamura 1930	<div><p>* Dictyopteris prolifera (Okamura) Okamura, 1930</p><p>Japanese name: ヘラヤハズ (herayahazu)</p><p>Basionym: Haliseris prolifera Okamura, 1894</p><p>Lectotype localities: — Totomi, Boshu, Idzumo, Japan</p><p>Description and remarks: —Specimens have a dark-brown thalli with light-brown apical branches. Thalli are upright and 9–25 cm tall and densely branched. The stipe is 1–1.5 mm in diameter and is winged, having 0.5 mm bladelike ridges on both sides. The blades are slender, 2–3.5 mm in width and lanceolate, gradually tapering to the base. Blades arise irregularly along the stipe.</p><p>This species is common along the exposed rocky subtidal at Yokoura at a depth of 1– 2 m. They first appear during mid to late spring and persist until late summer and occur together with Dictyopteris undulata .</p><p>Materials examined: —18ECS1-21 (1 to 15)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA85876FF5AFD88FD67FC16	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA85876FF5AFBE4FD67FA47.text	03FBD87AFFA85876FF5AFBE4FD67FA47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyopteris undulata Holmes 1896	<div><p>Dictyopteris undulata Holmes, 1896</p><p>Japanese name: シワヤハズ (shiwayahazu)</p><p>Type locality: —Misaki, Japan</p><p>Description and remarks: —Thalli are upright and 5–15 cm tall and attached to the rocky substrate through feltlike rhizoids. Stipes are 2–3 mm in diameter, robust and winged. The blades are branched dichotomously and have obtuse to rounded apices. The margins are undulating to ruffled while older specimens have torn and tattered margins and denuded stipes. Plants in their habitat have a bluish iridescent hue but become dark brown when dried.</p><p>This species was found in Yokoura and in both ports at Mushima. Plants are highly seasonal and can be found growing on the rocky subtidal in mid-spring and persists until late autumn.</p><p>Materials examined: —18ECS1-22 (1 to 22)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA85876FF5AFBE4FD67FA47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA85876FF5AFA14FD65F89B.text	03FBD87AFFA85876FF5AFA14FD65F89B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyota coriacea (Holmes) I. K. Kwang, Hy. S. Kim & W. J. Lee 2004	<div><p>* Dictyota coriacea (Holmes) I.K. Kwang, Hy. S. Kim &amp; W.J. Lee, 2004</p><p>Japanese name: サナダグサ (sanadagusa)</p><p>Basionym: Glossophora coriacea Holmes, 1896</p><p>Heterotypic synonym: Pachydictyon coriaceum (Holmes) Okamura, 1899</p><p>Type locality: —Enoura, Japan</p><p>Description and remarks: —Specimens are dark brown, erect, up to 5–13 cm tall with rhizoidal holdfasts. The dichotomously branched apices may bend inwards, causing an overlap between adjacent branches. One specimen shows multi-proliferous branching on one side of an apical branch, possibly due to physical damage, as shown in (Hwang et al. 2004b). This species is common throughout the study sites in spring, often growing together with Colpomenia sinuosa and Dictyopteris spp.</p><p>Materials examined: —18ECS1-111 (1 to 3)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA85876FF5AFA14FD65F89B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA85879FF5AF860FC4AFC4F.text	03FBD87AFFA85879FF5AF860FC4AFC4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyota dimorphosa	<div><p>* Dictyota dimorphosa Ni-Ni-Win &amp; Kawai, 2024</p><p>Japanese name: アミジグサ (amijigusa)</p><p>Type locality: —Reihoku, Amakusa, Kumamoto Prefecture, Japan</p><p>Description and remarks: —Specimens are 3–7 cm tall measured from the small rhizoidal holdfast up to the apices. The specimens show increasing thallus width from the base (3–4 mm) to the upper branches (8–9 mm). Branches are dichotomous with broad and obtuse tips. This species was collected at Yokoura, Naname, and at the old and new ports of Mushima. Specimens here can be considered small and presumably juvenile as their collection was done in the early part of the spring seaweed bloom in Japan (Choi et al. 2002) compared to more mature specimens (Hörnig &amp; Schnetter 1988).</p><p>This species has a very close resemblance with D. dichotoma, a species which has a wide geographic distribution, from the Northern Atlantic (van den Hoek 1982) to the tropics (Titlyanov et al. 2015, Phang et al. 2016, Lastimoso &amp; Santiañez 2021). Its morphological variability has been discussed in the Dictyota dichotoma complex (Hörnig &amp; Schnetter 1988). A study using a molecular approach show that D. dichotoma and D. implexa have a wider distribution unlike other European species (i.e., D. fasciola, D. mediterranea, D. spiralis, D. cyanoloma) which are restricted to the northeastern Atlantic (Tronholm et al. 2010). An earlier paper suggested that two D. dichotoma morphotypes from Korea as being a single species based on molecular analyses despite having differences in anatomical characteristics (Hwang et al. 2005). However, a recent paper suggests that the previously described D. dichotoma from Korea, along with representatives from Japan may actually be a new species, Dictyota dimorphosa (Ni-Ni-Win et al. 2024) based on a combination of morphological and molecular evidence. In our case, we did not include molecular methods to ascertain the identity of our specimens but the gross morphology seem to match those described by Ni-Ni-Win et al. (2024). For example, our specimens have dichotomous branching that extend in a straight direction throughout the length of the branch, while in the field, we also observed a slight iridescence in live samples. Since it is likely that the D. dichotoma in Japan may be based on misapplied names (Tronholm et al. 2010, Ni-Ni-Win et al. 2024), D. dimorphosa is the most plausible species epithet to our samples.</p><p>Materials examined: —18ECS1-33 (1 to 21) and 18ECS1-114-1</p></div>	https://treatment.plazi.org/id/03FBD87AFFA85879FF5AF860FC4AFC4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA75879FF5AFC1CFD71FA0F.text	03FBD87AFFA75879FF5AFC1CFD71FA0F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dictyota spinulosa Hooker f. & Arnott 1838	<div><p>* Dictyota spinulosa Hooker f. &amp; Arnott, 1838</p><p>Japanese name: ハリアミジグサ (hariamijigusa)</p><p>Type locality: —“Loo-Choo” [Okinawa, Ryukyu Retto, Japan]</p><p>Description and remarks: —Specimens are 15–16 cm tall and slender. Thalli are dichotomously branched with spinose margins. The upward-facing spines are denser in the apical branches and are sparse in the middle parts of the thallus, but the lower and basal parts are usually smooth. Branch apices generally have acute angles, appearing pointed and serrated along with the small marginal spines, but gradually become obtuse and rounded when branching.</p><p>This species of Dictyota was collected from sheltered areas in the Mushima old port and is easily distinguishable for its spinose margins especially in branches near the apex. Other Dictyota species which also have dentate structures (i.e., D. ciliolata, D. crenulata, D. crenulata var. canariensis, D. menstrualis, D. jamaicensis, and D. kohlmeyeri) compose the “ Dictyota ciliolata – crenulata complex” but are restricted mostly to the Pacific region of Central America and the Caribbean Sea of the Atlantic (Tronholm et al. 2013). D. spinulosa also occurs in the South China Sea (Phang et al. 2016) and Taiwan (Lewis &amp; Norris 1987)</p><p>Materials examined: —18ECS1-77 (1 to 2)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA75879FF5AFC1CFD71FA0F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA75879FF5AF9DCFDA5F822.text	03FBD87AFFA75879FF5AF9DCFDA5F822.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Distromium decumbens (Okamura) Levring 1940	<div><p>Distromium decumbens (Okamura) Levring, 1940</p><p>Japanese name: フタエオƯギ (futaeougi)</p><p>Basionym: Chlanidote decumbens Okamura, 1899</p><p>Type locality: — Japan (Boshyu, Enoshima, Prov. Hiuga, Prov. Tosa)</p><p>Description and remarks: —The thalli is prostrate, about 9 cm in width. The blades are flabellate and margins are entire with some areas having deep incisions, possibly due to herbivory. This is commonly observed attached to rocky substrate in Aokata where they share the substrate with Dictyota spp. in late spring.</p><p>This species lacks the in-rolled margins (Kitayama 2011) found in other similar species like Padina spp. The specimen exhibits fine filamentous outgrowths from a portion of its margins and is a means of vegetative reproduction for this species (Kajimura 1986).</p><p>Materials examined: —18ECS1-158-1</p></div>	https://treatment.plazi.org/id/03FBD87AFFA75879FF5AF9DCFDA5F822	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA65878FF5AFF58FD67FDA7.text	03FBD87AFFA65878FF5AFF58FD67FDA7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Padina arborescens Holmes 1896	<div><p>Padina arborescens Holmes, 1896</p><p>Japanese name: ƯミƯチワ (umiuchiwa)</p><p>Type locality: —Enoshima, Kanagawa, Japan</p><p>Description and remarks: —Plants are prostrate with thalli 5–10 cm in length and 6–11 cm in width. Algae are attached to the rocky substrate via a rhizomatous holdfast. Blades are fan-shaped to reniform with occasional clefts while calcification is very light or absent. Fresh specimens are light brown but turn dark brown once dried.</p><p>This species is common among the rocky subtidal throughout all the study sites starting from early spring to mid-late summer. They grow densely on any hard substrate together with Colpomenia sinuosa and almost cover the entire stretch of rocks in the shallow areas. In the mid to later parts of summer, most specimens have torn blades and heavy damage due to herbivory.</p><p>Materials examined: —18ECS1-34 (1 to 23)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA65878FF5AFF58FD67FDA7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA65878FF5AFDB4FD71FBFB.text	03FBD87AFFA65878FF5AFDB4FD71FBFB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Padina japonica Yamada 1931	<div><p>* Padina japonica Yamada, 1931</p><p>Japanese name: オキナƯチワ (okinauchiwa)</p><p>Syntype locality: —Various locations in Japan</p><p>Description and remarks: —Thalli are prostrate with flabellate to reniform blades 2–2.5 cm long and 1.5-3.5 cm in width. The thallus is moderately calcified on the superior surface, from the base to the middle regions while the inferior surface is very lightly calcified to almost absent.</p><p>This species grows on the shallow rocky subtidal in moderately exposed habitats at Yokoura among other species of Padina and dense stands of Colpomenia sinuosa and Hydroclathrus clathratus in late spring. This species was previously considered to be conspecific with P. santae-crucis Børgesen (Abbott &amp; Huisman 2003) but was later recognized to be distinct (Ni-Ni-Win et al. 2010) but occupy the same clade (Ni-Ni-Win et al. 2011).</p><p>Materials examined: —18ECS1-35 (1 to 2)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA65878FF5AFDB4FD71FBFB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA65878FF5AFB80FD78FA47.text	03FBD87AFFA65878FF5AFB80FD78FA47.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Padina minor Yamada 1925	<div><p>* Padina minor Yamada, 1925</p><p>Japanese name: ƯスユキƯチワ (usuyukiuchiwa)</p><p>Type locality: —Cape Oluanbi, Taiwan</p><p>Description and remarks: —Thallus is prostrate, 3.5–5 cm in length from the holdfast and 4–7 cm in width. Blades are flabellate to reniform and margins may be entire or appear lobed. Clefts may be present which may make the lobes overlap. Both sides of the thallus are calcified. In fresh specimens the thalli appear light brown but later turn even lighter when dried. This species is common in Yokoura from early-late summer where they are found in association with dense stands of P. Arborescens and H. clathratus .</p><p>Materials examined: —18ECS1-84 (1to 3)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA65878FF5AFB80FD78FA47	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA65878FF5AFA14FD71F807.text	03FBD87AFFA65878FF5AFA14FD71F807.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rugulopteryx okamurae (E. Y. Dawson) I. K. Hwang, W. J. Lee & H. S. Kim 2009	<div><p>* Rugulopteryx okamurae (E.Y. Dawson) I.K. Hwang, W.J. Lee &amp; H.S. Kim, 2009</p><p>Japanese name: フクリンアミジ (fukurin-amiji)</p><p>Basionym: Dilophus okamurae E.Y. Dawson, 1950</p><p>Syntype localities: —Hizen, Kyushu; Iyo, Shikoku; Sagami, Iwaki and Rikuzen, Eastern Honshu</p><p>Description and remarks: —The thalli are slender and upright, 1.5–14 cm tall, attached to the substrate through fine rhizoids and may form dense, bushy, tangled clumps. Branches are dichotomous with obtuse tips. Fresh specimens are light brown on the upper parts and dark brown close to the base and do not exhibit iridescence unlike some species of Dictyota . Some individuals exhibit the stoloniferous fibers from the holdfast (Hwang et al. 2009).</p><p>This species is common in all of the study sites except Aokata and was present year-round. This species is also common along the shallow waters of the western Pacific coasts of Japan, Korea, China, Taiwan and Philippines (Hwang et al. 2009). However, it has been detected in the Mediterranean coast of France and was determined to be inadvertently introduced together with Japanese Oysters ( Crassostrea gigas) used for aquaculture (Verlaque et al. 2009).</p><p>Materials examined: —18ECS1-5 (2 to 17)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA65878FF5AFA14FD71F807	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA5587BFF5AFF58FCA1FDC2.text	03FBD87AFFA5587BFF5AFF58FCA1FDC2.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Spatoglossum latum J. Tanaka 1992	<div><p>* Spatoglossum latum J. Tanaka, 1992</p><p>Japanese name: ヒロハコモングサ (hirohakomongusa)</p><p>Type locality: —Shimoda, Shizuoka, Japan</p><p>Description and remarks: —The thalli are upright, 9–15 cm tall, and attached to the rocky substrate using fine rhizoids. The when fresh, samples are light brown in color but turn darker when dried. The distal parts of the fronds are dichotomously branched, broad and appear to be round to obtuse but the ones found in the specimens examined here are torn or bear bite marks from herbivory. Figures 1 –5 in (Tanaka 1992) shows a good representation of this species and their features. Sori can be seen as black spots scattered across the thallus. This species was common in Yokoura and in Aokata in late spring to early summer.</p><p>Materials examined: —18ECS1-19-1 and 18ECS1-32 (1 to 5)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA5587BFF5AFF58FCA1FDC2	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA5587BFF5AFD8BFD6DFC4F.text	03FBD87AFFA5587BFF5AFD8BFD6DFC4F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Stypopodium zonale (J. V. Lamouroux) Papenfuss 1940	<div><p>Stypopodium zonale (J.V. Lamouroux) Papenfuss, 1940</p><p>Japanese name: ジガミグサ (jigamigusa)</p><p>Type locality: — Dominican Republic</p><p>Description and remarks: —Specimens are upright with thalli measuring 7–14 cm tall and 3–15 cm in width. Plants are attached to the rocky substrates by rhizoidal holdfasts. Younger blades may be flabellate but clefts form as they grow which run the entire length of the thallus, creating strap-shaped segments. Older plants also exhibit torn margins possibly due to herbivory. Underwater, plants exhibit iridescence and are present together with Padina spp. and Dictyopteris spp. in the early-mid summer. This species was collected from Yokoura and at Mushima new port.</p><p>Material examined: —18ECS1-76 (1 to 10)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA5587BFF5AFD8BFD6DFC4F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA5587BFF5AFC1CFE52FA40.text	03FBD87AFFA5587BFF5AFC1CFE52FA40.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Zonaria diesingiana J. Agardh 1841	<div><p>* Zonaria diesingiana J. Agardh, 1841</p><p>Japanese name: シマオƯギ (shimaougi)</p><p>Type locality: —“ad oras Novae Hollandiae” (Probably near Sydney, Australia)</p><p>Description and remarks: —Plants are prostrate with thalli having a 2–4 cm length and 3–5 cm in width and attached to the bottom by fine rhizoidal holdfasts. Fronds grow in a layered structure, with the youngest blades on the surface. The blade margins are entire or sometimes contain clefts which may show the margins as being lobed. In the field, plants are most often found among encrusting and geniculate coralline algae in sheltered areas of Yokoura and Mushima old port and are present from spring until autumn.</p><p>Materials examined: —18ECS1-62 (1 to 8)</p><p>Order Ectocarpales</p><p>Family Chordariaceae</p></div>	https://treatment.plazi.org/id/03FBD87AFFA5587BFF5AFC1CFE52FA40	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA5587AFF5AFA17FE73FF5B.text	03FBD87AFFA5587AFF5AFA17FE73FF5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Myelophycus caespitosus Kjellman (Kjellman 1893	<div><p>Myelophycus caespitosus Kjellman, 1893</p><p>Japanese name: ニシイワヒゲ (nishiiwahige)</p><p>Type locality: —Goto Islands, Nagasaki Prefecture</p><p>Description and remarks: —Specimens are bushy, with several individual terete unbranched thalli arising from a common point. Thalli are often twisted and can reach about 7 cm high with widths of 1–1.5 mm. Plants are common from early-late summer and can be found on the upper intertidal zone at Yokoura and Aokata and are attached to rocks via rhizoidal holdfasts.</p><p>This species was previously synonymized to Myelophycus simplex (Papenfuss 1967) but was recently reinstated as a distinct species upon examining cox1 and cox3 DNA sequences (Hanyuda et al. 2020). Their study shows that the distribution of M. simplex is distinct on the eastern coast of Honshu while M. caespitosus is distributed along the western coast of Honshu, Shikoku and Kyushu.</p><p>Material examined: —18ECS1-51 (1 to 4)</p><p>Family Petrospongiaceae</p></div>	https://treatment.plazi.org/id/03FBD87AFFA5587AFF5AFA17FE73FF5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA4587AFF5AFC8CFD71FB2F.text	03FBD87AFFA4587AFF5AFC8CFD71FB2F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Colpomenia sinuosa (Roth) Derbes & Solier 1851	<div><p>Colpomenia sinuosa (Mertens ex Roth) Derbès &amp; Solier, 1851</p><p>Japanese name: フクロノリ (fukuronori)</p><p>Basionym: Ulva sinuosa Mertens ex Roth, 1806</p><p>Type locality: —“Inter Algas e mari Atlantico prope Gades”; near Cadiz, Spain</p><p>Description and remarks: —Specimens in the collection generally have a saccate and globose shape, with sizes ranging from 2.5–13 cm in width. This species is highly seasonal and was observed across the study sites from March to June, along with Hydroclathrus clathratus .</p><p>Although this species is known to vary widely in its morphology, the appearance of this species in our study sites had smooth to sometimes lobed or convoluted surfaces (Lee et al. 2013, Fig. 1 C and D).</p><p>Materials examined: —18ECS1-6 (2 to 19)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA4587AFF5AFC8CFD71FB2F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA4587AFF5AFB3CFDB0F99F.text	03FBD87AFFA4587AFF5AFB3CFDB0F99F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Dactylosiphon durville	<div><p>* Dactylosiphon durville i (Bory) Santiañez, K.M. Lee, S.M. Boo &amp; Kogame, 2018</p><p>Japanese name: ホソクビワタモ (hosokubiwatamo)</p><p>Basionym: Asperococcus durvillei Bory, 1828</p><p>Type locality: —Concepcion, Chile</p><p>Description and remarks: —The specimen is composed of several tubular, saccate and unbranched thalli with varying sizes arising from a common point. The longest has heights of about 10–15.5 cm while shorter ones are about 0.5–6 cm. This species is common in sheltered areas of Yokoura in early-late spring.</p><p>The specimen bears several bite marks, 5–8 mm wide, along the middle and apices on several of its thalli, indicating it as a food source for herbivorous fish at Yokoura.</p><p>Material examined: —18ECS1-122-1</p></div>	https://treatment.plazi.org/id/03FBD87AFFA4587AFF5AFB3CFDB0F99F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA4587AFF5AFF20FE6DFCFF.text	03FBD87AFFA4587AFF5AFF20FE6DFCFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Petrospongium rugosum (Okamura) Setchell and Gardner 1924	<div><p>* Petrospongium rugosum (Okamura) Setchell and Gardner, 1924</p><p>Japanese name: f シワノカワ (shiwanokawa)</p><p>Basionym: Cylindrocarpus rugosus Okamura, 1903</p><p>Type locality: — Japan</p><p>Description and remarks: —The algal thallus is dark brown, 3–4 mm thick, prostrate and encrusting on hard surfaces using fine rhizoids. The algal surface is rugose with lobed margins and have a leathery or spongy texture. The thallus is composed of four tissue layers, 1.) a central layer composed of layers of cylindrical rhizoidal cells. 2.) rhizoidal layer used for adhering to the substrate. 3.) dichotomously or trichotomously divided subcortical cells that are elongated, cylindrical and are about 30-40 µ thick and 50-80 µ long in young plants but are about 40 µ thick and 150 µ long in older specimens. 4.) a layer of assimilating filaments that are about 5-8 µ thick and 8-10 cells in length and terminated by obtuse cells (Inagaki, 1958).</p><p>This species grows near the intertidal areas in Aokata in the early-mid summer season.</p><p>Materials examined: —18ECS1-157-1</p><p>Family Scytosiphonaceae</p></div>	https://treatment.plazi.org/id/03FBD87AFFA4587AFF5AFF20FE6DFCFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA4587DFF5AF96CFD71FF5B.text	03FBD87AFFA4587DFF5AF96CFD71FF5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Hydroclathrus clathratus (C. Agardh) M. Howe 1920	<div><p>Hydroclathrus clathratus (C. Agardh) M. Howe, 1920</p><p>Japanese name: カゴメノリ (kagomenori)</p><p>Basionym: Encoelium clathratum C. Agardh, 1823</p><p>Lectotype locality: — Belle-île, Brittany, France</p><p>Description and remarks: —Specimens in the collection differ greatly in size, having widths of 5 cm to over 30 cm. This species generally has a net-like, heavily perforated thalli that are flat and spread-out or clumped and convoluted. The holes of the thallus vary in widths of 1 mm to 6 cm in diameter. Larger holes are generally observed to have a dark and thickened borders while smaller holes do not.</p><p>This species is observed to form extensive mats on the bottom from March to June in our study sites. This species also has potential as feedstock for biogas production (Marquez et al. 2014).</p><p>Materials examined: —18ECS1-7 (2 to 16)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA4587DFF5AF96CFD71FF5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA3587DFF5AFF20FDB1FD8A.text	03FBD87AFFA3587DFF5AFF20FDB1FD8A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Petalonia Derbes & Solier 1850	<div><p>* Petalonia fascia (O.F. Müller) Kunzte, 1898</p><p>Japanese name: セイヨƯハバノリ (seiyouhabanori)</p><p>Basionyn: Fucus fascia O.F. Müller, 1778</p><p>Type locality: —Near Kristiansand, Norway</p><p>Description and remarks: —Specimens are delicate and have ribbon-like thalli, 7–8 cm long and tapered at the bottom. Thalli are attached via a small discoid holdfast 0.5 mm wide. Margins are smooth, wavy or lobed and sometimes overlap when pressed. Specimens are yellowish-brown and mucilaginous when fresh and turn light brown when dried and pressed. Samples were collected from a rubble bottom inside the new port in Mushima in early spring.</p><p>Material examined: —18ECS1-115-1</p></div>	https://treatment.plazi.org/id/03FBD87AFFA3587DFF5AFF20FDB1FD8A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA3587DFF5AFB10FC34F9D7.text	03FBD87AFFA3587DFF5AFB10FC34F9D7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum alternato-pinnatum Yamada 1942	<div><p>* Sargassum alternato-pinnatum Yamada, 1942</p><p>Japanese name: キレバモク (kirebamoku)</p><p>Type locality: —Hiuga, Miyazaki Prefecture, Japan (see Yamada, 1942)</p><p>Description and remarks: —This species is distinct for having warty growths on the main axis and branches. The leaves are narrow and elongated, about 2–5 cm long with serrate to dentate margins with scattered cryptostomata. The leaf apices may be lanceolate or forked while leaflets may bifurcate from the leaf margins. The vesicles are round and found on stalks, along the secondary or tertiary branches. This species is common in Aokata, Naname and Yokoura in the spring until mid summer months.</p><p>Materials examined: —18ECS1-124 (1 to 2) and 18ECS1-131 (1 to 10)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA3587DFF5AFB10FC34F9D7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA3587DFF5AF9A4FD71F807.text	03FBD87AFFA3587DFF5AF9A4FD71F807.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum crispifolium Yamada 1931	<div><p>* Sargassum crispifolium Yamada, 1931</p><p>Japanese name: コブクロモク (kobukuromoku)</p><p>Syntype locality: — Various locations across Japan ( Sagami Prov.; Tosa Prov.; Hizen Prov.; Satsuma Prov.; Hiuga Prov.)</p><p>Description and remarks: —Thalli specimens were up to 52 cm tall with smooth axes, 1.5–2 mm in width. The leaves may be crispate or entire and margins may be heavily or lightly dentate with clearly visible midribs and cryptostomata scattered around the leaf surface. The air bladders are about 1.5–2 mm in diameter and found around the primary laterals. This species was found in Yokoura and in Mushima old port in early summer, together with other species of Sargassum .</p><p>Materials examined: —18ECS1-20 (1 to 3)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA3587DFF5AF9A4FD71F807	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA3587DFF5AFD53FEA6FB44.text	03FBD87AFFA3587DFF5AFD53FEA6FB44.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Scytosiphon lomentaria (Lyngbye) Link	<div><p>Scytosiphon lomentaria (Lyngbye) Link, nom. cons. 1833</p><p>Japanese name: カヤモノリ (kayamonori)</p><p>Basionym: Chorda lomentaria Lyngbye</p><p>Syntype localities: —“Habitat in infimo refluxus limite ad littoral Faeroensia, ut a Qvivig, copiosa. Ad Hammerhuus Bornholmiae” [Quivig, Faeroe Islands and Bornholm, Denmark].</p><p>Description and remarks: —The thalli are bushy, with several individual hollow, air-filled and terete unbranched thalli arising from a common point. The thalli have several constrictions along its axis which creates segments that are 3–5 mm long. Thalli can reach about 10–20 cm high with widths of 1–2 mm. Plants were found on the upper intertidal to shallow zone and attached to rocks via rhizoidal holdfasts. This species is common in early to late spring across Naname and Yokoura.</p><p>Materials examined: —18ECS1-162 (1 to 3)</p><p>Order Fucales</p><p>Family Sargassaceae</p></div>	https://treatment.plazi.org/id/03FBD87AFFA3587DFF5AFD53FEA6FB44	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA2587CFF5AFF58FD71FD6F.text	03FBD87AFFA2587CFF5AFF58FD71FD6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum fusiforme (Harvey) Setchell 1931	<div><p>Sargassum fusiforme (Harvey) Setchell, 1931</p><p>Japanese name: ヒジキ (hijiki)</p><p>Basionym: Cystophyllum fusiforme Harvey, 1860</p><p>Type locality: —Shimoda, Shizuoka, Japan</p><p>Description and remarks: —Specimens in the collection are fleshy with a 2–2.5 mm diameter main axis. The leaves are modified into cylindrical phylloids of various lengths, 1.5–3.5 mm long. The vesicles are fusiform with mucronate or attenuate tips and are attached to short stalks, 1–1.5 cm in length. Plants were collected from rocky outcrops on the exposed portion of Yokoura. This species was found at Naname, in the shallow subtidal areas but the plant heights are small compared to the ones at Yokoura.</p><p>This species is widely consumed as food in China, Japan and Korea and is known for being a potential source of therapeutic compounds that are of pharmaceutical importance (Liu et al. 2020). However, people are cautioned on eating S. fusiforme as it is also known to contain inorganic arsenic (Lee et al. 2022).</p><p>Materials examined: —18ECS1-49 (1 to 8)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA2587CFF5AFF58FD71FD6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA2587CFF5AFD7CFD67FBDF.text	03FBD87AFFA2587CFF5AFD7CFD67FBDF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum hemiphyllum (Turner) C. Agardh (Agardh 1821	<div><p>Sargassum hemiphyllum (Turner) C. Agardh, 1820</p><p>Japanese name: イソモク (isomoku)</p><p>Basionym: Fucus hemiphyllus Turner, 1811</p><p>Type locality: — Nagasaki, Japan</p><p>Description and remarks: —Specimen thalli are bushy. The leaves are easily distinguishable for this species as they are coarsely dentate on the outer side while being smooth and entire on the inward side. The vesicles are borne on stalks and may be pyriform, ovoid or obovoid in shape. The vesicle apices may either be round, mucronate or bear a small leaflet, especially in the upper parts of the thalli. This species forms thick stands along the windward side of Yokoura from early spring to early summer.</p><p>Materials examined: —18ECS1-47 (1 to 14)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA2587CFF5AFD7CFD67FBDF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA2587CFF5AFBACFD67F90B.text	03FBD87AFFA2587CFF5AFBACFD67F90B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum horneri (Turner) C. Agardh 1820	<div><p>Sargassum horneri (Turner) C. Agardh, 1820</p><p>Japanese name: アカモク (akamoku)</p><p>Basionym: Fucus horneri Turner</p><p>Type locality: —In the Straits of Corea</p><p>Description and remarks: —Specimens are densely branched with a thicker main axis at the bottom (2.5–3 mm) and thinner at the top (1 mm). The main axis and the primary branches in the lower portions have small spines while the upper parts are smooth. Branching is alternately bipinnate with the leaves having a prominent midrib with deeply incised and alternating margins. The vesicles are 1–1.5 cm long, cylindrical and are found in the apical regions. The apex of each vesicle is terminated by a leaflet with thin, alternating, deeply pinnatisect margins. In Naname, this species forms dense forests grows to heights reaching over 3–5 m in early to late spring.</p><p>This species is known for producing large floating rafts in the East China Sea region during macroalgal blooms, known as “golden tide”, and is considered as a nuisance when they drift ashore (Zhuang et al. 2021, Qi et al. 2022). In 2015, massive rafts of drifting S. horneri from China were recorded to have drifted ashore in many parts of the Goto Islands (Naru, Fukue, Kamigoto), mainland Nagasaki and to the islands in Iki and Tsushima (Kiriyama et al. 2016). Despite this problem, these floating rafts provide refuge for juvenile pelagic species such as yellowtail ( Seriola quinqueradiata), an important fishery species (Mizuno et al. 2014).</p><p>Materials examined: —18ECS1-125 (1 to 7)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA2587CFF5AFBACFD67F90B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA2587FFF5AF8D0FD67FEC7.text	03FBD87AFFA2587FFF5AF8D0FD67FEC7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum ilicifolium (Turner) C. Agardh 1820	<div><p>* Sargassum ilicifolium (Turner) C. Agardh, 1820</p><p>Japanese name: ヒイラギモク (hiiragimoku)</p><p>Basionym: Fucus ilicifolius Turner</p><p>Type locality: —Sunda Strait, Indonesia</p><p>Description and remarks: —The specimen is small with a height of 6 cm. Leaves may be 1.5 cm long and 1 cm in width, with dentate margins. The leaves may sometimes bear duplicated leaf margins (Liao et al., 2013) where the leaf folds along one third of one side of the leaf or along the axis of the faint midrib. This results in one side being smooth or bending slightly inward creating a crescent margin, similar to Sargassum hemiphyllum leaves. This species was observed growing in small patches along the shallow windward side of Yokoura in mid spring until mid summer, together with other species of Sargassum, namely S. muticum and S. micracanthum .</p><p>Materials examined: —18ECS1-130 (1 to 2)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA2587FFF5AF8D0FD67FEC7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA1587FFF5AFE94FD43FCFF.text	03FBD87AFFA1587FFF5AFE94FD43FCFF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum macrocarpum C. Agardh 1820	<div><p>Sargassum macrocarpum C. Agardh, 1820</p><p>Japanese name: ノコギリモク (nokogirimoku)</p><p>Type locality: —ex Japonia</p><p>Description and remarks: —Plants are bushy and reaching up to more than 30 cm tall. The main axis has a diameter of about 2 mm, some sections are slightly compressed and winged, bearing serrate margins. Leaves occur on the secondary branches and have deeply incised serrate margins with a faint midrib. Vesicles are found on the base of the tertiary branches, having a spherical, elliptical or obovoid shape with a leaflet on apex. This species of dioecious and the reproductive parts are found on the terminal branches around the month of June until September.</p><p>This is a perennial species and is widely distributed around Yokoura, forming dense algal forests. From 2019, this species gradually declined until completely disappearing by the end of 2020.</p><p>Materials examined: —18ECS1-2 (2 to 8)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA1587FFF5AFE94FD43FCFF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA1587FFF5AFC8CFD7CFB13.text	03FBD87AFFA1587FFF5AFC8CFD7CFB13.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum micracanthum (Kutzing) Endlicher 1843	<div><p>Sargassum micracanthum (Kützing) Endlicher, 1843</p><p>Japanese name: トゲモク (togemoku)</p><p>Basionym: Halochloa micracantha Kützing, 1843</p><p>Type locality: — Japan</p><p>Description and remarks: —Specimens are bushy, reaching up to 25 cm tall and have robust discoid holdfasts up to 1 cm in diameter. The main axis is cylindrical with a diameter of 1.5–2 mm. Leaves grow directly from the main axis, pinnatisect, having deeply incised margins with distinct midribs. The vesicles are spherical to ovoid in shape with caudate tips or terminated by a leaflet. Vesicles are found growing densely on the base of leaves close to the terminal portions of the plant. This species was found along the windward side of Yokoura from early-mid summer, together with S. muticum and S. ilicifolium .</p><p>Material examined: —18ECS1-48 (1 to 8)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA1587FFF5AFC8CFD7CFB13	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA1587FFF5AFAD8FDB0F927.text	03FBD87AFFA1587FFF5AFAD8FDB0F927.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum muticum (Yendo) Fensholt 1955	<div><p>* Sargassum muticum (Yendo) Fensholt, 1955</p><p>Japanese name: タマハハキモク (tamahahakimoku)</p><p>Basionym: Sargassum kjellmanianum f. muticum Yendo, 1955</p><p>Type locality: —Izumo Prov., Kii, Honshu, Japan</p><p>Description and remarks: —Plants can grow to more than 50 cm tall. The main axis is about 1.5–2 mm in diameter and bipinnately branched. Leaves located on the primary branches and are about 1.4 cm long and slender, not more than 2 mm wide with either smooth or serrate margins. Vesicles are 1.5 mm wide, avoid to elliptical with either smooth apices or with slender leaflets. Branches are longer closer to the base and shorter near the apex of the plant. This species is common along the windward side of Yokoura in the mid spring months and is found together with other species of Sargassum like S. ilicifolium and S. micracanthum .</p><p>Material examined: —18ECS1-126-1</p></div>	https://treatment.plazi.org/id/03FBD87AFFA1587FFF5AFAD8FDB0F927	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA1587EFF5AF934FD71FF5B.text	03FBD87AFFA1587EFF5AF934FD71FF5B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum piluliferum (Turner) C. Agardh 1820	<div><p>Sargassum piluliferum (Turner) C. Agardh, 1820</p><p>Japanese name: マメタワラ (mametawara)</p><p>Basionym: Fucus pilulifer Turner, 1808</p><p>Type locality: —“Harbour at Nagasaki ”, Nagasaki, Japan</p><p>Description and remarks: —Specimens have one or multiple axes growing from a discoid holdfast. The primary and secondary branches are compressed with leaves that are also compressed with faint midribs, sometimes with bifurcate tips. The vesicles are globose to obovoid, attached to stalks 4–5 mm long. This species is common in mid-late spring along sheltered areas of Yokoura and in Naname.</p><p>Materials examined: —18ECS1-50 (1 to 6)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA1587EFF5AF934FD71FF5B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA0587EFF5AFF20FD8CFD6F.text	03FBD87AFFA0587EFF5AFF20FD8CFD6F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum siliquastrum (Turner) C. Agardh 1820	<div><p>Sargassum siliquastrum (Mertens ex Turner) C. Agardh, 1820</p><p>Japanese name: ヨレモク (yoremoku)</p><p>Type locality: — Nagasaki Harbor, Nagasaki, Japan</p><p>Description and remarks: —The specimen has a robust discoid holdfast with two branches arising from it. The main axis is 2.5 mm wide and has three angles on its cross-section (i.e., triquetrous). The leaves are dense around the upper parts of the thallus and can reach lengths of up to 7 cm, with widths of about 2–3 mm and have serrate or dentate margins. The midribs are also distinct on the leaves. The vesicles are located on upper, secondary branches and are borne on stalks, 3–5 mm long. The vesicles are avoid and may be observed with leaflets on their apices. This species is common in Yokoura in mid spring and has been found along the exposed and shallow rocky areas near the wave breaker.</p><p>Material examined: —18ECS1-3-2</p></div>	https://treatment.plazi.org/id/03FBD87AFFA0587EFF5AFF20FD8CFD6F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA0587EFF5AFD7CFCD9FB67.text	03FBD87AFFA0587EFF5AFD7CFCD9FB67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum thunbergii (Roth) Kuntze 1880	<div><p>Sargassum thunbergii (Mertens ex Roth) Kuntze, 1880</p><p>Japanese name: Ưミトラノオ (umitoranoo)</p><p>Basionym: Fucus thunbergii Mertens ex Roth, 1800</p><p>Syntype locality: — China and Japan seas</p><p>Description and remarks: —The thalli of the specimens are bushy and can reach to about 45–50 cm high. The main axis is terete with a diameter of about 1–1.5 mm. The phylloids are slender and filiform, 3–5 mm long and 0.5–1 mm wide with smooth or dentate margins. The vesicles are on stalks, ovate or fusiform with mucronate tips. Plants are dioecious and may form branches which contain the terete receptacles that are 5–10 mm long. Branches may arise from the main axis and branch lengths are usually longer near the base and shorter near the apex. This species forms a thick forests in Yokoura along the sheltered side of the small embayment on the rocky patches near the sandy areas in mid-late spring. In Naname, it was found along the periphery of the S. horneri forests in early spring.</p><p>Materials examined: —18ECS1-4 (2, 3 and 10–14)</p></div>	https://treatment.plazi.org/id/03FBD87AFFA0587EFF5AFD7CFCD9FB67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA0587EFF5AFB74FE83F998.text	03FBD87AFFA0587EFF5AFB74FE83F998.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sargassum yendoi Okamura and Yamada 1938	<div><p>* Sargassum yendoi Okamura and Yamada, 1938</p><p>Japanese name: エンドƯモク (endoumoku)</p><p>Type locality: —Enoshima, Shimoda or Hayama, Japan</p><p>Description and remarks: —The specimen is 25 cm tall, with a winged main axis, 2.5 mm wide. The leaves are elongated, 5–7 cm long, 8 mm wide with distinct midribs and are alternating on the main axis. The vesicles are globose and on 3–5 mm stalks. The receptacles are densely branched and highly conspicuous in mature individuals. This species forms sparse patches along the rocky subtidal areas close to the mouth of the embayment in Yokoura in mid spring.</p><p>Material examined: —18ECS1-129-1</p><p>Order Laminariales</p><p>Family Alariaceae</p></div>	https://treatment.plazi.org/id/03FBD87AFFA0587EFF5AFB74FE83F998	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFA05861FF5AF96FFEA7FDC3.text	03FBD87AFFA05861FF5AF96FFEA7FDC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Undaria pinnatifida (Harvey) Suringar 1873	<div><p>Undaria pinnatifida (Harvey) Suringar, 1873</p><p>Japanese name: ワカメ (wakame)</p><p>Basionym: Alaria pinnatifida Harvey, 1860</p><p>Type locality: —Shimoda, Shizuoka Prefecture, Japan</p><p>Description and remarks: —Specimen is 32 cm tall with the blade attached on a 2.5 cm long stipe, 2 mm in diameter. The blade contains a midrib that connects to the stipe. The blade margins are deeply pinnate at the base but are shallower closer to the apex. The specimens collected are young and do not yet have the reproductive sporophyll structures at the base. This species is ubiquitous in early to mid spring along the study sites, especially in Naname where they form understory forests below the S. horneri canopy. However, this species has gradually declined in Yokoura together with other canopy-forming species like Sargassum macrocarpum and Ecklonia cava subsp. kurome .</p><p>This species is indigenous to the northwest Pacific coast and grows along the shallow rocky subtidal zone and is also cultivated in Japan, South Korea and China for food (Wu &amp; Meng 1997, Tanaka et al. 2020). In recent decades, its appearance along the coasts of north and south America, Europe and the southern hemisphere was due to accidental introduction probably through shipping or with the introduction of cultured Pacific oysters (Hay &amp; Luckens 1987, Floc’h et al. 1991, Casas &amp; Piriz 1996, Casas 2020). In the Atlantic, U. pinnatifida is highly invasive and rapidly expanding its range, outcompeting native species and causes fouling of submerged structures (Minchin &amp; Nunn 2014). Strategies have been formulated towards controlling its spread and local eradication (Marine Pest Sectoral Committee 2015, Gnanalingam &amp; Hepburn 2019) but total extermination may be futile.</p><p>Materials examined: —18ECS1-119 (1, 3, 4)</p><p>Family Lessoniaceae</p></div>	https://treatment.plazi.org/id/03FBD87AFFA05861FF5AF96FFEA7FDC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFBF5861FF5AFD88FE4FFB26.text	03FBD87AFFBF5861FF5AFD88FE4FFB26.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ecklonia cava subsp. kurome (Okamura)	<div><p>Ecklonia cava subsp. kurome (Okamura) Akita, Hashimoto, Hanyuda, &amp; Kawai, 2020</p><p>Japanese name: クロメ (kurome)</p><p>Basionym: Ecklonia kurome Okamura, 1927</p><p>Lectotype locality: — Kanayama, Wakayama Prefecture, Japan</p><p>Description and remarks: —Thalli of specimens are 12–26 cm tall, with blades attached to cylindrical stipes which have a length of 1–2.5 cm and 1.5–3 cm in diameter. Holdfasts are rhizomatous and attached to rocky substrates. Half of the specimens have wrinkled blades while the others are smooth. All specimens were collected in May to July 2018, from boulders 1–2 m deep, in a semi-sheltered part of Yokoura. This species has not been observed at Yokoura since 2019. However, this species still persists in Naname and has been collected even until early-mid summer.</p><p>This species was previously known as E. kurome until the change was proposed based on multigene molecular phylogeny (Akita et al. 2020). Their study revealed that E. cava, E. kurome and E. stolonifera are all similar morphospecies and are genetically similar and therefore recognized to be subspecies to E. cava . The presence of wrinkled or smooth blades, previously used as distinguishing features separating E. cava and E. kurome was found to be unreliable and possibly caused by environmental conditions or plant age (Arai et al. 1997).</p><p>Materials examined: —18ECS1-36 (1 to 14)</p><p>Order Sporochnales</p><p>Family Sporochnaceae</p></div>	https://treatment.plazi.org/id/03FBD87AFFBF5861FF5AFD88FE4FFB26	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
03FBD87AFFBF5861FF5AFB2BFD67F98B.text	03FBD87AFFBF5861FF5AFB2BFD67F98B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sporochnus radiciformis (R. Brown ex Turner) C. Agardh 1817	<div><p>* Sporochnus radiciformis (R. Brown ex Turner) C. Agardh, 1817</p><p>Japanese name: ケヤリ (keyari)</p><p>Basionym: Fucus radiciformis R. Brown ex Turner</p><p>Type locality: — Australia, probably near Sydney</p><p>Description and remarks: —Specimens are tall and can reach up to 50 cm in height with stiff and terete main axis about 0.5–1 mm in width. Branches are pinnate with alternating branchlets that are terminated by ovate to globular receptacles with tufts of fine hair. Brownish when fresh but turns light green when dried and pressed. Specimens were found attached to rock substrate via a small discoid holdfast, 1 mm in width. This species was found growing along the sandy areas in the middle of Yokoura, especially in mid spring.</p><p>Materials examined: —18ECS1-150 (1 to 2)</p></div>	https://treatment.plazi.org/id/03FBD87AFFBF5861FF5AFB2BFD67F98B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Belleza, Dominic Franco C.;Tateishi, Hiroto;Tanimae, Shin-Ichiro;Kobayashi, Taishun;Aota, Tomoyuki;Urasawa, Toshiki;Nishihara, Gregory N.	Belleza, Dominic Franco C., Tateishi, Hiroto, Tanimae, Shin-Ichiro, Kobayashi, Taishun, Aota, Tomoyuki, Urasawa, Toshiki, Nishihara, Gregory N. (2025): An annotated list of brown seaweeds (Phaeophyceae) from Nakadori Island (northern Goto Islands, Nagasaki, Japan). Phytotaxa 694 (1): 1-24, DOI: 10.11646/phytotaxa.694.1.1, URL: https://doi.org/10.11646/phytotaxa.694.1.1
