identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03F8C565801DA006A7A1FB57FC5DF8BA.text	03F8C565801DA006A7A1FB57FC5DF8BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goodeniacarus Beard & Seeman 2025	<div><p>Goodeniacarus gen. nov. Beard &amp; Seeman</p><p>Type species: Goodeniacarus rostulatus sp. nov. Beard &amp; Seeman</p><p>Diagnosis. Female. Palps 3-segmented; palp tarsus with 2 terminal phaneres (1 long and 1 minute). Anterior margin of prodorsum with small median forked projection forming a notch; dorsal opisthosoma with 7 pairs of setae (c1, c3, d1, d3, e3, f3, h1 present; c2, d2, e1, e2, f2, h2 absent); dorsal setae short, fine with blunt rounded tips; 1–3 pairs of large pores present on opisthosoma (often with an associated minute pore visible, but difficult to see). Ventral and genital shields not developed, both regions membranous and merged together; 2 pairs of ps setae, anal valves membranous, ps2 and ps3 inserted longitudinally on anal valves; setae 4a elongate fine; lacking supernumerary coxal setae 3a2 and 4a2. Setal formulae for legs I – IV: cx 2-2-1-1, tr 1-1-2-1, fe 4-4-2-1/2, ge 3-3-0-0, ti 5-5-3-3, ta 9(1)-9(1)-5-5 respectively. Trochanter III with seta l′ blunt, delayed to adult. Femur IV with seta d present or absent. Setae on genua I–II all added in deutonymph. Tarsus I–II with seta tcʺ delayed to adult. Femora I–IV, genua I–II, tibiae I–IV usually with either or both setae d and l″ blunt. Tarsal claws and empodia pad-like.</p><p>Remarks. We cannot determine with certainty if the palp phaneres are eupathidia and/or setae. This genus has a thick microplate layer that persists intact even after mounting. This layer often obscures features, especially the minute dorsal pores. The ultrafine detail of this layer can be seen in the images taken using low temperature scanning electron microscopy (LTSEM) in Figures 6, 9, 11, 14, 31.</p></div>	https://treatment.plazi.org/id/03F8C565801DA006A7A1FB57FC5DF8BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801DA006A7A1FC05FD8CFBF6.text	03F8C565801DA006A7A1FC05FD8CFBF6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tenuipalpidae Berlese	<div><p>Family Tenuipalpidae Berlese</p><p>Type genus— Tenuipalpus Donnadieu, 1875</p></div>	https://treatment.plazi.org/id/03F8C565801DA006A7A1FC05FD8CFBF6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.text	03F8C565801EA003A7A1FF65FEADF9BD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goodeniacarus leiperi Beard & Seeman 2025	<div><p>Goodeniacarus leiperi sp. nov. Beard &amp; Seeman</p><p>(Figs 1–4)</p><p>Types. Holotype. Female, Australia, Ranger Road, approx. 5 km S of Warwick along New England Highway, Queensland, 28°16’50.1”S 151°58’31.5”E, ex. Goodenia rotundifolia R.Br. ( Goodeniaceae), 25.iii.2016, J.J. Beard and P.I. Forster . Paratypes. 5 females, same data as holotype. All specimens deposited in QM .</p><p>Diagnosis. Female. Dorsal ornamentation comprising plicae forming irregular fine reticulation and folds; dorsal setae length 5–8. Setae g1 positioned slightly posterior to g2. Setae ps2 and ps3 blunt, lengths ps2 5–6, ps3 5–6. Femora I–II with seta bv″ long, tapered; femora IV with 1 seta (ev′ present, d absent); tarsus I–II with companion seta ftʺ slightly longer than solenidion ωʺ. Blunt setae on femora and tibiae: femur I–III d; genu I–II d, l″; tibia I–II l″; tibia III–IV d. Blunt leg setae can have 1–2 small barbs (can be difficult to see).</p><p>Description</p><p>Adult female. (6 measured)</p><p>Dorsum. (Figs 1, 2) Body measurements: v2–h1 192–209 [209], sc2–sc2 122–133 [132], c3–c3 126–141 [134], f3–f3 74–88 [88]. Prodorsal and opisthosomal shields weakly developed. Prodorsum with fine irregular plicae forming irregular reticulation and folds; with shallow sublateral furrows; anterior margin with small median forked projection forming a notch (5–10) [5–7]; with a pair of minute pores sublaterally (can be difficult to see) and discrete pore-like invagination mesad sc2 (indicated by arrows in Fig. 1). Opisthosoma cuticle pattern similar to that of prodorsum, with complicated plicae forming fine irregular reticulation, becoming smoother anteriorly between c1–c1 and d1–d1; with transversely plicate band of sclerotised cuticle around opisthosomal shield margin, most apparent when specimen flattened (Fig. 2); with 3 pairs of large pores—just mesad setae c3, mesad d3 and anteromesad e3; with up to 4 pairs minute pores visible (can be difficult to see). Dorsal setae all short, blunt. Setal measurements: v2 6–7 [6–7], sc1 6–8 [6–7], sc2 6–8 [6], c1 6–8 [6–7], c3 5–7 [5–6], d1 5–8 [6], d3 5–6 [5–6], e3 5 –6 [5–6], f3 5–6 [5–6], h1 4–6 [5].</p><p>Infracapitulum. (Figs 1, 3A) Dorsal cuticle appears smooth, ventral cuticle with fine oblique plicae laterally; infracapitular setae m fine (11–13) [12–13]. Palp tarsus phaneres (2–3 and 8–9); femur-genu-tibia with single barbed dorsal seta (13–15) [13–14].</p><p>Venter. (Fig. 3A) Ventral cuticle with various patterns of fine to strong plicae; with fine longitudinal plicae between setae 1b–1a and longitudinal to oblique plicae on coxae I–II; with transverse plicae between setae 1a–3a becoming widely spaced 3a–4a; with small region of broadly U-shaped plicae anterior to 4a–4a; with fine arching plicae demarking anterior margin of genital flap; genital region flanked by widely spaced longitudinal plicae. Setae g1 inserted just posterior to g2 on genital flap; flap with longitudinal to irregular plicae between ag–g1. Most ventral setae short, fine, appear smooth or with 1–2 barbs; except setae 1a and 4a elongate, fine, and often broken; ps2 and ps3 blunt. Setae ps3 often obscured within lateral folds of genital opening; with large, rounded pore in soft cuticle laterad anal opening. Setal measurements: 1a 64–96 [75–86], 1b 18–20 [19], 1c 14–16 [15], 2b 18–21 [18–19], 2c 14–17 [15–16], 3a 13–17 [15], 3b 11–19 [16], 4a 41–56 [56], 4b 15–17 [17], ag 9–12 [10–11], g1 8–12 [8–9], g2 9–13 [9], ps2 5–6 [5], ps3 5–6 [6].</p><p>Spermathecal apparatus. (Fig. 3B) A moderately short membranous duct with a broader bulb midway, and narrower distal portion with a granulate texture, terminating in a small, rounded bulb.</p><p>Legs. (Fig. 4) See genus Diagnosis for full setal formulae for legs I – IV, except femur IV with 1 seta (ev′ present, d absent). See description of type species, G. rosulatus sp. nov., for full leg chaetotaxy. Tarsi I and II each with one abaxial solenidion ωʺ (both 5–6 [5–6, 6]); barbed companion seta ftʺ slightly longer than solenidion (7–9 [8], 8–9 [8], respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 8–9). Solenidion and setae ft′ inserted on small tubercles. Some leg setae blunt, of similar form to dorsal idiosomal setae except can have 1–2 small barbs (barbs can be difficult to see): seta l′ on trochanter III; seta d on femora I– II, d on femora III; d, l″ on genua I–II; l″ on tibiae I– II, and d on tibiae III– IV.</p><p>Colour. These mites were bright red when alive.</p><p>Host. Goodenia rotundifolia R.Br. ( Goodeniaceae).</p><p>Distribution. This mite is so far known only from Warwick in southeast Queensland, Australia.</p><p>Etymology. This species is named for the field botanist and photographer Glenn Leiper, who has helped the senior author collect and identify host plants on many occasions over the years.</p><p>Remarks. Host plant identification made by Queensland Herbarium (BRI voucher: PIF43590). Goodeniacarus leiperi sp. nov. and G. uri sp. nov. were collected from the same species of host plant approximately 160 km apart. These two species can easily be separated using the following characters: G. leiperi sp. nov. dorsum with fine irregular reticulation, seta bv″ on femur I–II finely tapered, setae l′ on femur II and genu II finely tapered vs G. uri sp. nov. dorsum with mostly elongate longitudinal plicae, bv″ on femur I–II short and blunt, setae l′ on femur II and genu II short, blunt.</p></div>	https://treatment.plazi.org/id/03F8C565801EA003A7A1FF65FEADF9BD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658018A017A7A1F998FB2BFAF9.text	03F8C5658018A017A7A1F998FB2BFAF9.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goodeniacarus reticulatus Beard & Seeman 2025	<div><p>Goodeniacarus reticulatus sp. nov. Beard &amp; Seeman</p><p>(Figs 5–15)</p><p>Types. Holotype. Female, Australia, Herberton Range Conservation Park, above Hall’s Falls on North Wondecla Creek, Far North Queensland, 17°24’37.5”S 145°26’55.5”E, ex. Goodenia grandifolia Sims. ( Goodeniaceae), 27.iv.2017, J.J. Beard, P.I. Forster and K. McDonald . Paratypes. 14 females, 7 males, 2 deutonymphs, same data as holotype. All specimens deposited in QM .</p><p>Other material examined. 7 females, 1 male, same data as holotype (QM) .</p><p>Diagnosis. Female. Dorsal ornamentation mostly reticulate; dorsal setae length 6–10. Setae g1 positioned anterior to g2. Setae ps2 and ps3 slender, tapered, lengths ps2 9–11, ps3 8–10. Femora I–II with seta bv″ long, tapered; femora IV with 2 seta (ev′, d present); tarsus I–II with companion seta ftʺ much longer than solenidion ωʺ.</p><p>Blunt setae on femora and tibiae: femur I–IV d; genu I–II d, l″; tibia I d, l″, tibia II l″, tibia III–IV d. Blunt setae smooth.</p><p>Description</p><p>Adult female. (6 measured)</p><p>Dorsum. (Figs 5, 6) Body measurements: v2–h1 208–215 [215], sc2–sc2 132–143 [143], c3–c3 134–150 [150], f3–f3 79–86 [81]. Prodorsal and opisthosomal shields weakly developed. Prodorsum mostly with reticulate cuticle; anterior margin with small median forked projection forming a broad shallow notch (4–7) [7]; with two pairs minute pores visible (can be difficult to see; indicated by arrows in Fig. 5). Opisthosoma similarly with mostly reticulate cuticle, with band of irregular plicae anteriorly; with two pairs large pores—just mesad setae d3 and anterior to e3; with up to four pairs minute pores (can be difficult to see). Dorsal setae all short, fine, blunt. Setal measurements: v2 7–8 [8], sc1 8–10 [9], sc2 8–9 [8–9], c1 7–9 [8], c3 7–8 [7], d1 6–7 [7], d3 7–8 [7], e3 7 –8 [7–8], f3 7–8 [8], h1 6–8 [7–8].</p><p>Infracapitulum. (Figs 5, 7) Dorsal cuticle with fine longitudinal plicae, ventral cuticle appears smooth; infracapitular setae m barbed (12–14) [13]. Palp tarsus phaneres (3 and 8–10) [8–9]; femur-genu-tibia with single barbed dorsal seta (12–17) [12].</p><p>Venter. (Fig. 7) Ventral cuticle with oblique plicae on coxae I–II; with widely spaced transverse plicae between setae 1a–4a; with small region of broadly U-shaped plicae anterior to 4a–4a; with irregular arching plicae surrounding ag setae; with transverse plicae and weakly verrucose cuticle surrounding g1–g2 setae; genital region flanked by widely spaced longitudinal plicae. Setae g1 inserted anterior to g2 on genital flap. Most ventral setae short, fine, apparently smooth; except setae 1a and 4a elongate, fine, and often broken. Setae ps3 often obscured within lateral folds of genital opening. Pair of large, round pores often visible in soft cuticle laterad anus. Setal measurements: 1a 62–88 [68–88], 1b 17–24 [20–22], 1c 19–23 [21], 2b 25–23 [28–32], 2c 26–30 [28], 3a 15–23 [23], 3b 27–34 [34], 4a 40–85 [81], 4b 22–27 [24], ag 14–18 [16], g1 13–16 [13], g2 13–16 [15], ps2 9–11 [9–10], ps3 8–10 [9].</p><p>Spermathecal apparatus. A moderately short membranous duct with a broader bulb midway, and narrower distal portion with a granulate texture, terminating in a small, rounded bulb.</p><p>Legs. (Fig. 8) See genus Diagnosis for full setal formulae for legs I – IV, and description of type species, G. rosulatus sp. nov., for full leg chaetotaxy. Tarsi I and II each with one abaxial solenidion ωʺ (5–6 [5], 4–5 [4], respectively); barbed companion seta ftʺ much longer than solenidion (13–15 [15], 11–14 [12], respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 9–10). Solenidion and setae ft′ inserted on small tubercles. Some leg setae blunt and of similar form to dorsal idiosomal setae—seta l′ on trochanter III; seta d on femora I– IV; d, l″ on genua I– II; d, l″ on tibiae I; l″ on tibiae II; d on tibiae III– IV .</p><p>Microplates. (Figs 6, 9) The microplate layer appears coarsely granular at 1,000 X magnification using DIC, and likewise at 2,000 X using scanning electron microscopy (Fig. 9A). Individual microplates are hexagonal in shape (Figs 9C, D), and have a finely granular surface visible at 20,000 X (Fig. 9D). The microplates on the male (Fig. 11) appear differently to those on the female, however this may simply be due to the image being taken at a different developmental stage to those of the female. The individual plates on the male appear rounded and domed, and interconnected by smaller rectangular microplates; however, the hexagonal structure seen in the female plates is also apparent.</p><p>Adult male. (7 measured).</p><p>Dorsum. (Figs 10, 11) Body measurements: v2–h1 145–154, sc2–sc2 97–105, c3–c3 94–104, f3–f3 52–57. Prodorsum mostly smooth with few wrinkles and plicae, small sublateral region with some weak rugose reticulation indicated; with 3–4 pairs of minute pores visible sublaterally to centrally (can be difficult to see); anterior margin with small median forked projection forming a notch (2–5). Dorsal opisthosoma divided into mesonotal and pygidial shields; mesonotal shield with some weak irregular plicae, captures setal pairs c1–c3 and d1–d3; pygidial shield posterior to D row setae with few arching plicae anterior to mostly smooth central region and oblique plicae laterally. Opisthosoma with 3 pairs of large pores—just mesad setae c3, mesad d3, and anteromesad e3; with up to four pairs minute pores (difficult to see). Dorsal setae as in female, all short, fine, blunt. Setal measurements: v2 7–8, sc1 7–9, sc2 7–10, c1 7–8, c3 6–8, d1 7–8, d3 6–8, e3 7 –8, f3 7–9, h1 7–8.</p><p>Infracapitulum. (Figs 10, 12) Dorsal cuticle with fine longitudinal plicae, ventral cuticle with fine oblique plicae laterally or smooth; infracapitular setae m (8–10). Palp tarsus phaneres (2–3 and 7–8); femur-genu-tibia with single barbed dorsal seta (10–13).</p><p>Venter. (Figs 12, 13, 14) Ventral cuticle with fine longitudinal plicae 1b–1a; with widely spaced transverse plicae from 1a–3a; with region of broadly V-shaped plicae anterior to 4a–4a; with band of fine transverse plicae 4a–ag, and with few weak transverse plicae ag –genital region; genital region can be concealed under fold of cuticle; genital setae and opening flanked by plicae arching around region anteriorly. Setae g1 inserted just anterior to g2 on anterior margin of anogenital opening. Most ventral setae short, fine, apparently smooth or with 1–2 minute barbs (can be seen in Fig. 13); except setae 1a and 4a elongate, fine, and often broken. Setae ps3 modified into short, thick, clavate accessory genital stylets (Fig. 14), often obscured under posterior projections of anogenital opening. Setal measurements: 1a 50–58, 1b 14–18, 1c 14–18, 2b 17–21, 2c 21–23, 3a 13–20, 3b 19–23, 4a 37–60, 4b 15–26, ag 11–13, g1 11–13, g2 12–14, ps2 7–11, ps3 6–7.</p><p>Aedeagus. (Fig. 12) Sclerotised, elongate, subulate (58–64), with narrow membranous tube emerging from sclerotised section.</p><p>Legs. (Fig. 15) Setal formulae and chaetotaxy same as adult female except ta I–II with solenidion ω′ also present. Tarsi I and II each with two solenidia (both with adaxial ω′ 6–8, abaxial ωʺ 5–6); companion seta ftʺ barbed (9–12, 9–10, respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 8–9). Form of leg setae similar to that of female, except setae bv″ is blunt on femur II.</p><p>Deutonymph. (2 measured)</p><p>Dorsum. Body measurements: v2–h1 173–187, sc2–sc2 112–127, c3–c3 118–123, f3–f3 56–60. Prodorsal and opisthosomal shields weakly developed. Prodorsum with smooth cuticle; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with widely spaced broken transverse plicae between c1–d1; posterior d1–d1 with few longitudinal folds centrally to sublaterally. Dorsal setae as in adults. Setal measurements: v2 4–5, sc1 5–6, sc2 5–6, c1 6–7, c3 4–5, d1 4–5, d3 3–5, e3 4 –5, f3 4–5, h1 4.</p><p>Infracapitulum. Cuticle as in adults; infracapitular setae m (7–8). Palp tarsus phaneres (1–2 and 6–7); femur-genu-tibia with single barbed dorsal seta (9–8).</p><p>Venter. Ventral cuticle finely plicate; with fine longitudinal plicae 1b–1a; transverse plicae 1a to just posterior 4a; plicae circling around setae ag with longitudinal plicae inside circle; with band of transverse plicae between ag–ps3. Form of ventral setae as in adults. Setal measurements: 1a 58, 1b 14–18, 1c 7–10, 2b 10–15, 2c 8–11, 3a 8–13, 3b 9–13, 4a 24–34, 4b 7–11, ag 5–8, g1 6–8, ps2 3, ps3 3–4.</p><p>Legs. Setal formulae and chaetotaxy for legs I – IV same as adult female except trochanters 1-1-1-0. Tarsi I and II each with one abaxial solenidion ωʺ (4–5, 4, respectively); barbed companion seta ftʺ (both 5–6); pair of prorals p′ζ, pʺζ (6–7, 5–6, respectively). Some leg setae short to minute, blunt—seta l′ on trochanter III, seta d on femora I–IV, genua I–II, tibiae III–IV, and seta lʺ on genua I–II and tibiae I–II.</p><p>Host. The host plant, Goodenia grandifolia Sims. ( Goodeniaceae), has sticky and fragrant leaves, and the mites were observed in higher numbers on the older, less sticky leaves.</p><p>Etymology. This species is named for its obviously reticulate dorsum.</p><p>Remarks. Host plant identification made by Queensland Herbarium (BRI voucher: PIF45191).</p></div>	https://treatment.plazi.org/id/03F8C5658018A017A7A1F998FB2BFAF9	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.text	03F8C565800CA019A7A1FA5FFD59FE08.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goodeniacarus rosulatus Beard & Seeman 2025	<div><p>Goodeniacarus rosulatus sp. nov. Beard &amp; Seeman</p><p>(Figs 16–25)</p><p>Types. Holotype. Female, Australia, Rungulla National Park, 37. 3 km SSE ranger station and airstrip, Queensland, 19°28’00’’S 143°31’33”E, ex. Goodenia rosulata Domin. ( Goodeniaceae), 08.v.2022, J.J. Beard (QM) . Paratypes. 16 females, 5 males, 4 deutonymphs, 3 protonymphs, 3 larvae, same data as holotype (QM) .</p><p>Other material examined. 6 females, 1 male (pharate), 4 deutonymphs, 1 protonymph, same data as holotype (QM) .</p><p>Diagnosis. Female. Prodorsal ornamentation weaky reticulate, becoming weak and irregular medially and laterally; opisthosomal ornamentation reticulate, becoming weak laterally; dorsal setae length 4–9. Setae g1 positioned just anterior to g2. Setae ps2 and ps3 slender, tapered, lengths ps2 7–10, ps3 5–9 (sometimes with a blunt tip). Femora I with setae bv″ long, tapered to fine tip; femora II with bv″ shorter, blunt; femora IV with 2 seta (ev′, d present; companion seta ftʺ much longer than solenidion ωʺ. Blunt setae present on femora and tibiae: femur I–IV d, femur II bv″; genu I–II d, l″; tibia I–II l″; tibia III–IV d. All blunt setae with minute barbs.</p><p>Description</p><p>Adult female. (14 measured)</p><p>Dorsum. (Fig. 16) Body measurements: v2–h1 187–213 [193], sc2–sc2 121–140 [128], c3–c3 119–139 [128], f3–f3 62–81 [72]. Prodorsal and opisthosomal shields weakly developed. Prodorsum with reticulate cuticle becoming weak and irregular medially and laterally; with shallow longitudinal sublateral furrows; anterior margin with small median forked projection forming a notch (5–10) [8]; with up to 4 pairs of minute pores visible sublaterally. Opisthosoma with stronger reticulation over large central region, becoming weak and irregular medially and laterally; with 3 pairs of large pores—mesad setae c3, mesad d3, and posteromesad d3; with up to 3 pairs minute pores (can be difficult to see). Dorsal setae all short, fine, blunt. Setal measurements: v2 6–8 [7], sc1 5–9 [5–6], sc2 6–9 [6], c1 5–8 [5], c3 5–7 [5], d1 4–7 [5], d3 5–6 [5], e3 5 –7 [5], f3 5–7 [5], h1 5–7 [5].</p><p>Infracapitulum. (Figs 16, 17A–B) Cuticle on dorsal infracapitulum with fine longitudinal plicae; infracapitular setae m (13–18) [14]. Palp tarsus phaneres (2 and 8–9); femur-genu-tibia with single barbed dorsal seta (12–14) (Fig. 17A–B).</p><p>Venter. (Fig. 18) Ventral cuticle with fine longitudinal plicae between setae 1b–1a and longitudinal to oblique on coxae I–II; with widely spaced transverse plicae between setae 1a–3a; with small region of broadly U-shaped plicae anterior to 4a–4a; with fine arching plicae demarking anterior margin of genital flap; genital region flanked by widely spaced longitudinal plicae. Setae g1 inserted just anterior to g2 on genital flap. Most ventral setae short, fine, smooth or minutely barbed (difficult to see); except setae 1a and 4a elongate, fine, and often broken. Setae ps2, ps3 slender, tapered; ps3 often obscured within lateral folds of genital opening, sometimes with minutely blunt tip. Pair of large, round pores often visible in soft cuticle laterad anus. Setal measurements: 1a 64–86 [73–83], 1b 15–29 [20–24], 1c 15–22 [17], 2b 15–23 [18], 2c 18–24 [21], 3a 14–19 [13–15], 3b 18–21 [18], 4a 56–76 [61–66], 4b 13–17 [15], ag 13–21 [21], g1 10–12 [10–11], g2 10–13 [12], ps2 7–10 [7], ps3 5–9 [6].</p><p>Spermathecal apparatus. (Fig. 17C) A moderately short membranous duct with a broader bulb midway, and narrower distal portion with a granulate texture, terminating in a small, rounded vesicle.</p><p>Legs. (Fig. 19) See genus Diagnosis for full leg setal formulae. Chaetotaxy for legs I – IV: cx I 1b, 1c; cx II 2b, 2c; cx III 3c; cx IV 4b; tr I–II, IV v′, tr III l′, v′; fe I–II d, l′, v′, bvʺ; fe III–IV d, ev′; ge I–II d, l′–lʺ, ge III–IV nude; ti I–II d, l′, lʺ, v′, vʺ; ti III–IV d, v′, vʺ; ta I–II ft′, ftʺ, tc′, tc″, p′ζ, pʺζ, u′, uʺ, ωʺ; ta III–IV ft′, tc′, tcʺ, u′, uʺ. Tarsi I and II each with one abaxial solenidion ωʺ (5–7 [5], 4–6 [5], respectively); barbed companion seta ftʺ much longer than solenidion (10–13 [10], 9–12 [10], respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 8–10). Solenidion and setae ft′, ftʺ inserted on small tubercles. Some leg setae blunt and of similar form to dorsal idiosomal setae except with minute barbs—seta l′ on trochanter III, seta d on femora I–IV, bv″ on femur II; seta d on genua I–II, tibiae III–IV; and seta lʺ on genua I–II and tibiae I–II.</p><p>Adult male. (4 measured)</p><p>Dorsum. (Fig. 20) Body measurements: v2–h1 144–148, sc2–sc2 96–104, c3–c3 91–100, f3–f3 47–51. Prodorsum with weakly developed irregular reticulation and folds sublaterally, with smooth central region and becoming smooth laterally; anterior margin of prodorsum with small median forked projection forming a notch (4–7); with at least 2 pairs of minute pores sublaterally (can be difficult to see). Dorsal opisthosoma with pair of weakly developed metanotal shields indicated between setal pairs c1–c3 and d1–d3; and posterior to D row setae with few oblique plicae from e3–f3 to central smooth region. Dorsal opisthosoma divided into mesonotal and pygidial shields; with at least 2 pairs large pores visible—mesad c3, anteromesad e3; with at least 3 pairs minute pores (can be difficult to see). Dorsal setae as in female, all short, fine, blunt. Setal measurements: v2 5–6, sc1 6–7, sc2 6–7, c1 5–8, c3 4–6, d1 5–6, d3 4–6, e3 5 –6, f3 6–7, h1 5–6.</p><p>Infracapitulum. (Fig. 20) Cuticle on dorsal infracapitulum with fine longitudinal plicae; infracapitular setae m (8–10). Palp tarsus phaneres (2 and 6–7); femur-genu-tibia with single barbed dorsal seta (9–11).</p><p>Venter. (Fig. 21) Ventral cuticle with widely spaced transverse plicae from 1a–3a; with small region broadly U-shaped plicae anterior to 4a–4a between coxae III–IV; with band of fine transverse plicae 4a–ag, and with few weak transverse plicae ag –genital setae; genital region with few flanking plicae arching around region anteriorly. Setae g1 inserted just anterior to g2 on anterior margin of anogenital opening. Most ventral setae short, fine, smooth or minutely barbed (difficult to see); except setae 1a and 4a elongate, fine, and often broken. Setae ps3 modified into short, thick, clavate accessory genital stylets, often obscured under posterior projections of anogenital opening. Setal measurements: 1a 48–55, 1b 12–15, 1c 13–14, 2b 15–21, 2c 14–17, 3a 11–13, 3b 14–16, 4a 28–45, 4b 12–15, ag 9–14, g1 9–11, g2 11–12, ps2 6–8, ps3 5–6.</p><p>Aedeagus. (Fig. 21) Sclerotised, elongate, subulate (57–58), with narrow membranous tube emerging from sclerotised section.</p><p>Legs. (Fig. 22) Setal formulae and chaetotaxy same as adult female except ta I–II with solenidion ω′ also present; ω′ inserted somewhat proximally and partially concealed by strong transverse cuticular ridge across segment. Tarsi I and II each with two solenidia (ta I adaxial ω′ 7–9, abaxial ω″ 6–7; ta II adaxial ω′ 7–8, abaxial ω″ 5–6); companion seta ftʺ barbed (9–10, 6–9, respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 7–9). Form of leg setae similar to that of female.</p><p>Deutonymph. (4 measured)</p><p>Dorsum. Body measurements: v2–h1 168–188, sc2–sc2 112–118, c3–c3 117–121, f3–f3 47–51. Prodorsal and opisthosomal shields weakly developed, with smooth cuticle; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with a few broad transverse striae. Dorsal setae form as in adults. Setal measurements: v2 3–4, sc1 3–4, sc2 3–4, c1 3–4, c3 3, d1 2–3, d3 2–3, e3 2 –3, f3 2–3, h1 2–3.</p><p>Infracapitulum. Cuticle as in adults; infracapitular setae m (7–8). Palp tarsus phaneres (1–2 and 5–6); femur-genu-tibia with single barbed dorsal seta (8).</p><p>Venter. (Fig. 23A) Widely spaced longitudinal plicae between 1b–1a, widely spaced transverse plicae 1a to just posterior 4a; plicae forming a triangle around setae ag with transverse plicae within triangle; with band of transverse plicae between ag–g1 and mixed plicae in genital region. Form of ventral setae as in adults. Setal measurements: 1a 55–72, 1b 10–12, 1c 8–10, 2b 10–12, 2c 11–14, 3a 7–10, 3b 7–12, 4a 27–32, 4b 8–12, ag 6–7, g1 4–5, ps2 2–3, ps3 2–4.</p><p>Legs. (Fig. 23B) Setal formulae and chaetotaxy for legs I – IV same as adult female except tarsi 8(1)-8(1)-3-3 (seta tcʺ absent on ta I–II). In addition to G. rosulatus sp. nov., the deutonymph is known for G. reticulatus sp. nov., G. uri sp. nov. and G. wondul sp. nov., and these species all share trochanter III with only v′ present, and the addition of l′ being delayed to the adult stage. It should be noted, however, that seta l′ is inconsistently expressed on tr III in the deutonymph of G. rosulatus sp. nov. Of the 8 individual deutonymphs we examined, 2 had v′ symmetrically expressed (1 seta on both sides), 3had setae v′ and l′ on one side and only v′ on the other(l′ asymmetrically expressed), 3 specimens had only 1 trochanter III clearly visible and 2 of these had 1 seta present and 1 had 2 setae present.As the most common pattern was seta v′ only present, we consider the presence of seta l′ in the deutonymph to be an aberration. Tarsi I and II each with one abaxial solenidion ωʺ (both 3–4); barbed companion seta ftʺ (both 4–6); pair of prorals p′ζ, pʺζ (6–9, 6–7, respectively). Some leg setae short to minute, blunt—seta l′ on trochanter III, seta d on femora I–IV, genua I–II, tibiae III–IV, and seta lʺ on genua I–II and tibiae I–II.</p><p>Protonymph. (2 measured)</p><p>Dorsum. Body measurements: v2–h1 140–144, sc2–sc2 98–99, c3–c3 98–105, f3–f3 35–40. Prodorsal and opisthosomal shields weakly developed; prodorsum mostly smooth, with smoothly rounded anterior margin without median notch; dorsal opisthosoma with few broadly spaced transverse plicae. Dorsal setae form as in adults. Setal measurements: v2 3–4, sc1 3–4, sc2 4, c1 4–5, c3 3–4, d1 3–4, d3 3–4, e3 2 –3, f3 2–3, h1 2–3.</p><p>Infracapitulum. Cuticle as in deutonymph; infracapitular setae m (5–6). Palp tarsus phaneres (1 and 5); femur-genu-tibia with single barbed dorsal seta (7–8).</p><p>Venter. Ventral cuticle and setae as in deutonymph. Setal measurements: 1a 58, 1b 7–10, 1c 8–9, 2c 8–12, 3a 8–10, 3b 8–9, ag 5–7, ps2 2, ps3 2.</p><p>Legs. (Fig. 24) Setal formulae for legs I – IV: cx 2-1-1-0, tr 0-0-0-0, fe 3-3-2-2, ge 0-0-0-0, ti 5-5-3-3, ta 7(1)- 7(1)-3-3 respectively. Leg chaetotaxy: same as larva except 1c, 2c, 3b added to cx I, II, III respectively; and tc′ added to ta I–II. Tarsi I and II each with one abaxial solenidion ωʺ (both 3); barbed companion seta ftʺ (5, 4–5, respectively); pair of prorals p′ζ, pʺζ (both 5–6).</p><p>Larva. (2 measured)</p><p>Dorsum. (Fig. 25) Body measurements: v2–h1 117–122, sc2–sc2 86–108, c3–c3 84–96, f3–f3 27–31. No shields developed. Prodorsum sparsely finely colliculate anteriorly, and few short, irregular plicae posteriorly; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with few widely spaced transverse striae between c1–d1, and short oblique plicae d1–h1. Dorsal setae form as in adults, except much shorter. Setal measurements: v2 3, sc1 3–4, sc2 3–4, c1 3–4, c3 2–3, d1 3, d3 2–3, e3 2 –3, f3 2–3, h1 2–3.</p><p>Infracapitulum. Cuticle with fine transverse plicae; infracapitular setae m absent. Palp tarsus phaneres (1 and 5); femur-genu-tibia with single barbed dorsal seta (5–6).</p><p>Venter. Ventral cuticle and setae as in deutonymph; fine longitudinal plicae 1b–1a; fine transverse plicae from 1a–ps3; region between coxae III–III with finely granulate cuticle with mixed oblique plicae. Setal measurements: 1a 38–39, 1b 8–9, 3a 6–7, ps2 2–3, ps3 2–3.</p><p>Legs. (Fig. 25) Setal formulae for legs I – III: cx 1-0-0, tr 0-0-0, fe 3-3-2, ge 0-0-0, ti 5-5-3, ta 7(1)-7(1)-3 respectively. Leg chaetotaxy: cx I 1b; fe I–II d, v′, bvʺ; fe III d, ev′; ge I–III nude; ti I–II d, l′, lʺ, v′, vʺ; ti III d, v′, vʺ; ta I–II ft′, ftʺ, p′ζ, pʺζ, u′, uʺ, ωʺ; ta III ft′, u′, uʺ. Tarsi I and II each with one abaxial solenidion ωʺ (both 2–3); barbed companion seta ftʺ (4–5, 5, respectively); pair of prorals p′ζ, pʺζ (both 5–6).</p><p>Colour. These mites were bright red when alive.</p><p>Host. Goodenia rosulata Domin ( Goodeniaceae).</p><p>Distribution. This mite is so far known only from Rungulla National Park in northern Queensland, Australia, where it and its host plant were collected at a single site within the park.</p><p>Etymology. This species is named for the specific epithet of the host plant, rosulata .</p><p>Remarks. Host plant identification made by Queensland Herbarium (BRI voucher: RNP41). Atypical ontogeny of leg setae additions: larval setae l′, l″ on genua I–II are delayed to the deutonymph to be added with seta d; seta tcʺ delayed to adult on tarsus I–II (Table 1).</p></div>	https://treatment.plazi.org/id/03F8C565800CA019A7A1FA5FFD59FE08	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.text	03F8C5658002A02EA7A1FE04FEAAF821.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goodeniacarus uri Beard & Seeman 2025	<div><p>Goodeniacarus uri sp. nov. Beard &amp; Seeman</p><p>(Figs 26–36)</p><p>Types. Holotype. Female, Australia, Serpentine Creek Historic Cemetery, Serpentine Cemetery Road, Redland Bay, Queensland, 27°40’10”S 153°18’05”E, ex. Goodenia rotundifolia R.Br. ( Goodeniaceae), 24.ix.2015, J.J. Beard, G. Leiper and P.I. Forster (QM) . Paratypes. 6 females, 2 males, 3 deutonymphs, 2 protonymphs, 2 larvae, same data as holotype (QM) .</p><p>Other material examined. 1 pharate protonymph, same data as holotype (QM) .</p><p>Diagnosis. Female. Dorsal ornamentation comprising elongate longitudinal plicae; dorsal setae length 4–7. Setae g1 positioned in transverse line with g2. Setae ps2 and ps3 blunt, lengths ps2 5–6, ps3 5–6. Femora I–II with seta bv″ fine blunt; femora IV with 1 seta (ev′ present, d absent; companion seta ftʺ slightly longer than solenidion ωʺ. Blunt setae on femora and tibiae: femur I–II d, l′, bv″; femur III d; genu I–II d, l″; tibia I–II d (1–2 barbs), l″; tibia III–IV d. Most blunt setae apparently smooth.</p><p>Description</p><p>Adult female. (7 measured)</p><p>Dorsum. (Figs 26, 27) Body measurements: v2–h1 206–224 [221], sc2–sc2 135–154 [142], c3–c3 133–161 [143], f3–f3 76–87 [84]. Prodorsal and opisthosomal shields weakly developed. Prodorsum with mostly elongate longitudinal plicae narrowly spaced, becoming mostly smooth centrally; anterior margin of prodorsum with small median forked projection forming a notch (8–11) [10]; with up to 4 pairs of minute pores in longitudinal band posterior to v2 (can be difficult to see), and with pore-like invagination just mesad sc2 (indicated by arrows in Fig. 26). Opisthosoma with complicated plicae mostly longitudinal to oblique, but almost forming weak elongate reticulation; with 3 pairs of large pores—mesad setae c3, mesad setae d3 and posteromesad d3; with at least 3 pairs minute pores (can be difficult to see). Dorsal setae all short, blunt. Setal measurements: v2 6–7 [6–7], sc1 6–7 [6–7], sc2 6–7 [6–7], c1 6–7 [6–7], c3 4–7 [4–5], d1 5–6 [-], d3 5–6 [5–6], e3 4 –6 [5–6], f3 4–6 [5–6], h1 4–6 [5].</p><p>Infracapitulum. (Figs 26, 28A) Dorsal cuticle with fine longitudinal plicae laterally, ventral cuticle with fine oblique plicae; infracapitular setae m weakly lanceolate with 1–2 small barbs (11–12) [11]. Palp tarsus phaneres (1–2 and 8–9); femur-genu-tibia with single barbed dorsal seta (13–15) [13–14].</p><p>Venter. (Figs 28A, 29) Ventral cuticle with longitudinal and oblique plicae on coxae I–II; with fine longitudinal plicae between 1b–1a; with widely spaced transverse plicae between setae 1a–3a, becoming finer 3a–ag; with small region broadly U-shaped plicae between coxae III–IV just anterior to 4a–4a; with arching plicae surrounding region of transverse plicae between 4a–ag setae; with sparse weak transverse plicae between ag and g1–g2 setae; genital region flanked by widely spaced longitudinal plicae. Setae g1 and g2 inserted in transverse row posteriorly on genital flap. Most ventral setae short, fine, smooth; except setae 1a and 4a elongate, fine, and often broken; and ps2 and ps3 blunt. Setae ps3 often obscured within lateral folds of genital opening. Setal measurements: 1a 64–96 [75–86], 1b 18–20 [19], 1c 14–16 [15], 2b 18–21 [18–19], 2c 14–17 [15–16], 3a 13–17 [15], 3b 11–19 [16], 4a 41–56 [56], 4b 15–17 [17], ag 9–12 [10–11], g1 8–12 [8–9], g2 9–13 [9], ps2 5–6 [5], ps3 5–6 [6].</p><p>Spermathecal apparatus. (Fig. 28B) A moderately short membranous duct slightly broadened midway, with a narrower distal portion, terminating in a small rounded vesicle with granular appearance.</p><p>Legs. (Fig. 30) See genus Diagnosis for full setal formulae for legs I – IV, except femur IV with 1 seta (ev′ present, d absent). See description of type species, G. rosulatus sp. nov., for full leg chaetotaxy. Tarsi I and II each with one abaxial solenidion ωʺ (both 5–6 [5–6, 6]); barbed companion seta ftʺ (7–9 [8], 8–9 [8], respectively) slightly longer than solenidion; pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 8–9). Solenidia inserted on small tubercles. Some leg setae blunt and of similar form to the dorsal idiosomal setae—seta l′ on trochanter III; setae d, l′, bv″ on femora I– II, d on femora III; setae d, l″ on genua I–II; setae d, l″ on tibiae I– II and d on tibiae III– IV.</p><p>Microplates. (Fig. 31) Individual microplates are hexagonal in shape (Figs 31A, B), and have a mostly smooth surface with a few short parallel ridges visible at 10,000 X (Fig. 31B).</p><p>Adult male. (2 measured)</p><p>Dorsum. (Figs 32, 33) Body measurements: v2–h1 153–156, sc2–sc2 95–111, c3–c3 97–102, f3–f3 55–57. Prodorsum similar to female, with mostly longitudinal to oblique plicae narrowly spaced, becoming mostly smooth centrally; anterior margin of prodorsum with small median forked projection forming a notch (4–6); with up to 4 pairs of minute pores sublaterally (can be difficult to see). Dorsal opisthosoma divided into mesonotal and pygidial shields; metanotal shield captures setal pairs c1–c3 and d1–d3; pygidial shield posterior to D row setae with few arching plicae anterior to smooth central region; with 3 pairs of large pores—just mesad setae c3, mesad d3 and posteromesad d3; with up to 4 pairs of minute pores (can be difficult to see; arrow indicates pores in Fig. 33); with a pair of large deep pore-like invaginations just laterad d1. Dorsal setae as in female, all short, blunt. Setal measurements: v2 5–6, sc1 6–7, sc2 5–6, c1 5–6, c3 5–6, d1 5–6, d3 5, e3 4 –5, f3 5–6, h1 5–6.</p><p>Infracapitulum. (Figs 32, 33, 34) Dorsal cuticle with fine longitudinal plicae, ventral cuticle with fine oblique plicae; infracapitular setae m (7–9) appears smooth. Palp tarsus phaneres (2 and 7–8); femur-genu-tibia with single barbed dorsal seta (8–10).</p><p>Venter. (Fig. 34A, B) Ventral cuticle with widely spaced transverse plicae from 1a–3a; with region of broadly V-shaped plicae between coxae III–IV just anterior to 4a–4a; with band of fine transverse plicae 4a–ag, and with few weak transverse plicae ag –genital region; genital region often concealed under large fold of cuticle (Fig. 34A, B), genital setae and anogenital opening flanked by plicae arching around region anteriorly. Setae g1 inserted just anterior to g2 on anterior margin of anogenital opening (Fig. 34B). Most ventral setae short, fine, appear smooth; except setae 1a and 4a elongate, fine, and often broken; ps2 tapered, blunt. Setae ps3 modified into short, thick, clavate accessory genital stylets, often obscured under posterior projections of anogenital opening. Setal measurements: 1a 68–78, 1b 11–16, 1c 12, 2b 11–15, 2c 11–12, 3a 11–13, 3b 11–12, 4a 31–34, 4b 12–13, ag 6–10, g1 11–13, g2 11, ps2 5–7, ps3 5.</p><p>Aedeagus. (Fig. 34C) Sclerotised, elongate, subulate (62), with narrow membranous tube emerging from sclerotised section.</p><p>Legs. (Fig. 35) Setal formulae and chaetotaxy same as adult female except ta I–II with solenidion ω′ also present. Tarsi I and II each with two solenidia (ta I adaxial ω′ 7–8, abaxial ωʺ 7; ta II adaxial ω′ 8, abaxial ωʺ 6–7); companion seta ftʺ barbed (both 7); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 7–8). Form of leg setae similar to that of female.</p><p>Deutonymph. (3 measured)</p><p>Dorsum. Body measurements: v2–h1 157–188, sc2–sc2 101–114, c3–c3 109–125, f3–f3 47–58. Prodorsal and opisthosomal shields weakly developed, with smooth cuticle; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with a few broad transverse striae. Dorsal setae form as in adults. Setal measurements: v2 3–5, sc1 4–6, sc2 4–5, c1 4–5, c3 3–4, d1 3–5, d3 3–4, e3 3 –5, f3 3–4, h1 3–4.</p><p>Infracapitulum. Cuticle as in adults; infracapitular setae m (7–8). Palp tarsus phaneres (1 and 6–7); femur-genu-tibia with single barbed dorsal seta (9–11).</p><p>Venter. Strong longitudinal plicae between 1b–1a; strong transverse, finely granulate plicae 1a to just posterior 4a; plicae from 4a to ag arching, becoming transverse between ag to g1; with sparse oblique plicae surrounding genital region. Form of ventral setae as in adults. Setal measurements: 1a 58, 1b 14–18, 1c 7–10, 2b 10–15, 2c 8–11, 3a 8–13, 3b 9–13, 4a 24–34, 4b 7–11, ag 5–8, g1 6–8, ps2 3, ps3 3–4.</p><p>Legs. Setal formulae and chaetotaxy for legs I – IV same as adult female except trochanters 1-1-1-0. Tarsi I and II each with one abaxial solenidion ωʺ (4–5, 4, respectively); barbed companion seta ftʺ (both 5–6); pair of prorals p′ζ, pʺζ (6–7, 5–6, respectively). Some leg setae short to minute, blunt—seta d on femora I–IV, genua I–II, tibiae I, III–IV, and seta lʺ on genua I–II and tibiae I–II.</p><p>Protonymph. (2 measured)</p><p>Dorsum. Body measurements: v2–h1 139–143, sc2–sc2 95–100, c3–c3 103–107, f3–f3 40–41. Prodorsum mostly smooth, with smoothly rounded anterior margin without median notch; dorsal opisthosoma with broadly spaced transverse plicae between c1–d1; pygidial region between d1–h1 smooth. Dorsal setae form as in adults. Setal measurements: v2 3–5, sc1 4–5, sc2 4–5, c1 4, c3 3–4, d1 3, d3 3–4, e3 3 –4, f3 3, h1 2–3.</p><p>Infracapitulum. Cuticle as in deutonymph; infracapitular setae m (6–8). Palp tarsus phaneres (1 and 5–6); femur-genu-tibia with single barbed dorsal seta (8).</p><p>Venter. Ventral cuticle with longitudinal plicae 1b–1a; transverse plicae 1a–ps3; small region between coxae IV with arching plicae. Setal measurements: 1a 42–51, 1b 9–10, 1c 9–10, 2c 9–10, 3a 7–11, 3b 8–10, ag 6–7, ps2 1–2, ps3 1–2.</p><p>Legs. Setal formulae for legs I – IV: cx 2-1-1-0, tr 0-0-0-0, fe 3-3-2-1, ge 0-0-0-0, ti 5-5-3-3, ta 7(1)-7(1)-3-3 respectively. Leg chaetotaxy same as larva except 1c, 2c, 3b added to cx I, II, III respectively. Tarsi I and II each with one abaxial solenidion ωʺ (both 3); barbed companion seta ftʺ (both 4–5); pair of prorals p′ζ, pʺζ (both 5).</p><p>Larva. (2 measured)</p><p>Dorsum. (Fig. 36A) Body measurements: v2–h1 107–123, sc2–sc2 76–89, c3–c3 83–92, f3–f3 31–33. Dorsum mostly membranous with fine plicae, no shields developed but central prodorsum with thickened patterned cuticle. Prodorsum colliculate anteriorly and centrally with oblique to longitudinal plicae laterally; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with fine transverse plicae between c1–c1 and d1–d1, and a narrow band of widely spaced transverse striae between setal rows C and D; with longitudinal to oblique plicae d1–h1. Pair of large rounded pores anterior to setae e3. Dorsal setae form as in adults. Setal measurements: v2 4, sc1 4–5, sc2 4–5, c1 4–6, c3 3–4, d1 3–4, d3 3–4, e3 3, f3 3, h1 3.</p><p>Infracapitulum. Cuticle with fine transverse plicae; infracapitular setae m absent. Palp tarsus phaneres (1 and 5); femur-genu-tibia with single barbed dorsal seta (5–6).</p><p>Venter. Ventral cuticle mostly finely plicate. Setal measurements: 1a 29–33, 1b 9–10, 3a 8–9, ps2 1, ps3 1.</p><p>Legs. (Fig. 36B) Setal formulae for legs I – III: cx 1-0-0, tr 0-0-0, fe 3-3-2, ge 0-0-0, ti 5-5-3, ta 7(1)-7(1)-3 respectively. Leg chaetotaxy: cx I 1b; fe I–II d, v′, bvʺ; fe III d, ev′; ge I–III nude; ti I–II d, l′, lʺ, v′, vʺ; ti III d, v′, vʺ; ta I–II ft′, ftʺ, p′ζ, pʺζ, u′, uʺ, ωʺ; ta III ft′, u′, uʺ. Tarsi I and II each with one abaxial solenidion ωʺ (both 3); barbed companion seta ftʺ (5–6, 6–7, respectively); pair of prorals p′ζ, pʺζ (both 5). Some leg setae blunt and similar in form to dorsal setae: d on fe I–III and ti III; d, l″ ti I–II.</p><p>Colour. These mites were bright red when alive.</p><p>Host. Goodenia rotundifolia R.Br. ( Goodeniaceae).</p><p>Distribution. This mite is so far known only from Redland Bay in southeast Queensland, Australia.</p><p>Etymology. This species is posthumously named for our friend and mentor, the late Uri Gerson. Uri contributed enormously to our understanding of the Tenuipalpidae, especially so for the Australian fauna, and it is because of his direct invitation and encouragement that the senior author embarked on her journey with this fabulous family of plant-feeding mites. It was an honour to know and work with him and he is painfully missed.</p><p>Remarks. Host plant identification made by Queensland Herbarium (BRI voucher: PIF43028). Atypical ontogeny of leg setae additions: larval setae l′, l″ on genua I–II are delayed to the deutonymph to be added with seta d; tectal setae tc′, tcʺ delayed to deutonymph on tarsus I–II (Table 1).</p><p>Goodeniacarus uri sp. nov. and G. leiperi sp. nov. are found on the same host plant, but were collected approximately 160 km apart from each other. See Remarks for G. leiperi sp. nov. and the key to species for their diagnostic features.</p></div>	https://treatment.plazi.org/id/03F8C5658002A02EA7A1FE04FEAAF821	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.text	03F8C5658036A036A7A1FF65FD33FC24.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goodeniacarus wondul Beard & Seeman 2025	<div><p>Goodeniacarus wondul sp. nov. Beard &amp; Seeman</p><p>(Figs 37–42)</p><p>Types. Holotype. Female, Australia, Wondul Stock Route, 36 km north of Inglewood, Queensland, 28°06’41’’S 151°04’39”E, ex. Goodenia glabra R. Br. ( Goodeniaceae), 10.xii.2001, J.J. Beard and P.I. Forster (UQIC87363; paratype female on same slide) . Paratypes. 4 females, 4 males, deutonymph, protonymph, same data as holotype (UQIC87365, 87369). All material in QM .</p><p>Diagnosis. Female. Dorsal ornamentation fine, irregular reticulation, becoming weaker and smoother on medial prodorsum and anteromedial opisthosoma; dorsal setae length 5–9. Setae g 1 in transverse line with g2, or slightly posterior to g2. Setae ps2 and ps3 short, ps2 tapered, ps3 tapered blunt, lengths ps2 6–7, ps3 6–8. Femora I with seta bv″ long, tapered; femur II with seta bv″ shorter, tapered blunt; femora IV with 1 seta (ev′ present, d absent; companion seta ftʺ longer than solenidion ωʺ. Blunt setae on legs: seta l′ on trochanter III; femur I–III d; genu I–II d, l″; tibia I l″; tibia IV d. Additional ambiguously blunt setae: tibia II d, l″; tibia III d. Blunt leg setae can have 1–2 minute barbs.</p><p>Description</p><p>Adult female. (5 measured)</p><p>Dorsum. (Fig. 37) Body measurements: v2–h1 190–200 [194], sc2–sc2 111–117 [113], c3–c3 112–123 [112], f3–f3 66–75 [70]. Prodorsal and opisthosomal shields weakly developed. Prodorsum with fine irregular reticulation and plicae becoming smoother centrally; anterior margin of prodorsum with small median forked projection forming a notch (5–8); with at least single pair minute pores mesad eyes (can be difficult to see). Opisthosoma with fine irregular reticulation; with broad band of sclerotised cuticle around opisthosomal shield margin, with series of strong transverse plicae; with 3 pairs of large pores—just mesad seta c3, mesad d3, and posteromesad d3; with at least 3 pairs minute pores (can be difficult to see). Dorsal setae all short, fine, smooth, blunt (n.b. one female with c1, c3, d3 tapered blunt and apparently with 1–2 barbs). Setal measurements: v2 6–7 [7], sc1 7–9 [8–9], sc2 8–9 [8], c1 7–9 [7], c3 6–8 [6–7], d1 6–8 [6], d3 7–8 [7], e3 5 –8 [6–7], f3 6–8 [6], h1 6–7 [6].</p><p>Infracapitulum. (Fig. 37) Dorsal cuticle with fine longitudinal to oblique plicae; ventral cuticle smooth centrally with fine oblique striae laterally; infracapitular setae m (13–14) [13]. Palp tarsus phaneres (1–2 and 8–9); femur-genu-tibia with single barbed dorsal seta (13–15) [14].</p><p>Venter. (Fig. 38) Ventral cuticle with few fine longitudinal plicae between setae 1b–1a and longitudinal to oblique plicae on coxae I–II; with transverse plicae between setae 1a to just posterior to 3a; with region of sparse broadly U-shaped plicae anterior to 4a–4a; with fine arching plicae between 4a–ag; genital region flanked by widely spaced longitudinal plicae; genital flap with irregular plicae between ag and genital setae. Setae g1 and g2 inserted in almost transverse row along posterior margin of genital flap. Most ventral setae short, fine, smooth, except setae 1a, 4a elongate, fine (often broken). Setal measurements: 1a 68–81 [68–77], 1b 20–27 [21], 1c 15–18 [15], 2b 20–23 [21], 2c 17–19 [17], 3a 17–22 [17], 3b 17–18 [17], 4a 64–67 [-], 4b 19–21 [21], ag 10–14 [12–13], g1 10–13 [10], g2 11–13 [12], ps2 6–7 [6], ps3 6–8 [8].</p><p>Spermathecal apparatus. A moderately short membranous duct terminating in small bulb; basal half much broader than distal half, with a slightly enlarged bulb midway, and granulate texture basally.</p><p>Legs. (Fig. 39) See genus Diagnosis for full setal formulae for legs I – IV, except femur IV with 1 seta (ev′ present, d absent). See description of type species, G. rosulatus sp. nov., for full leg chaetotaxy. Tarsi I and II each with one abaxial solenidion ωʺ (6–7 [6], 6–7 [6], respectively); barbed companion seta ftʺ (10–11 [11], 10–12 [11], respectively) longer than solenidion; pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 9–10). Solenidia inserted on small tubercles. Some leg setae blunt and of similar form to dorsal idiosomal setae (except with minute barbs)—seta l′ on trochanter III; seta d on femora I– III; d, l″ on genua I–II; seta l″ on tibiae I– II and d on tibiae III– IV.</p><p>Adult male. (4 measured)</p><p>Dorsum. (Fig. 40) Body measurements: v2–h1 128–137, sc2–sc2 81–88, c3–c3 85–88, f3–f3 51–55. Prodorsum somewhat similar to female, with fine irregular mostly longitudinal plicae and weak rugose elements, becoming smoother centrally; anterior margin with small median forked projection forming a notch (4–5); with up to 4 pairs of minute pores sublaterally (can be difficult to see). Dorsal opisthosoma divided into mesonotal and pygidial shields; mesonotal shield captures setal pairs c1–c3 and d1–d3; pygidial shield posterior to D row setae with few arching and oblique plicae anteriorly becoming mostly smooth posteriorly; with 2 pairs large pores—mesad setae c3, and just anteromesad e3; with up to 4 pairs minute pores (can be difficult to see). Dorsal setae as in female, all short, smooth, fine, blunt (some setae appear to have 1–2 minute barbs). Setal measurements: v2 6–8, sc1 6–9, sc2 6–8, c1 6–7, c3 6–8, d1 7, d3 6–7, e3 6 –7, f3 6–7, h1 7.</p><p>Infracapitulum. (Figs 40, 41A) Dorsal cuticle with fine longitudinal plicae, ventral cuticle appears smooth; infracapitular setae m (8–11). Palp tarsus phaneres (1–2 and 7–8); femur-genu-tibia with single barbed dorsal seta (12).</p><p>Venter. (Fig. 41A, B) Ventral cuticle with fine longitudinal plicae between 1b–1a; transverse to oblique plicae on coxae I–II; fine transverse plicae from 1a–3a; with region of broadly V-shaped plicae anterior to 4a–4a; with band of fine transverse plicae 4a–ag, and with few weak transverse plicae ag –genital region; genital region often concealed under fold of cuticle (Fig. 41A, B), genital setae and anogenital opening flanked by plicae arching around region anteriorly. Setae g1 inserted just anterior to g2 on anterior margin of anogenital opening; valves with fine longitudinal plicae; pair of large pores laterad anal opening (Fig. 41A, B). Most ventral setae short, fine, smooth or with few minute barbs; except setae 1a and 4a elongate, fine, and often broken. Setae ps3 modified into short, thick and weakly clavate accessory genital stylets, often obscured under posterior projections of anogenital opening. Setal measurements: 1a 52–66, 1b 14–16, 1c 11–14, 2b 12–17, 2c 13–17, 3a 11–14, 3b 12–20, 4a 35–39, 4b 15–20, ag 10–12, g1 10–13, g2 11–13, ps2 6–8, ps3 6–7.</p><p>Aedeagus. (Fig. 41C) Sclerotised, elongate subulate (55–56), with narrow membranous tube emerging from sclerotised section.</p><p>Legs. (Fig. 42) Setal formulae and chaetotaxy same as adult female except ta I–II with solenidion ω′ also present. Tarsi I and II each with two solenidia (ta I adaxial ω′ 8–9, abaxial ωʺ 6–7; ta II adaxial ω′ 8–9, abaxial ωʺ 6–7); companion seta ftʺ barbed (both 7–8); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 7–8). Form of leg setae similar to that of female, except setae l″ on ge II and ti II tapered (not blunt).</p><p>Deutonymph. (1 measured)</p><p>Dorsum. Body measurements: v2–h1 126, sc2–sc2 86, c3–c3 89, f3–f3 42. Prodorsal and opisthosomal shields weakly developed. Prodorsum finely punctate with rounded central region indicated by folds; anterior margin of prodorsum smoothly rounded without median notch. Opisthosoma with broad weak plicae between c1–d1; pygidial shield finely punctate. Dorsal setae as in adults. Setal measurements: v2 3–5, sc1 4–6, sc2 4–5, c1 4–5, c3 3–4, d1 3–5, d3 3–4, e3 3 –5, f3 3–4, h1 3–4.</p><p>Infracapitulum. Cuticle as in adults; infracapitular setae m (7). Palp tarsus phaneres (1 and 6); femur-genu-tibia with single barbed dorsal seta (7).</p><p>Venter. Fine longitudinal plicae between 1b–1a; transverse plicae 1a–g1; longitudinal plicae surrounding genital region. Form of ventral setae as in adults. Setal measurements: 1a 44, 1b 14, 1c 10, 2b 12, 2c 10, 3a 8, 3b 9, 4a 21, 4b 8, ag 7, g1 6, ps2 3–4, ps3 3–4.</p><p>Legs. Setal formulae and chaetotaxy for legs I – IV same as adult female except trochanters 1-1-1-0. Tarsi I and II each with one abaxial solenidion ωʺ (both 4); barbed companion seta ftʺ (both 5–6); pair of prorals p′ζ, pʺζ (6–7).</p><p>Protonymph. (1 measured)</p><p>Dorsum. Body measurements: v2–h1 132, sc2–sc2 78, c3–c3 90, f3–f3 57. Prodorsum smooth, weak shield indicated with few strong longitudinal folds. Opisthosoma as in deutonymph. Setal measurements: v2 4–5, sc1 6, sc2 5, c1 5, c3 4, d1 4, d3 4, e3 3 –4, f3 3–4, h1 2–3.</p><p>Infracapitulum. Cuticle as in deutonymph. Palp tarsus phaneres (1 and 6); femur-genu-tibia with single barbed dorsal seta (7).</p><p>Venter. Ventral cuticle as in deutonymph. Setal measurements: 1a 48, 1b 10, 1c 10, 2c 9, 3a 10, 3b 11, ag 7, ps2 3, ps3 4.</p><p>Legs. Tarsi I and II each with one abaxial solenidion ωʺ (both 4); barbed companion seta ftʺ (both 6); pair of prorals p′ζ, pʺζ (both 5–6).</p><p>Colour. These mites were bright red when alive.</p><p>Host. Goodenia glabra R.Br. ( Goodeniaceae).</p><p>Distribution. This mite is so far known only from Inglewood in southeast Queensland, Australia.</p><p>Etymology. This species is named for the collection site, along the Wondul Stock Route.</p><p>Remarks. Host plant identification made by Queensland Herbarium. Goodeniacarus wondul sp. nov. was collected only 1. 3 km from, and on the same host plant as, the new genus Inglacarus . The host Goodenia glabra is an annual or perennial plant that grows after rain in sandy loamy soils in the drier inland parts of Australia, and yet this ephemeral resource is significant for these mites.</p></div>	https://treatment.plazi.org/id/03F8C5658036A036A7A1FF65FD33FC24	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA036A7A1FC20FBFEF918.text	03F8C565802DA036A7A1FC20FBFEF918.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Inglacarus Beard & Seeman 2025	<div><p>Inglacarus gen. nov.</p><p>Diagnosis. Palps 1-segmented, with single elongate tapered seta (apparently smooth).Anterior margin of prodorsum rounded, without median projection or notch; opisthosoma with 8 pairs of setae (c1, c3, d1, d3, e3, f3, h1, h2 present; c2, d2, e1, e2, f2 absent); dorsal setae short, fine, blunt. Ventral and genital shields not developed, both regions entirely membranous; two pairs of ps setae, anal valves membranous, ps2 and ps3 inserted longitudinally on anal valves; setae 4a short; 4a2 absent. Trochanter setal formula 1-1-2-1 (ontogeny of l′ on tr III unknown); trochanter III with seta l′ blunt. Femur III with seta d present, femur IV with seta d absent, formula 4-4-2-1. Genua III lacking seta l′, genual formula 3-3-0-0. Blunt leg setae: femur I setae d, l′; femur II d, l′, bv″; femur III d; genu I–II d, l′, l″; tibia I–II d, l″; tibia III–IV d. Tarsal claws and empodia pad-like. Usually three pairs of large pores visible on dorsal opisthosoma.</p><p>Remarks. Inglacarus gen. nov. is closest to Goodeniacarus gen. nov. in sharing a reduction of idiosomal dorsal setae, having short blunt dorsal idiosomal and leg setae, undeveloped ventral shields, the same leg setation, pectinate ventral tibial and tarsal setae, the morphology of the pretarsus, and the same host plant genus. The new genus is distinguished by having two regressive features: the reduction of the palp to just one segment with one phanere (probably the tibial dorsal seta due to length, suggesting loss of tarsal phaneres concomitant with the fusion or loss of the tarsus), and loss of the median forked projection. The new genus also has seta h2, which represents a significant plesiomorphic retention. These two genera are also distinguished from each other by the short seta 4a in Inglacarus jonesi sp. nov. versus long, attenuated in Goodeniacarus gen. nov. While the significance of this feature is not fully understood, it seems to be consistent within most other tenuipalpid genera; although there is some intraspecific variation within genera such as Aegyptobia and Dolichotetranychus .</p></div>	https://treatment.plazi.org/id/03F8C565802DA036A7A1FC20FBFEF918	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA032A7A1F8FCFD7BF9BA.text	03F8C565802DA032A7A1F8FCFD7BF9BA.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Inglacarus jonesi Beard & Seeman 2025	<div><p>Inglacarus jonesi sp. nov. Beard &amp; Seeman</p><p>(Figs 43–45)</p><p>Types. Holotype. Female, Australia, Wondul Stock Route, 36 km north of Inglewood, Queensland, 28°05’59’’S 151°04’26”E, ex. Goodenia glabra ( Goodeniaceae), 10.xii.2001, J.J. Beard and P.I. Forster (UQIC87350; 2 paratypes on same slide) . Paratypes. 5 females, same data as holotype (UQIC87347–87350). All material in QM .</p><p>Diagnosis. Female. Dorsal ornamentation rugose-reticulate medially, becoming weaker to smooth laterally; dorsal setae short, length 3–6. Setae g1 positioned posterior to g2. Seta ps2 slender, tapered to fine tip, ps3 tapered to blunt tip. Blunt setae on femora-tibiae: femur I d, l′; femur II d, l′, bv″; femur III d; genu I–II d, l′, l″; tibia I–II d, l″; tibia III–IV d. All blunt setae appear smooth.</p><p>Description</p><p>Adult female. (6 measured)</p><p>Dorsum. (Fig. 43) Body measurements: v2–h1 195–221 [217], sc2–sc2 111–128 [128], c3–c3 118–130 [126], f3–f3 75–80 [77]. Prodorsal and opisthosomal shields weakly developed; prodorsum mostly reticulate to rugose; anterior margin of prodorsum smoothly rounded without median forked projection or notch; with up to 3 pairs of minute pores sublaterally (can be difficult to see). Opisthosoma with rugose reticulation over large central region, becoming weaker posteriorly and mostly smooth at lateral margins; with 3 pairs of large pores—mesad c3, mesad d3, posteromesad d3; with at least 2 pairs of minute pores (can be difficult to see). Dorsal setae all short, smooth, blunt. Setal measurements: v2 4–6 [5–6], sc1 5–6 [5–6], sc2 4–6 [4–5], c1 5 [5], c3 4–5 [4–5], d1 4–5 [4–5], d3 4–5 [4–5], e3 4 –5 [4], f3 4–5 [4–5], h1 3–5 [4], h2 4–5 [4–5].</p><p>Infracapitulum. (Figs 43, 44) Dorsal cuticle with fine longitudinal striae; ventral cuticle smooth centrally with fine oblique striae laterally; infracapitular setae m (10–13) [11]. Palps one segmented, with single distal seta (11–14) [12–14].</p><p>Venter. (Fig. 44) Ventral cuticle with various patterns of fine to strong plicae; with few fine longitudinal plicae between setae 1b–1a and longitudinal to oblique plicae on coxae I–II; with widely spaced transverse plicae between setae 1a to just posterior to 3a; with small region broadly V-shaped plicae anterior to 4a–4a; with fine arching plicae between 4a–ag; genital region flanked by widely spaced longitudinal plicae. Setae g1 inserted just posterior to g2 on genital flap. Most ventral setae short, fine, apparently smooth, tapered; except setae 1a elongate, fine (often broken), and seta l′ on tr III short, blunt. Setal measurements: 1a 68–76 [68–70], 1b 14–19 [19], 1c 13–18 [16], 2b 11–18 [14], 2c 14–17 [14–15], 3a 12–16 [15–16], 3b 13–16 [16], 4a 14–19 [14–15], 4b 11–13 [11], ag 9–14 [13], g1 9–11 [10–11], g2 9–11 [10–11], ps2 7–9 [8–9], ps3 6–7 [6–7].</p><p>Spermathecal apparatus. (Fig. 45B) A moderately short membranous duct terminating in small bulb; basal half much broader than distal half, with a slightly enlarged bulb midway, and granulate texture basally.</p><p>Legs. (Fig. 45A) Setal formulae for legs I – IV: cx 2-2-1-1, tr 1-1-2-1, fe 4-4-2-1, ge 3-3-0-0, ti 5-5-3-3, ta 9(1)- 9(1)-5-5 respectively. Leg chaetotaxy for legs I – IV: cx I 1b, 1c; cx II 2b, 2c; cx III 3c; cx IV 4b; tr I–II IV v′, tr III l′, v′; fe I–II d, l′, v′, bvʺ; fe III d, ev′, fe IV ev′; ge I–II d, l′, lʺ, ge III–IV nude; ti I–II d, l′, lʺ, v′, vʺ; ti III–IV d, v′, vʺ; ta I–II ftʺ, tc′, p′ζ, pʺζ, u′, uʺ, ωʺ; ta III–IV ft′, tc′, tcʺ, u′, uʺ. Tarsi I and II each with one abaxial solenidion ωʺ (7 [7], 6–7 [7], respectively); barbed companion seta ftʺ subequal in length to solenidion (7–9 [7–8], 7–8 [7–8], respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 7–8). Solenidion and setae ft′, ftʺ inserted on small tubercles. Some leg setae blunt and of similar form to dorsal idiosomal setae (some may have 1–2 minute barbs)—seta l′ on trochanter III; seta d, l′ on femora I–II, d on femora III; all setae genua I–II; d, l″ on tibiae I–II and d on tibiae III–IV.</p><p>Etymology. This species is named after Mr John Jones, for his generous permission and assistance with collecting on his property.</p><p>Remarks. This species was collected at the same site and on the same host species as Goodeniacarus wondul sp. nov. (see Remarks for G. wondul sp. nov.).</p></div>	https://treatment.plazi.org/id/03F8C565802DA032A7A1F8FCFD7BF9BA	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658029A032A7A1F991FA44F85C.text	03F8C5658029A032A7A1F991FA44F85C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Goodeniacarus Beard & Seeman 2025	<div><p>Key to species of Goodeniacarus and Inglacarus, based on adult females</p><p>1. Palps with 3 segments............................................................ Goodeniacarus gen. nov. 2</p><p>- Palps with single segment.................................................. Inglacarus gen. nov. jonesi sp. nov.</p><p>2. Femur I with seta bv″ short, blunt............................................................. G. uri sp. nov.</p><p>- Femur I with seta bv″ finely tapered...................................................................... 3</p><p>3. Femur IV with 1 seta present (ev′ present, d absent).......................................................... 4</p><p>- Femur IV with 2 setae present (both ev′, d present).......................................................... 5</p><p>4. Tibia I with setae d and l″ short, blunt; femur II with seta d short, blunt............................ G. leiperi sp. nov.</p><p>- Tibia I with setae d short, blunt and setae l′ tapered; femur II with seta d tapered.................... G. wondul sp. nov.</p><p>5. Femur II with seta bv″ long, finely tapered; tibia II with seta l″ finely tapered; both prodorsum and dorsal opisthosoma with fully reticulate cuticle................................................................ G. reticulatus sp. nov.</p><p>- Femur II with seta bv″ short, bluntly tapered; tibia II with seta l″ short, blunt; prodorsum and dorsal opisthosoma with some regions of weak reticulate cuticle........................................................ G. rosulatus sp. nov.</p></div>	https://treatment.plazi.org/id/03F8C5658029A032A7A1F991FA44F85C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Beard, Jennifer J.;Seeman, Owen D.	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
