taxonID	type	description	language	source
03F8C565801DA006A7A1FB57FC5DF8BA.taxon	type_taxon	Type species: Goodeniacarus rostulatus sp. nov. Beard & Seeman	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801DA006A7A1FB57FC5DF8BA.taxon	diagnosis	Diagnosis. Female. Palps 3 - segmented; palp tarsus with 2 terminal phaneres (1 long and 1 minute). Anterior margin of prodorsum with small median forked projection forming a notch; dorsal opisthosoma with 7 pairs of setae (c 1, c 3, d 1, d 3, e 3, f 3, h 1 present; c 2, d 2, e 1, e 2, f 2, h 2 absent); dorsal setae short, fine with blunt rounded tips; 1 – 3 pairs of large pores present on opisthosoma (often with an associated minute pore visible, but difficult to see). Ventral and genital shields not developed, both regions membranous and merged together; 2 pairs of ps setae, anal valves membranous, ps 2 and ps 3 inserted longitudinally on anal valves; setae 4 a elongate fine; lacking supernumerary coxal setae 3 a 2 and 4 a 2. Setal formulae for legs I – IV: cx 2 - 2 - 1 - 1, tr 1 - 1 - 2 - 1, fe 4 - 4 - 2 - 1 / 2, ge 3 - 3 - 0 - 0, ti 5 - 5 - 3 - 3, ta 9 (1) - 9 (1) - 5 - 5 respectively. Trochanter III with seta l′ blunt, delayed to adult. Femur IV with seta d present or absent. Setae on genua I – II all added in deutonymph. Tarsus I – II with seta tcʺ delayed to adult. Femora I – IV, genua I – II, tibiae I – IV usually with either or both setae d and l ″ blunt. Tarsal claws and empodia pad-like.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801DA006A7A1FB57FC5DF8BA.taxon	discussion	Remarks. We cannot determine with certainty if the palp phaneres are eupathidia and / or setae. This genus has a thick microplate layer that persists intact even after mounting. This layer often obscures features, especially the minute dorsal pores. The ultrafine detail of this layer can be seen in the images taken using low temperature scanning electron microscopy (LTSEM) in Figures 6, 9, 11, 14, 31.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801DA006A7A1FC05FD8CFBF6.taxon	type_taxon	Type genus — Tenuipalpus Donnadieu, 1875	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.taxon	description	(Figs 1 – 4)	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.taxon	materials_examined	Types. Holotype. Female, Australia, Ranger Road, approx. 5 km S of Warwick along New England Highway, Queensland, 28 ° 16 ’ 50.1 ” S 151 ° 58 ’ 31.5 ” E, ex. Goodenia rotundifolia R. Br. (Goodeniaceae), 25. iii. 2016, J. J. Beard and P. I. Forster. Paratypes. 5 females, same data as holotype. All specimens deposited in QM.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.taxon	diagnosis	Diagnosis. Female. Dorsal ornamentation comprising plicae forming irregular fine reticulation and folds; dorsal setae length 5 – 8. Setae g 1 positioned slightly posterior to g 2. Setae ps 2 and ps 3 blunt, lengths ps 2 5 – 6, ps 3 5 – 6. Femora I – II with seta bv ″ long, tapered; femora IV with 1 seta (ev′ present, d absent); tarsus I – II with companion seta ftʺ slightly longer than solenidion ωʺ. Blunt setae on femora and tibiae: femur I – III d; genu I – II d, l ″; tibia I – II l ″; tibia III – IV d. Blunt leg setae can have 1 – 2 small barbs (can be difficult to see).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.taxon	description	Description Adult female. (6 measured) Dorsum. (Figs 1, 2) Body measurements: v 2 – h 1 192 – 209 [209], sc 2 – sc 2 122 – 133 [132], c 3 – c 3 126 – 141 [134], f 3 – f 3 74 – 88 [88]. Prodorsal and opisthosomal shields weakly developed. Prodorsum with fine irregular plicae forming irregular reticulation and folds; with shallow sublateral furrows; anterior margin with small median forked projection forming a notch (5 – 10) [5 – 7]; with a pair of minute pores sublaterally (can be difficult to see) and discrete pore-like invagination mesad sc 2 (indicated by arrows in Fig. 1). Opisthosoma cuticle pattern similar to that of prodorsum, with complicated plicae forming fine irregular reticulation, becoming smoother anteriorly between c 1 – c 1 and d 1 – d 1; with transversely plicate band of sclerotised cuticle around opisthosomal shield margin, most apparent when specimen flattened (Fig. 2); with 3 pairs of large pores — just mesad setae c 3, mesad d 3 and anteromesad e 3; with up to 4 pairs minute pores visible (can be difficult to see). Dorsal setae all short, blunt. Setal measurements: v 2 6 – 7 [6 – 7], sc 1 6 – 8 [6 – 7], sc 2 6 – 8 [6], c 1 6 – 8 [6 – 7], c 3 5 – 7 [5 – 6], d 1 5 – 8 [6], d 3 5 – 6 [5 – 6], e 3 5 – 6 [5 – 6], f 3 5 – 6 [5 – 6], h 1 4 – 6 [5]. Infracapitulum. (Figs 1, 3 A) Dorsal cuticle appears smooth, ventral cuticle with fine oblique plicae laterally; infracapitular setae m fine (11 – 13) [12 – 13]. Palp tarsus phaneres (2 – 3 and 8 – 9); femur-genu-tibia with single barbed dorsal seta (13 – 15) [13 – 14]. Venter. (Fig. 3 A) Ventral cuticle with various patterns of fine to strong plicae; with fine longitudinal plicae between setae 1 b – 1 a and longitudinal to oblique plicae on coxae I – II; with transverse plicae between setae 1 a – 3 a becoming widely spaced 3 a – 4 a; with small region of broadly U-shaped plicae anterior to 4 a – 4 a; with fine arching plicae demarking anterior margin of genital flap; genital region flanked by widely spaced longitudinal plicae. Setae g 1 inserted just posterior to g 2 on genital flap; flap with longitudinal to irregular plicae between ag – g 1. Most ventral setae short, fine, appear smooth or with 1 – 2 barbs; except setae 1 a and 4 a elongate, fine, and often broken; ps 2 and ps 3 blunt. Setae ps 3 often obscured within lateral folds of genital opening; with large, rounded pore in soft cuticle laterad anal opening. Setal measurements: 1 a 64 – 96 [75 – 86], 1 b 18 – 20 [19], 1 c 14 – 16 [15], 2 b 18 – 21 [18 – 19], 2 c 14 – 17 [15 – 16], 3 a 13 – 17 [15], 3 b 11 – 19 [16], 4 a 41 – 56 [56], 4 b 15 – 17 [17], ag 9 – 12 [10 – 11], g 1 8 – 12 [8 – 9], g 2 9 – 13 [9], ps 2 5 – 6 [5], ps 3 5 – 6 [6]. Spermathecal apparatus. (Fig. 3 B) A moderately short membranous duct with a broader bulb midway, and narrower distal portion with a granulate texture, terminating in a small, rounded bulb. Legs. (Fig. 4) See genus Diagnosis for full setal formulae for legs I – IV, except femur IV with 1 seta (ev′ present, d absent). See description of type species, G. rosulatus sp. nov., for full leg chaetotaxy. Tarsi I and II each with one abaxial solenidion ωʺ (both 5 – 6 [5 – 6, 6]); barbed companion seta ftʺ slightly longer than solenidion (7 – 9 [8], 8 – 9 [8], respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 8 – 9). Solenidion and setae ft′ inserted on small tubercles. Some leg setae blunt, of similar form to dorsal idiosomal setae except can have 1 – 2 small barbs (barbs can be difficult to see): seta l′ on trochanter III; seta d on femora I – II, d on femora III; d, l ″ on genua I – II; l ″ on tibiae I – II, and d on tibiae III – IV. Colour. These mites were bright red when alive.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.taxon	biology_ecology	Host. Goodenia rotundifolia R. Br. (Goodeniaceae).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.taxon	distribution	Distribution. This mite is so far known only from Warwick in southeast Queensland, Australia.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.taxon	etymology	Etymology. This species is named for the field botanist and photographer Glenn Leiper, who has helped the senior author collect and identify host plants on many occasions over the years.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565801EA003A7A1FF65FEADF9BD.taxon	discussion	Remarks. Host plant identification made by Queensland Herbarium (BRI voucher: PIF 43590). Goodeniacarus leiperi sp. nov. and G. uri sp. nov. were collected from the same species of host plant approximately 160 km apart. These two species can easily be separated using the following characters: G. leiperi sp. nov. dorsum with fine irregular reticulation, seta bv ″ on femur I – II finely tapered, setae l′ on femur II and genu II finely tapered vs G. uri sp. nov. dorsum with mostly elongate longitudinal plicae, bv ″ on femur I – II short and blunt, setae l′ on femur II and genu II short, blunt.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658018A017A7A1F998FB2BFAF9.taxon	description	(Figs 5 – 15) Types. Holotype. Female, Australia, Herberton Range Conservation Park, above Hall’s Falls on North Wondecla Creek, Far North Queensland, 17 ° 24 ’ 37.5 ” S 145 ° 26 ’ 55.5 ” E, ex. Goodenia grandifolia Sims. (Goodeniaceae), 27. iv. 2017, J. J. Beard, P. I. Forster and K. McDonald. Paratypes. 14 females, 7 males, 2 deutonymphs, same data as holotype. All specimens deposited in QM. Other material examined. 7 females, 1 male, same data as holotype (QM).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658018A017A7A1F998FB2BFAF9.taxon	diagnosis	Diagnosis. Female. Dorsal ornamentation mostly reticulate; dorsal setae length 6 – 10. Setae g 1 positioned anterior to g 2. Setae ps 2 and ps 3 slender, tapered, lengths ps 2 9 – 11, ps 3 8 – 10. Femora I – II with seta bv ″ long, tapered; femora IV with 2 seta (ev′, d present); tarsus I – II with companion seta ftʺ much longer than solenidion ωʺ. Blunt setae on femora and tibiae: femur I – IV d; genu I – II d, l ″; tibia I d, l ″, tibia II l ″, tibia III – IV d. Blunt setae smooth.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658018A017A7A1F998FB2BFAF9.taxon	description	Description Adult female. (6 measured) Dorsum. (Figs 5, 6) Body measurements: v 2 – h 1 208 – 215 [215], sc 2 – sc 2 132 – 143 [143], c 3 – c 3 134 – 150 [150], f 3 – f 3 79 – 86 [81]. Prodorsal and opisthosomal shields weakly developed. Prodorsum mostly with reticulate cuticle; anterior margin with small median forked projection forming a broad shallow notch (4 – 7) [7]; with two pairs minute pores visible (can be difficult to see; indicated by arrows in Fig. 5). Opisthosoma similarly with mostly reticulate cuticle, with band of irregular plicae anteriorly; with two pairs large pores — just mesad setae d 3 and anterior to e 3; with up to four pairs minute pores (can be difficult to see). Dorsal setae all short, fine, blunt. Setal measurements: v 2 7 – 8 [8], sc 1 8 – 10 [9], sc 2 8 – 9 [8 – 9], c 1 7 – 9 [8], c 3 7 – 8 [7], d 1 6 – 7 [7], d 3 7 – 8 [7], e 3 7 – 8 [7 – 8], f 3 7 – 8 [8], h 1 6 – 8 [7 – 8]. Infracapitulum. (Figs 5, 7) Dorsal cuticle with fine longitudinal plicae, ventral cuticle appears smooth; infracapitular setae m barbed (12 – 14) [13]. Palp tarsus phaneres (3 and 8 – 10) [8 – 9]; femur-genu-tibia with single barbed dorsal seta (12 – 17) [12]. Venter. (Fig. 7) Ventral cuticle with oblique plicae on coxae I – II; with widely spaced transverse plicae between setae 1 a – 4 a; with small region of broadly U-shaped plicae anterior to 4 a – 4 a; with irregular arching plicae surrounding ag setae; with transverse plicae and weakly verrucose cuticle surrounding g 1 – g 2 setae; genital region flanked by widely spaced longitudinal plicae. Setae g 1 inserted anterior to g 2 on genital flap. Most ventral setae short, fine, apparently smooth; except setae 1 a and 4 a elongate, fine, and often broken. Setae ps 3 often obscured within lateral folds of genital opening. Pair of large, round pores often visible in soft cuticle laterad anus. Setal measurements: 1 a 62 – 88 [68 – 88], 1 b 17 – 24 [20 – 22], 1 c 19 – 23 [21], 2 b 25 – 23 [28 – 32], 2 c 26 – 30 [28], 3 a 15 – 23 [23], 3 b 27 – 34 [34], 4 a 40 – 85 [81], 4 b 22 – 27 [24], ag 14 – 18 [16], g 1 13 – 16 [13], g 2 13 – 16 [15], ps 2 9 – 11 [9 – 10], ps 3 8 – 10 [9]. Spermathecal apparatus. A moderately short membranous duct with a broader bulb midway, and narrower distal portion with a granulate texture, terminating in a small, rounded bulb. Legs. (Fig. 8) See genus Diagnosis for full setal formulae for legs I – IV, and description of type species, G. rosulatus sp. nov., for full leg chaetotaxy. Tarsi I and II each with one abaxial solenidion ωʺ (5 – 6 [5], 4 – 5 [4], respectively); barbed companion seta ftʺ much longer than solenidion (13 – 15 [15], 11 – 14 [12], respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 9 – 10). Solenidion and setae ft′ inserted on small tubercles. Some leg setae blunt and of similar form to dorsal idiosomal setae — seta l′ on trochanter III; seta d on femora I – IV; d, l ″ on genua I – II; d, l ″ on tibiae I; l ″ on tibiae II; d on tibiae III – IV. Microplates. (Figs 6, 9) The microplate layer appears coarsely granular at 1,000 X magnification using DIC, and likewise at 2,000 X using scanning electron microscopy (Fig. 9 A). Individual microplates are hexagonal in shape (Figs 9 C, D), and have a finely granular surface visible at 20,000 X (Fig. 9 D). The microplates on the male (Fig. 11) appear differently to those on the female, however this may simply be due to the image being taken at a different developmental stage to those of the female. The individual plates on the male appear rounded and domed, and interconnected by smaller rectangular microplates; however, the hexagonal structure seen in the female plates is also apparent. Adult male. (7 measured). Dorsum. (Figs 10, 11) Body measurements: v 2 – h 1 145 – 154, sc 2 – sc 2 97 – 105, c 3 – c 3 94 – 104, f 3 – f 3 52 – 57. Prodorsum mostly smooth with few wrinkles and plicae, small sublateral region with some weak rugose reticulation indicated; with 3 – 4 pairs of minute pores visible sublaterally to centrally (can be difficult to see); anterior margin with small median forked projection forming a notch (2 – 5). Dorsal opisthosoma divided into mesonotal and pygidial shields; mesonotal shield with some weak irregular plicae, captures setal pairs c 1 – c 3 and d 1 – d 3; pygidial shield posterior to D row setae with few arching plicae anterior to mostly smooth central region and oblique plicae laterally. Opisthosoma with 3 pairs of large pores — just mesad setae c 3, mesad d 3, and anteromesad e 3; with up to four pairs minute pores (difficult to see). Dorsal setae as in female, all short, fine, blunt. Setal measurements: v 2 7 – 8, sc 1 7 – 9, sc 2 7 – 10, c 1 7 – 8, c 3 6 – 8, d 1 7 – 8, d 3 6 – 8, e 3 7 – 8, f 3 7 – 9, h 1 7 – 8. Infracapitulum. (Figs 10, 12) Dorsal cuticle with fine longitudinal plicae, ventral cuticle with fine oblique plicae laterally or smooth; infracapitular setae m (8 – 10). Palp tarsus phaneres (2 – 3 and 7 – 8); femur-genu-tibia with single barbed dorsal seta (10 – 13). Venter. (Figs 12, 13, 14) Ventral cuticle with fine longitudinal plicae 1 b – 1 a; with widely spaced transverse plicae from 1 a – 3 a; with region of broadly V-shaped plicae anterior to 4 a – 4 a; with band of fine transverse plicae 4 a – ag, and with few weak transverse plicae ag – genital region; genital region can be concealed under fold of cuticle; genital setae and opening flanked by plicae arching around region anteriorly. Setae g 1 inserted just anterior to g 2 on anterior margin of anogenital opening. Most ventral setae short, fine, apparently smooth or with 1 – 2 minute barbs (can be seen in Fig. 13); except setae 1 a and 4 a elongate, fine, and often broken. Setae ps 3 modified into short, thick, clavate accessory genital stylets (Fig. 14), often obscured under posterior projections of anogenital opening. Setal measurements: 1 a 50 – 58, 1 b 14 – 18, 1 c 14 – 18, 2 b 17 – 21, 2 c 21 – 23, 3 a 13 – 20, 3 b 19 – 23, 4 a 37 – 60, 4 b 15 – 26, ag 11 – 13, g 1 11 – 13, g 2 12 – 14, ps 2 7 – 11, ps 3 6 – 7. Aedeagus. (Fig. 12) Sclerotised, elongate, subulate (58 – 64), with narrow membranous tube emerging from sclerotised section. Legs. (Fig. 15) Setal formulae and chaetotaxy same as adult female except ta I – II with solenidion ω′ also present. Tarsi I and II each with two solenidia (both with adaxial ω′ 6 – 8, abaxial ωʺ 5 – 6); companion seta ftʺ barbed (9 – 12, 9 – 10, respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 8 – 9). Form of leg setae similar to that of female, except setae bv ″ is blunt on femur II. Deutonymph. (2 measured) Dorsum. Body measurements: v 2 – h 1 173 – 187, sc 2 – sc 2 112 – 127, c 3 – c 3 118 – 123, f 3 – f 3 56 – 60. Prodorsal and opisthosomal shields weakly developed. Prodorsum with smooth cuticle; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with widely spaced broken transverse plicae between c 1 – d 1; posterior d 1 – d 1 with few longitudinal folds centrally to sublaterally. Dorsal setae as in adults. Setal measurements: v 2 4 – 5, sc 1 5 – 6, sc 2 5 – 6, c 1 6 – 7, c 3 4 – 5, d 1 4 – 5, d 3 3 – 5, e 3 4 – 5, f 3 4 – 5, h 1 4. Infracapitulum. Cuticle as in adults; infracapitular setae m (7 – 8). Palp tarsus phaneres (1 – 2 and 6 – 7); femur-genu-tibia with single barbed dorsal seta (9 – 8). Venter. Ventral cuticle finely plicate; with fine longitudinal plicae 1 b – 1 a; transverse plicae 1 a to just posterior 4 a; plicae circling around setae ag with longitudinal plicae inside circle; with band of transverse plicae between ag – ps 3. Form of ventral setae as in adults. Setal measurements: 1 a 58, 1 b 14 – 18, 1 c 7 – 10, 2 b 10 – 15, 2 c 8 – 11, 3 a 8 – 13, 3 b 9 – 13, 4 a 24 – 34, 4 b 7 – 11, ag 5 – 8, g 1 6 – 8, ps 2 3, ps 3 3 – 4. Legs. Setal formulae and chaetotaxy for legs I – IV same as adult female except trochanters 1 - 1 - 1 - 0. Tarsi I and II each with one abaxial solenidion ωʺ (4 – 5, 4, respectively); barbed companion seta ftʺ (both 5 – 6); pair of prorals p′ζ, pʺζ (6 – 7, 5 – 6, respectively). Some leg setae short to minute, blunt — seta l′ on trochanter III, seta d on femora I – IV, genua I – II, tibiae III – IV, and seta lʺ on genua I – II and tibiae I – II.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658018A017A7A1F998FB2BFAF9.taxon	biology_ecology	Host. The host plant, Goodenia grandifolia Sims. (Goodeniaceae), has sticky and fragrant leaves, and the mites were observed in higher numbers on the older, less sticky leaves.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658018A017A7A1F998FB2BFAF9.taxon	etymology	Etymology. This species is named for its obviously reticulate dorsum.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658018A017A7A1F998FB2BFAF9.taxon	discussion	Remarks. Host plant identification made by Queensland Herbarium (BRI voucher: PIF 45191).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.taxon	description	(Figs 16 – 25)	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.taxon	materials_examined	Types. Holotype. Female, Australia, Rungulla National Park, 37. 3 km SSE ranger station and airstrip, Queensland, 19 ° 28 ’ 00 ’’ S 143 ° 31 ’ 33 ” E, ex. Goodenia rosulata Domin. (Goodeniaceae), 08. v. 2022, J. J. Beard (QM). Paratypes. 16 females, 5 males, 4 deutonymphs, 3 protonymphs, 3 larvae, same data as holotype (QM). Other material examined. 6 females, 1 male (pharate), 4 deutonymphs, 1 protonymph, same data as holotype (QM).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.taxon	diagnosis	Diagnosis. Female. Prodorsal ornamentation weaky reticulate, becoming weak and irregular medially and laterally; opisthosomal ornamentation reticulate, becoming weak laterally; dorsal setae length 4 – 9. Setae g 1 positioned just anterior to g 2. Setae ps 2 and ps 3 slender, tapered, lengths ps 2 7 – 10, ps 3 5 – 9 (sometimes with a blunt tip). Femora I with setae bv ″ long, tapered to fine tip; femora II with bv ″ shorter, blunt; femora IV with 2 seta (ev′, d present; companion seta ftʺ much longer than solenidion ωʺ. Blunt setae present on femora and tibiae: femur I – IV d, femur II bv ″; genu I – II d, l ″; tibia I – II l ″; tibia III – IV d. All blunt setae with minute barbs.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.taxon	description	Description Adult female. (14 measured) Dorsum. (Fig. 16) Body measurements: v 2 – h 1 187 – 213 [193], sc 2 – sc 2 121 – 140 [128], c 3 – c 3 119 – 139 [128], f 3 – f 3 62 – 81 [72]. Prodorsal and opisthosomal shields weakly developed. Prodorsum with reticulate cuticle becoming weak and irregular medially and laterally; with shallow longitudinal sublateral furrows; anterior margin with small median forked projection forming a notch (5 – 10) [8]; with up to 4 pairs of minute pores visible sublaterally. Opisthosoma with stronger reticulation over large central region, becoming weak and irregular medially and laterally; with 3 pairs of large pores — mesad setae c 3, mesad d 3, and posteromesad d 3; with up to 3 pairs minute pores (can be difficult to see). Dorsal setae all short, fine, blunt. Setal measurements: v 2 6 – 8 [7], sc 1 5 – 9 [5 – 6], sc 2 6 – 9 [6], c 1 5 – 8 [5], c 3 5 – 7 [5], d 1 4 – 7 [5], d 3 5 – 6 [5], e 3 5 – 7 [5], f 3 5 – 7 [5], h 1 5 – 7 [5]. Infracapitulum. (Figs 16, 17 A – B) Cuticle on dorsal infracapitulum with fine longitudinal plicae; infracapitular setae m (13 – 18) [14]. Palp tarsus phaneres (2 and 8 – 9); femur-genu-tibia with single barbed dorsal seta (12 – 14) (Fig. 17 A – B). Venter. (Fig. 18) Ventral cuticle with fine longitudinal plicae between setae 1 b – 1 a and longitudinal to oblique on coxae I – II; with widely spaced transverse plicae between setae 1 a – 3 a; with small region of broadly U-shaped plicae anterior to 4 a – 4 a; with fine arching plicae demarking anterior margin of genital flap; genital region flanked by widely spaced longitudinal plicae. Setae g 1 inserted just anterior to g 2 on genital flap. Most ventral setae short, fine, smooth or minutely barbed (difficult to see); except setae 1 a and 4 a elongate, fine, and often broken. Setae ps 2, ps 3 slender, tapered; ps 3 often obscured within lateral folds of genital opening, sometimes with minutely blunt tip. Pair of large, round pores often visible in soft cuticle laterad anus. Setal measurements: 1 a 64 – 86 [73 – 83], 1 b 15 – 29 [20 – 24], 1 c 15 – 22 [17], 2 b 15 – 23 [18], 2 c 18 – 24 [21], 3 a 14 – 19 [13 – 15], 3 b 18 – 21 [18], 4 a 56 – 76 [61 – 66], 4 b 13 – 17 [15], ag 13 – 21 [21], g 1 10 – 12 [10 – 11], g 2 10 – 13 [12], ps 2 7 – 10 [7], ps 3 5 – 9 [6]. Spermathecal apparatus. (Fig. 17 C) A moderately short membranous duct with a broader bulb midway, and narrower distal portion with a granulate texture, terminating in a small, rounded vesicle. Legs. (Fig. 19) See genus Diagnosis for full leg setal formulae. Chaetotaxy for legs I – IV: cx I 1 b, 1 c; cx II 2 b, 2 c; cx III 3 c; cx IV 4 b; tr I – II, IV v′, tr III l′, v′; fe I – II d, l′, v′, bvʺ; fe III – IV d, ev′; ge I – II d, l′ – lʺ, ge III – IV nude; ti I – II d, l′, lʺ, v′, vʺ; ti III – IV d, v′, vʺ; ta I – II ft′, ftʺ, tc′, tc ″, p′ζ, pʺζ, u′, uʺ, ωʺ; ta III – IV ft′, tc′, tcʺ, u′, uʺ. Tarsi I and II each with one abaxial solenidion ωʺ (5 – 7 [5], 4 – 6 [5], respectively); barbed companion seta ftʺ much longer than solenidion (10 – 13 [10], 9 – 12 [10], respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 8 – 10). Solenidion and setae ft′, ftʺ inserted on small tubercles. Some leg setae blunt and of similar form to dorsal idiosomal setae except with minute barbs — seta l′ on trochanter III, seta d on femora I – IV, bv ″ on femur II; seta d on genua I – II, tibiae III – IV; and seta lʺ on genua I – II and tibiae I – II. Adult male. (4 measured) Dorsum. (Fig. 20) Body measurements: v 2 – h 1 144 – 148, sc 2 – sc 2 96 – 104, c 3 – c 3 91 – 100, f 3 – f 3 47 – 51. Prodorsum with weakly developed irregular reticulation and folds sublaterally, with smooth central region and becoming smooth laterally; anterior margin of prodorsum with small median forked projection forming a notch (4 – 7); with at least 2 pairs of minute pores sublaterally (can be difficult to see). Dorsal opisthosoma with pair of weakly developed metanotal shields indicated between setal pairs c 1 – c 3 and d 1 – d 3; and posterior to D row setae with few oblique plicae from e 3 – f 3 to central smooth region. Dorsal opisthosoma divided into mesonotal and pygidial shields; with at least 2 pairs large pores visible — mesad c 3, anteromesad e 3; with at least 3 pairs minute pores (can be difficult to see). Dorsal setae as in female, all short, fine, blunt. Setal measurements: v 2 5 – 6, sc 1 6 – 7, sc 2 6 – 7, c 1 5 – 8, c 3 4 – 6, d 1 5 – 6, d 3 4 – 6, e 3 5 – 6, f 3 6 – 7, h 1 5 – 6. Infracapitulum. (Fig. 20) Cuticle on dorsal infracapitulum with fine longitudinal plicae; infracapitular setae m (8 – 10). Palp tarsus phaneres (2 and 6 – 7); femur-genu-tibia with single barbed dorsal seta (9 – 11). Venter. (Fig. 21) Ventral cuticle with widely spaced transverse plicae from 1 a – 3 a; with small region broadly U-shaped plicae anterior to 4 a – 4 a between coxae III – IV; with band of fine transverse plicae 4 a – ag, and with few weak transverse plicae ag – genital setae; genital region with few flanking plicae arching around region anteriorly. Setae g 1 inserted just anterior to g 2 on anterior margin of anogenital opening. Most ventral setae short, fine, smooth or minutely barbed (difficult to see); except setae 1 a and 4 a elongate, fine, and often broken. Setae ps 3 modified into short, thick, clavate accessory genital stylets, often obscured under posterior projections of anogenital opening. Setal measurements: 1 a 48 – 55, 1 b 12 – 15, 1 c 13 – 14, 2 b 15 – 21, 2 c 14 – 17, 3 a 11 – 13, 3 b 14 – 16, 4 a 28 – 45, 4 b 12 – 15, ag 9 – 14, g 1 9 – 11, g 2 11 – 12, ps 2 6 – 8, ps 3 5 – 6. Aedeagus. (Fig. 21) Sclerotised, elongate, subulate (57 – 58), with narrow membranous tube emerging from sclerotised section. Legs. (Fig. 22) Setal formulae and chaetotaxy same as adult female except ta I – II with solenidion ω′ also present; ω′ inserted somewhat proximally and partially concealed by strong transverse cuticular ridge across segment. Tarsi I and II each with two solenidia (ta I adaxial ω′ 7 – 9, abaxial ω ″ 6 – 7; ta II adaxial ω′ 7 – 8, abaxial ω ″ 5 – 6); companion seta ftʺ barbed (9 – 10, 6 – 9, respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 7 – 9). Form of leg setae similar to that of female. Deutonymph. (4 measured) Dorsum. Body measurements: v 2 – h 1 168 – 188, sc 2 – sc 2 112 – 118, c 3 – c 3 117 – 121, f 3 – f 3 47 – 51. Prodorsal and opisthosomal shields weakly developed, with smooth cuticle; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with a few broad transverse striae. Dorsal setae form as in adults. Setal measurements: v 2 3 – 4, sc 1 3 – 4, sc 2 3 – 4, c 1 3 – 4, c 3 3, d 1 2 – 3, d 3 2 – 3, e 3 2 – 3, f 3 2 – 3, h 1 2 – 3. Infracapitulum. Cuticle as in adults; infracapitular setae m (7 – 8). Palp tarsus phaneres (1 – 2 and 5 – 6); femur-genu-tibia with single barbed dorsal seta (8). Venter. (Fig. 23 A) Widely spaced longitudinal plicae between 1 b – 1 a, widely spaced transverse plicae 1 a to just posterior 4 a; plicae forming a triangle around setae ag with transverse plicae within triangle; with band of transverse plicae between ag – g 1 and mixed plicae in genital region. Form of ventral setae as in adults. Setal measurements: 1 a 55 – 72, 1 b 10 – 12, 1 c 8 – 10, 2 b 10 – 12, 2 c 11 – 14, 3 a 7 – 10, 3 b 7 – 12, 4 a 27 – 32, 4 b 8 – 12, ag 6 – 7, g 1 4 – 5, ps 2 2 – 3, ps 3 2 – 4. Legs. (Fig. 23 B) Setal formulae and chaetotaxy for legs I – IV same as adult female except tarsi 8 (1) - 8 (1) - 3 - 3 (seta tcʺ absent on ta I – II). In addition to G. rosulatus sp. nov., the deutonymph is known for G. reticulatus sp. nov., G. uri sp. nov. and G. wondul sp. nov., and these species all share trochanter III with only v′ present, and the addition of l′ being delayed to the adult stage. It should be noted, however, that seta l′ is inconsistently expressed on tr III in the deutonymph of G. rosulatus sp. nov. Of the 8 individual deutonymphs we examined, 2 had v′ symmetrically expressed (1 seta on both sides), 3 had setae v′ and l′ on one side and only v′ on the other (l′ asymmetrically expressed), 3 specimens had only 1 trochanter III clearly visible and 2 of these had 1 seta present and 1 had 2 setae present. As the most common pattern was seta v′ only present, we consider the presence of seta l′ in the deutonymph to be an aberration. Tarsi I and II each with one abaxial solenidion ωʺ (both 3 – 4); barbed companion seta ftʺ (both 4 – 6); pair of prorals p′ζ, pʺζ (6 – 9, 6 – 7, respectively). Some leg setae short to minute, blunt — seta l′ on trochanter III, seta d on femora I – IV, genua I – II, tibiae III – IV, and seta lʺ on genua I – II and tibiae I – II. Protonymph. (2 measured) Dorsum. Body measurements: v 2 – h 1 140 – 144, sc 2 – sc 2 98 – 99, c 3 – c 3 98 – 105, f 3 – f 3 35 – 40. Prodorsal and opisthosomal shields weakly developed; prodorsum mostly smooth, with smoothly rounded anterior margin without median notch; dorsal opisthosoma with few broadly spaced transverse plicae. Dorsal setae form as in adults. Setal measurements: v 2 3 – 4, sc 1 3 – 4, sc 2 4, c 1 4 – 5, c 3 3 – 4, d 1 3 – 4, d 3 3 – 4, e 3 2 – 3, f 3 2 – 3, h 1 2 – 3. Infracapitulum. Cuticle as in deutonymph; infracapitular setae m (5 – 6). Palp tarsus phaneres (1 and 5); femur-genu-tibia with single barbed dorsal seta (7 – 8). Venter. Ventral cuticle and setae as in deutonymph. Setal measurements: 1 a 58, 1 b 7 – 10, 1 c 8 – 9, 2 c 8 – 12, 3 a 8 – 10, 3 b 8 – 9, ag 5 – 7, ps 2 2, ps 3 2. Legs. (Fig. 24) Setal formulae for legs I – IV: cx 2 - 1 - 1 - 0, tr 0 - 0 - 0 - 0, fe 3 - 3 - 2 - 2, ge 0 - 0 - 0 - 0, ti 5 - 5 - 3 - 3, ta 7 (1) - 7 (1) - 3 - 3 respectively. Leg chaetotaxy: same as larva except 1 c, 2 c, 3 b added to cx I, II, III respectively; and tc′ added to ta I – II. Tarsi I and II each with one abaxial solenidion ωʺ (both 3); barbed companion seta ftʺ (5, 4 – 5, respectively); pair of prorals p′ζ, pʺζ (both 5 – 6). Larva. (2 measured) Dorsum. (Fig. 25) Body measurements: v 2 – h 1 117 – 122, sc 2 – sc 2 86 – 108, c 3 – c 3 84 – 96, f 3 – f 3 27 – 31. No shields developed. Prodorsum sparsely finely colliculate anteriorly, and few short, irregular plicae posteriorly; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with few widely spaced transverse striae between c 1 – d 1, and short oblique plicae d 1 – h 1. Dorsal setae form as in adults, except much shorter. Setal measurements: v 2 3, sc 1 3 – 4, sc 2 3 – 4, c 1 3 – 4, c 3 2 – 3, d 1 3, d 3 2 – 3, e 3 2 – 3, f 3 2 – 3, h 1 2 – 3. Infracapitulum. Cuticle with fine transverse plicae; infracapitular setae m absent. Palp tarsus phaneres (1 and 5); femur-genu-tibia with single barbed dorsal seta (5 – 6). Venter. Ventral cuticle and setae as in deutonymph; fine longitudinal plicae 1 b – 1 a; fine transverse plicae from 1 a – ps 3; region between coxae III – III with finely granulate cuticle with mixed oblique plicae. Setal measurements: 1 a 38 – 39, 1 b 8 – 9, 3 a 6 – 7, ps 2 2 – 3, ps 3 2 – 3. Legs. (Fig. 25) Setal formulae for legs I – III: cx 1 - 0 - 0, tr 0 - 0 - 0, fe 3 - 3 - 2, ge 0 - 0 - 0, ti 5 - 5 - 3, ta 7 (1) - 7 (1) - 3 respectively. Leg chaetotaxy: cx I 1 b; fe I – II d, v′, bvʺ; fe III d, ev′; ge I – III nude; ti I – II d, l′, lʺ, v′, vʺ; ti III d, v′, vʺ; ta I – II ft′, ftʺ, p′ζ, pʺζ, u′, uʺ, ωʺ; ta III ft′, u′, uʺ. Tarsi I and II each with one abaxial solenidion ωʺ (both 2 – 3); barbed companion seta ftʺ (4 – 5, 5, respectively); pair of prorals p′ζ, pʺζ (both 5 – 6). Colour. These mites were bright red when alive.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.taxon	biology_ecology	Host. Goodenia rosulata Domin (Goodeniaceae).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.taxon	distribution	Distribution. This mite is so far known only from Rungulla National Park in northern Queensland, Australia, where it and its host plant were collected at a single site within the park.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.taxon	etymology	Etymology. This species is named for the specific epithet of the host plant, rosulata.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565800CA019A7A1FA5FFD59FE08.taxon	discussion	Remarks. Host plant identification made by Queensland Herbarium (BRI voucher: RNP 41). Atypical ontogeny of leg setae additions: larval setae l′, l ″ on genua I – II are delayed to the deutonymph to be added with seta d; seta tcʺ delayed to adult on tarsus I – II (Table 1).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.taxon	description	(Figs 26 – 36)	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.taxon	materials_examined	Types. Holotype. Female, Australia, Serpentine Creek Historic Cemetery, Serpentine Cemetery Road, Redland Bay, Queensland, 27 ° 40 ’ 10 ” S 153 ° 18 ’ 05 ” E, ex. Goodenia rotundifolia R. Br. (Goodeniaceae), 24. ix. 2015, J. J. Beard, G. Leiper and P. I. Forster (QM). Paratypes. 6 females, 2 males, 3 deutonymphs, 2 protonymphs, 2 larvae, same data as holotype (QM). Other material examined. 1 pharate protonymph, same data as holotype (QM).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.taxon	diagnosis	Diagnosis. Female. Dorsal ornamentation comprising elongate longitudinal plicae; dorsal setae length 4 – 7. Setae g 1 positioned in transverse line with g 2. Setae ps 2 and ps 3 blunt, lengths ps 2 5 – 6, ps 3 5 – 6. Femora I – II with seta bv ″ fine blunt; femora IV with 1 seta (ev′ present, d absent; companion seta ftʺ slightly longer than solenidion ωʺ. Blunt setae on femora and tibiae: femur I – II d, l′, bv ″; femur III d; genu I – II d, l ″; tibia I – II d (1 – 2 barbs), l ″; tibia III – IV d. Most blunt setae apparently smooth.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.taxon	description	Description Adult female. (7 measured) Dorsum. (Figs 26, 27) Body measurements: v 2 – h 1 206 – 224 [221], sc 2 – sc 2 135 – 154 [142], c 3 – c 3 133 – 161 [143], f 3 – f 3 76 – 87 [84]. Prodorsal and opisthosomal shields weakly developed. Prodorsum with mostly elongate longitudinal plicae narrowly spaced, becoming mostly smooth centrally; anterior margin of prodorsum with small median forked projection forming a notch (8 – 11) [10]; with up to 4 pairs of minute pores in longitudinal band posterior to v 2 (can be difficult to see), and with pore-like invagination just mesad sc 2 (indicated by arrows in Fig. 26). Opisthosoma with complicated plicae mostly longitudinal to oblique, but almost forming weak elongate reticulation; with 3 pairs of large pores — mesad setae c 3, mesad setae d 3 and posteromesad d 3; with at least 3 pairs minute pores (can be difficult to see). Dorsal setae all short, blunt. Setal measurements: v 2 6 – 7 [6 – 7], sc 1 6 – 7 [6 – 7], sc 2 6 – 7 [6 – 7], c 1 6 – 7 [6 – 7], c 3 4 – 7 [4 – 5], d 1 5 – 6 [-], d 3 5 – 6 [5 – 6], e 3 4 – 6 [5 – 6], f 3 4 – 6 [5 – 6], h 1 4 – 6 [5]. Infracapitulum. (Figs 26, 28 A) Dorsal cuticle with fine longitudinal plicae laterally, ventral cuticle with fine oblique plicae; infracapitular setae m weakly lanceolate with 1 – 2 small barbs (11 – 12) [11]. Palp tarsus phaneres (1 – 2 and 8 – 9); femur-genu-tibia with single barbed dorsal seta (13 – 15) [13 – 14]. Venter. (Figs 28 A, 29) Ventral cuticle with longitudinal and oblique plicae on coxae I – II; with fine longitudinal plicae between 1 b – 1 a; with widely spaced transverse plicae between setae 1 a – 3 a, becoming finer 3 a – ag; with small region broadly U-shaped plicae between coxae III – IV just anterior to 4 a – 4 a; with arching plicae surrounding region of transverse plicae between 4 a – ag setae; with sparse weak transverse plicae between ag and g 1 – g 2 setae; genital region flanked by widely spaced longitudinal plicae. Setae g 1 and g 2 inserted in transverse row posteriorly on genital flap. Most ventral setae short, fine, smooth; except setae 1 a and 4 a elongate, fine, and often broken; and ps 2 and ps 3 blunt. Setae ps 3 often obscured within lateral folds of genital opening. Setal measurements: 1 a 64 – 96 [75 – 86], 1 b 18 – 20 [19], 1 c 14 – 16 [15], 2 b 18 – 21 [18 – 19], 2 c 14 – 17 [15 – 16], 3 a 13 – 17 [15], 3 b 11 – 19 [16], 4 a 41 – 56 [56], 4 b 15 – 17 [17], ag 9 – 12 [10 – 11], g 1 8 – 12 [8 – 9], g 2 9 – 13 [9], ps 2 5 – 6 [5], ps 3 5 – 6 [6]. Spermathecal apparatus. (Fig. 28 B) A moderately short membranous duct slightly broadened midway, with a narrower distal portion, terminating in a small rounded vesicle with granular appearance. Legs. (Fig. 30) See genus Diagnosis for full setal formulae for legs I – IV, except femur IV with 1 seta (ev′ present, d absent). See description of type species, G. rosulatus sp. nov., for full leg chaetotaxy. Tarsi I and II each with one abaxial solenidion ωʺ (both 5 – 6 [5 – 6, 6]); barbed companion seta ftʺ (7 – 9 [8], 8 – 9 [8], respectively) slightly longer than solenidion; pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 8 – 9). Solenidia inserted on small tubercles. Some leg setae blunt and of similar form to the dorsal idiosomal setae — seta l′ on trochanter III; setae d, l′, bv ″ on femora I – II, d on femora III; setae d, l ″ on genua I – II; setae d, l ″ on tibiae I – II and d on tibiae III – IV. Microplates. (Fig. 31) Individual microplates are hexagonal in shape (Figs 31 A, B), and have a mostly smooth surface with a few short parallel ridges visible at 10,000 X (Fig. 31 B). Adult male. (2 measured) Dorsum. (Figs 32, 33) Body measurements: v 2 – h 1 153 – 156, sc 2 – sc 2 95 – 111, c 3 – c 3 97 – 102, f 3 – f 3 55 – 57. Prodorsum similar to female, with mostly longitudinal to oblique plicae narrowly spaced, becoming mostly smooth centrally; anterior margin of prodorsum with small median forked projection forming a notch (4 – 6); with up to 4 pairs of minute pores sublaterally (can be difficult to see). Dorsal opisthosoma divided into mesonotal and pygidial shields; metanotal shield captures setal pairs c 1 – c 3 and d 1 – d 3; pygidial shield posterior to D row setae with few arching plicae anterior to smooth central region; with 3 pairs of large pores — just mesad setae c 3, mesad d 3 and posteromesad d 3; with up to 4 pairs of minute pores (can be difficult to see; arrow indicates pores in Fig. 33); with a pair of large deep pore-like invaginations just laterad d 1. Dorsal setae as in female, all short, blunt. Setal measurements: v 2 5 – 6, sc 1 6 – 7, sc 2 5 – 6, c 1 5 – 6, c 3 5 – 6, d 1 5 – 6, d 3 5, e 3 4 – 5, f 3 5 – 6, h 1 5 – 6. Infracapitulum. (Figs 32, 33, 34) Dorsal cuticle with fine longitudinal plicae, ventral cuticle with fine oblique plicae; infracapitular setae m (7 – 9) appears smooth. Palp tarsus phaneres (2 and 7 – 8); femur-genu-tibia with single barbed dorsal seta (8 – 10). Venter. (Fig. 34 A, B) Ventral cuticle with widely spaced transverse plicae from 1 a – 3 a; with region of broadly V-shaped plicae between coxae III – IV just anterior to 4 a – 4 a; with band of fine transverse plicae 4 a – ag, and with few weak transverse plicae ag – genital region; genital region often concealed under large fold of cuticle (Fig. 34 A, B), genital setae and anogenital opening flanked by plicae arching around region anteriorly. Setae g 1 inserted just anterior to g 2 on anterior margin of anogenital opening (Fig. 34 B). Most ventral setae short, fine, appear smooth; except setae 1 a and 4 a elongate, fine, and often broken; ps 2 tapered, blunt. Setae ps 3 modified into short, thick, clavate accessory genital stylets, often obscured under posterior projections of anogenital opening. Setal measurements: 1 a 68 – 78, 1 b 11 – 16, 1 c 12, 2 b 11 – 15, 2 c 11 – 12, 3 a 11 – 13, 3 b 11 – 12, 4 a 31 – 34, 4 b 12 – 13, ag 6 – 10, g 1 11 – 13, g 2 11, ps 2 5 – 7, ps 3 5. Aedeagus. (Fig. 34 C) Sclerotised, elongate, subulate (62), with narrow membranous tube emerging from sclerotised section. Legs. (Fig. 35) Setal formulae and chaetotaxy same as adult female except ta I – II with solenidion ω′ also present. Tarsi I and II each with two solenidia (ta I adaxial ω′ 7 – 8, abaxial ωʺ 7; ta II adaxial ω′ 8, abaxial ωʺ 6 – 7); companion seta ftʺ barbed (both 7); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 7 – 8). Form of leg setae similar to that of female. Deutonymph. (3 measured) Dorsum. Body measurements: v 2 – h 1 157 – 188, sc 2 – sc 2 101 – 114, c 3 – c 3 109 – 125, f 3 – f 3 47 – 58. Prodorsal and opisthosomal shields weakly developed, with smooth cuticle; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with a few broad transverse striae. Dorsal setae form as in adults. Setal measurements: v 2 3 – 5, sc 1 4 – 6, sc 2 4 – 5, c 1 4 – 5, c 3 3 – 4, d 1 3 – 5, d 3 3 – 4, e 3 3 – 5, f 3 3 – 4, h 1 3 – 4. Infracapitulum. Cuticle as in adults; infracapitular setae m (7 – 8). Palp tarsus phaneres (1 and 6 – 7); femur-genu-tibia with single barbed dorsal seta (9 – 11). Venter. Strong longitudinal plicae between 1 b – 1 a; strong transverse, finely granulate plicae 1 a to just posterior 4 a; plicae from 4 a to ag arching, becoming transverse between ag to g 1; with sparse oblique plicae surrounding genital region. Form of ventral setae as in adults. Setal measurements: 1 a 58, 1 b 14 – 18, 1 c 7 – 10, 2 b 10 – 15, 2 c 8 – 11, 3 a 8 – 13, 3 b 9 – 13, 4 a 24 – 34, 4 b 7 – 11, ag 5 – 8, g 1 6 – 8, ps 2 3, ps 3 3 – 4. Legs. Setal formulae and chaetotaxy for legs I – IV same as adult female except trochanters 1 - 1 - 1 - 0. Tarsi I and II each with one abaxial solenidion ωʺ (4 – 5, 4, respectively); barbed companion seta ftʺ (both 5 – 6); pair of prorals p′ζ, pʺζ (6 – 7, 5 – 6, respectively). Some leg setae short to minute, blunt — seta d on femora I – IV, genua I – II, tibiae I, III – IV, and seta lʺ on genua I – II and tibiae I – II. Protonymph. (2 measured) Dorsum. Body measurements: v 2 – h 1 139 – 143, sc 2 – sc 2 95 – 100, c 3 – c 3 103 – 107, f 3 – f 3 40 – 41. Prodorsum mostly smooth, with smoothly rounded anterior margin without median notch; dorsal opisthosoma with broadly spaced transverse plicae between c 1 – d 1; pygidial region between d 1 – h 1 smooth. Dorsal setae form as in adults. Setal measurements: v 2 3 – 5, sc 1 4 – 5, sc 2 4 – 5, c 1 4, c 3 3 – 4, d 1 3, d 3 3 – 4, e 3 3 – 4, f 3 3, h 1 2 – 3. Infracapitulum. Cuticle as in deutonymph; infracapitular setae m (6 – 8). Palp tarsus phaneres (1 and 5 – 6); femur-genu-tibia with single barbed dorsal seta (8). Venter. Ventral cuticle with longitudinal plicae 1 b – 1 a; transverse plicae 1 a – ps 3; small region between coxae IV with arching plicae. Setal measurements: 1 a 42 – 51, 1 b 9 – 10, 1 c 9 – 10, 2 c 9 – 10, 3 a 7 – 11, 3 b 8 – 10, ag 6 – 7, ps 2 1 – 2, ps 3 1 – 2. Legs. Setal formulae for legs I – IV: cx 2 - 1 - 1 - 0, tr 0 - 0 - 0 - 0, fe 3 - 3 - 2 - 1, ge 0 - 0 - 0 - 0, ti 5 - 5 - 3 - 3, ta 7 (1) - 7 (1) - 3 - 3 respectively. Leg chaetotaxy same as larva except 1 c, 2 c, 3 b added to cx I, II, III respectively. Tarsi I and II each with one abaxial solenidion ωʺ (both 3); barbed companion seta ftʺ (both 4 – 5); pair of prorals p′ζ, pʺζ (both 5). Larva. (2 measured) Dorsum. (Fig. 36 A) Body measurements: v 2 – h 1 107 – 123, sc 2 – sc 2 76 – 89, c 3 – c 3 83 – 92, f 3 – f 3 31 – 33. Dorsum mostly membranous with fine plicae, no shields developed but central prodorsum with thickened patterned cuticle. Prodorsum colliculate anteriorly and centrally with oblique to longitudinal plicae laterally; anterior margin smoothly rounded without median notch. Dorsal opisthosoma with fine transverse plicae between c 1 – c 1 and d 1 – d 1, and a narrow band of widely spaced transverse striae between setal rows C and D; with longitudinal to oblique plicae d 1 – h 1. Pair of large rounded pores anterior to setae e 3. Dorsal setae form as in adults. Setal measurements: v 2 4, sc 1 4 – 5, sc 2 4 – 5, c 1 4 – 6, c 3 3 – 4, d 1 3 – 4, d 3 3 – 4, e 3 3, f 3 3, h 1 3. Infracapitulum. Cuticle with fine transverse plicae; infracapitular setae m absent. Palp tarsus phaneres (1 and 5); femur-genu-tibia with single barbed dorsal seta (5 – 6). Venter. Ventral cuticle mostly finely plicate. Setal measurements: 1 a 29 – 33, 1 b 9 – 10, 3 a 8 – 9, ps 2 1, ps 3 1. Legs. (Fig. 36 B) Setal formulae for legs I – III: cx 1 - 0 - 0, tr 0 - 0 - 0, fe 3 - 3 - 2, ge 0 - 0 - 0, ti 5 - 5 - 3, ta 7 (1) - 7 (1) - 3 respectively. Leg chaetotaxy: cx I 1 b; fe I – II d, v′, bvʺ; fe III d, ev′; ge I – III nude; ti I – II d, l′, lʺ, v′, vʺ; ti III d, v′, vʺ; ta I – II ft′, ftʺ, p′ζ, pʺζ, u′, uʺ, ωʺ; ta III ft′, u′, uʺ. Tarsi I and II each with one abaxial solenidion ωʺ (both 3); barbed companion seta ftʺ (5 – 6, 6 – 7, respectively); pair of prorals p′ζ, pʺζ (both 5). Some leg setae blunt and similar in form to dorsal setae: d on fe I – III and ti III; d, l ″ ti I – II. Colour. These mites were bright red when alive.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.taxon	biology_ecology	Host. Goodenia rotundifolia R. Br. (Goodeniaceae).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.taxon	distribution	Distribution. This mite is so far known only from Redland Bay in southeast Queensland, Australia.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.taxon	etymology	Etymology. This species is posthumously named for our friend and mentor, the late Uri Gerson. Uri contributed enormously to our understanding of the Tenuipalpidae, especially so for the Australian fauna, and it is because of his direct invitation and encouragement that the senior author embarked on her journey with this fabulous family of plant-feeding mites. It was an honour to know and work with him and he is painfully missed.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658002A02EA7A1FE04FEAAF821.taxon	discussion	Remarks. Host plant identification made by Queensland Herbarium (BRI voucher: PIF 43028). Atypical ontogeny of leg setae additions: larval setae l′, l ″ on genua I – II are delayed to the deutonymph to be added with seta d; tectal setae tc′, tcʺ delayed to deutonymph on tarsus I – II (Table 1). Goodeniacarus uri sp. nov. and G. leiperi sp. nov. are found on the same host plant, but were collected approximately 160 km apart from each other. See Remarks for G. leiperi sp. nov. and the key to species for their diagnostic features.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.taxon	description	(Figs 37 – 42)	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.taxon	materials_examined	Types. Holotype. Female, Australia, Wondul Stock Route, 36 km north of Inglewood, Queensland, 28 ° 06 ’ 41 ’’ S 151 ° 04 ’ 39 ” E, ex. Goodenia glabra R. Br. (Goodeniaceae), 10. xii. 2001, J. J. Beard and P. I. Forster (UQIC 87363; paratype female on same slide). Paratypes. 4 females, 4 males, deutonymph, protonymph, same data as holotype (UQIC 87365, 87369). All material in QM.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.taxon	diagnosis	Diagnosis. Female. Dorsal ornamentation fine, irregular reticulation, becoming weaker and smoother on medial prodorsum and anteromedial opisthosoma; dorsal setae length 5 – 9. Setae g 1 in transverse line with g 2, or slightly posterior to g 2. Setae ps 2 and ps 3 short, ps 2 tapered, ps 3 tapered blunt, lengths ps 2 6 – 7, ps 3 6 – 8. Femora I with seta bv ″ long, tapered; femur II with seta bv ″ shorter, tapered blunt; femora IV with 1 seta (ev′ present, d absent; companion seta ftʺ longer than solenidion ωʺ. Blunt setae on legs: seta l′ on trochanter III; femur I – III d; genu I – II d, l ″; tibia I l ″; tibia IV d. Additional ambiguously blunt setae: tibia II d, l ″; tibia III d. Blunt leg setae can have 1 – 2 minute barbs.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.taxon	description	Description Adult female. (5 measured) Dorsum. (Fig. 37) Body measurements: v 2 – h 1 190 – 200 [194], sc 2 – sc 2 111 – 117 [113], c 3 – c 3 112 – 123 [112], f 3 – f 3 66 – 75 [70]. Prodorsal and opisthosomal shields weakly developed. Prodorsum with fine irregular reticulation and plicae becoming smoother centrally; anterior margin of prodorsum with small median forked projection forming a notch (5 – 8); with at least single pair minute pores mesad eyes (can be difficult to see). Opisthosoma with fine irregular reticulation; with broad band of sclerotised cuticle around opisthosomal shield margin, with series of strong transverse plicae; with 3 pairs of large pores — just mesad seta c 3, mesad d 3, and posteromesad d 3; with at least 3 pairs minute pores (can be difficult to see). Dorsal setae all short, fine, smooth, blunt (n. b. one female with c 1, c 3, d 3 tapered blunt and apparently with 1 – 2 barbs). Setal measurements: v 2 6 – 7 [7], sc 1 7 – 9 [8 – 9], sc 2 8 – 9 [8], c 1 7 – 9 [7], c 3 6 – 8 [6 – 7], d 1 6 – 8 [6], d 3 7 – 8 [7], e 3 5 – 8 [6 – 7], f 3 6 – 8 [6], h 1 6 – 7 [6]. Infracapitulum. (Fig. 37) Dorsal cuticle with fine longitudinal to oblique plicae; ventral cuticle smooth centrally with fine oblique striae laterally; infracapitular setae m (13 – 14) [13]. Palp tarsus phaneres (1 – 2 and 8 – 9); femur-genu-tibia with single barbed dorsal seta (13 – 15) [14]. Venter. (Fig. 38) Ventral cuticle with few fine longitudinal plicae between setae 1 b – 1 a and longitudinal to oblique plicae on coxae I – II; with transverse plicae between setae 1 a to just posterior to 3 a; with region of sparse broadly U-shaped plicae anterior to 4 a – 4 a; with fine arching plicae between 4 a – ag; genital region flanked by widely spaced longitudinal plicae; genital flap with irregular plicae between ag and genital setae. Setae g 1 and g 2 inserted in almost transverse row along posterior margin of genital flap. Most ventral setae short, fine, smooth, except setae 1 a, 4 a elongate, fine (often broken). Setal measurements: 1 a 68 – 81 [68 – 77], 1 b 20 – 27 [21], 1 c 15 – 18 [15], 2 b 20 – 23 [21], 2 c 17 – 19 [17], 3 a 17 – 22 [17], 3 b 17 – 18 [17], 4 a 64 – 67 [-], 4 b 19 – 21 [21], ag 10 – 14 [12 – 13], g 1 10 – 13 [10], g 2 11 – 13 [12], ps 2 6 – 7 [6], ps 3 6 – 8 [8]. Spermathecal apparatus. A moderately short membranous duct terminating in small bulb; basal half much broader than distal half, with a slightly enlarged bulb midway, and granulate texture basally. Legs. (Fig. 39) See genus Diagnosis for full setal formulae for legs I – IV, except femur IV with 1 seta (ev′ present, d absent). See description of type species, G. rosulatus sp. nov., for full leg chaetotaxy. Tarsi I and II each with one abaxial solenidion ωʺ (6 – 7 [6], 6 – 7 [6], respectively); barbed companion seta ftʺ (10 – 11 [11], 10 – 12 [11], respectively) longer than solenidion; pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 9 – 10). Solenidia inserted on small tubercles. Some leg setae blunt and of similar form to dorsal idiosomal setae (except with minute barbs) — seta l′ on trochanter III; seta d on femora I – III; d, l ″ on genua I – II; seta l ″ on tibiae I – II and d on tibiae III – IV. Adult male. (4 measured) Dorsum. (Fig. 40) Body measurements: v 2 – h 1 128 – 137, sc 2 – sc 2 81 – 88, c 3 – c 3 85 – 88, f 3 – f 3 51 – 55. Prodorsum somewhat similar to female, with fine irregular mostly longitudinal plicae and weak rugose elements, becoming smoother centrally; anterior margin with small median forked projection forming a notch (4 – 5); with up to 4 pairs of minute pores sublaterally (can be difficult to see). Dorsal opisthosoma divided into mesonotal and pygidial shields; mesonotal shield captures setal pairs c 1 – c 3 and d 1 – d 3; pygidial shield posterior to D row setae with few arching and oblique plicae anteriorly becoming mostly smooth posteriorly; with 2 pairs large pores — mesad setae c 3, and just anteromesad e 3; with up to 4 pairs minute pores (can be difficult to see). Dorsal setae as in female, all short, smooth, fine, blunt (some setae appear to have 1 – 2 minute barbs). Setal measurements: v 2 6 – 8, sc 1 6 – 9, sc 2 6 – 8, c 1 6 – 7, c 3 6 – 8, d 1 7, d 3 6 – 7, e 3 6 – 7, f 3 6 – 7, h 1 7. Infracapitulum. (Figs 40, 41 A) Dorsal cuticle with fine longitudinal plicae, ventral cuticle appears smooth; infracapitular setae m (8 – 11). Palp tarsus phaneres (1 – 2 and 7 – 8); femur-genu-tibia with single barbed dorsal seta (12). Venter. (Fig. 41 A, B) Ventral cuticle with fine longitudinal plicae between 1 b – 1 a; transverse to oblique plicae on coxae I – II; fine transverse plicae from 1 a – 3 a; with region of broadly V-shaped plicae anterior to 4 a – 4 a; with band of fine transverse plicae 4 a – ag, and with few weak transverse plicae ag – genital region; genital region often concealed under fold of cuticle (Fig. 41 A, B), genital setae and anogenital opening flanked by plicae arching around region anteriorly. Setae g 1 inserted just anterior to g 2 on anterior margin of anogenital opening; valves with fine longitudinal plicae; pair of large pores laterad anal opening (Fig. 41 A, B). Most ventral setae short, fine, smooth or with few minute barbs; except setae 1 a and 4 a elongate, fine, and often broken. Setae ps 3 modified into short, thick and weakly clavate accessory genital stylets, often obscured under posterior projections of anogenital opening. Setal measurements: 1 a 52 – 66, 1 b 14 – 16, 1 c 11 – 14, 2 b 12 – 17, 2 c 13 – 17, 3 a 11 – 14, 3 b 12 – 20, 4 a 35 – 39, 4 b 15 – 20, ag 10 – 12, g 1 10 – 13, g 2 11 – 13, ps 2 6 – 8, ps 3 6 – 7. Aedeagus. (Fig. 41 C) Sclerotised, elongate subulate (55 – 56), with narrow membranous tube emerging from sclerotised section. Legs. (Fig. 42) Setal formulae and chaetotaxy same as adult female except ta I – II with solenidion ω′ also present. Tarsi I and II each with two solenidia (ta I adaxial ω′ 8 – 9, abaxial ωʺ 6 – 7; ta II adaxial ω′ 8 – 9, abaxial ωʺ 6 – 7); companion seta ftʺ barbed (both 7 – 8); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 7 – 8). Form of leg setae similar to that of female, except setae l ″ on ge II and ti II tapered (not blunt). Deutonymph. (1 measured) Dorsum. Body measurements: v 2 – h 1 126, sc 2 – sc 2 86, c 3 – c 3 89, f 3 – f 3 42. Prodorsal and opisthosomal shields weakly developed. Prodorsum finely punctate with rounded central region indicated by folds; anterior margin of prodorsum smoothly rounded without median notch. Opisthosoma with broad weak plicae between c 1 – d 1; pygidial shield finely punctate. Dorsal setae as in adults. Setal measurements: v 2 3 – 5, sc 1 4 – 6, sc 2 4 – 5, c 1 4 – 5, c 3 3 – 4, d 1 3 – 5, d 3 3 – 4, e 3 3 – 5, f 3 3 – 4, h 1 3 – 4. Infracapitulum. Cuticle as in adults; infracapitular setae m (7). Palp tarsus phaneres (1 and 6); femur-genu-tibia with single barbed dorsal seta (7). Venter. Fine longitudinal plicae between 1 b – 1 a; transverse plicae 1 a – g 1; longitudinal plicae surrounding genital region. Form of ventral setae as in adults. Setal measurements: 1 a 44, 1 b 14, 1 c 10, 2 b 12, 2 c 10, 3 a 8, 3 b 9, 4 a 21, 4 b 8, ag 7, g 1 6, ps 2 3 – 4, ps 3 3 – 4. Legs. Setal formulae and chaetotaxy for legs I – IV same as adult female except trochanters 1 - 1 - 1 - 0. Tarsi I and II each with one abaxial solenidion ωʺ (both 4); barbed companion seta ftʺ (both 5 – 6); pair of prorals p′ζ, pʺζ (6 – 7). Protonymph. (1 measured) Dorsum. Body measurements: v 2 – h 1 132, sc 2 – sc 2 78, c 3 – c 3 90, f 3 – f 3 57. Prodorsum smooth, weak shield indicated with few strong longitudinal folds. Opisthosoma as in deutonymph. Setal measurements: v 2 4 – 5, sc 1 6, sc 2 5, c 1 5, c 3 4, d 1 4, d 3 4, e 3 3 – 4, f 3 3 – 4, h 1 2 – 3. Infracapitulum. Cuticle as in deutonymph. Palp tarsus phaneres (1 and 6); femur-genu-tibia with single barbed dorsal seta (7). Venter. Ventral cuticle as in deutonymph. Setal measurements: 1 a 48, 1 b 10, 1 c 10, 2 c 9, 3 a 10, 3 b 11, ag 7, ps 2 3, ps 3 4. Legs. Tarsi I and II each with one abaxial solenidion ωʺ (both 4); barbed companion seta ftʺ (both 6); pair of prorals p′ζ, pʺζ (both 5 – 6). Colour. These mites were bright red when alive.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.taxon	biology_ecology	Host. Goodenia glabra R. Br. (Goodeniaceae).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.taxon	distribution	Distribution. This mite is so far known only from Inglewood in southeast Queensland, Australia.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.taxon	etymology	Etymology. This species is named for the collection site, along the Wondul Stock Route.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C5658036A036A7A1FF65FD33FC24.taxon	discussion	Remarks. Host plant identification made by Queensland Herbarium. Goodeniacarus wondul sp. nov. was collected only 1. 3 km from, and on the same host plant as, the new genus Inglacarus. The host Goodenia glabra is an annual or perennial plant that grows after rain in sandy loamy soils in the drier inland parts of Australia, and yet this ephemeral resource is significant for these mites.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA036A7A1FC20FBFEF918.taxon	diagnosis	Diagnosis. Palps 1 - segmented, with single elongate tapered seta (apparently smooth). Anterior margin of prodorsum rounded, without median projection or notch; opisthosoma with 8 pairs of setae (c 1, c 3, d 1, d 3, e 3, f 3, h 1, h 2 present; c 2, d 2, e 1, e 2, f 2 absent); dorsal setae short, fine, blunt. Ventral and genital shields not developed, both regions entirely membranous; two pairs of ps setae, anal valves membranous, ps 2 and ps 3 inserted longitudinally on anal valves; setae 4 a short; 4 a 2 absent. Trochanter setal formula 1 - 1 - 2 - 1 (ontogeny of l′ on tr III unknown); trochanter III with seta l′ blunt. Femur III with seta d present, femur IV with seta d absent, formula 4 - 4 - 2 - 1. Genua III lacking seta l′, genual formula 3 - 3 - 0 - 0. Blunt leg setae: femur I setae d, l′; femur II d, l′, bv ″; femur III d; genu I – II d, l′, l ″; tibia I – II d, l ″; tibia III – IV d. Tarsal claws and empodia pad-like. Usually three pairs of large pores visible on dorsal opisthosoma.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA036A7A1FC20FBFEF918.taxon	discussion	Remarks. Inglacarus gen. nov. is closest to Goodeniacarus gen. nov. in sharing a reduction of idiosomal dorsal setae, having short blunt dorsal idiosomal and leg setae, undeveloped ventral shields, the same leg setation, pectinate ventral tibial and tarsal setae, the morphology of the pretarsus, and the same host plant genus. The new genus is distinguished by having two regressive features: the reduction of the palp to just one segment with one phanere (probably the tibial dorsal seta due to length, suggesting loss of tarsal phaneres concomitant with the fusion or loss of the tarsus), and loss of the median forked projection. The new genus also has seta h 2, which represents a significant plesiomorphic retention. These two genera are also distinguished from each other by the short seta 4 a in Inglacarus jonesi sp. nov. versus long, attenuated in Goodeniacarus gen. nov. While the significance of this feature is not fully understood, it seems to be consistent within most other tenuipalpid genera; although there is some intraspecific variation within genera such as Aegyptobia and Dolichotetranychus.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA032A7A1F8FCFD7BF9BA.taxon	description	(Figs 43 – 45)	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA032A7A1F8FCFD7BF9BA.taxon	materials_examined	Types. Holotype. Female, Australia, Wondul Stock Route, 36 km north of Inglewood, Queensland, 28 ° 05 ’ 59 ’’ S 151 ° 04 ’ 26 ” E, ex. Goodenia glabra (Goodeniaceae), 10. xii. 2001, J. J. Beard and P. I. Forster (UQIC 87350; 2 paratypes on same slide). Paratypes. 5 females, same data as holotype (UQIC 87347 – 87350). All material in QM.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA032A7A1F8FCFD7BF9BA.taxon	diagnosis	Diagnosis. Female. Dorsal ornamentation rugose-reticulate medially, becoming weaker to smooth laterally; dorsal setae short, length 3 – 6. Setae g 1 positioned posterior to g 2. Seta ps 2 slender, tapered to fine tip, ps 3 tapered to blunt tip. Blunt setae on femora-tibiae: femur I d, l′; femur II d, l′, bv ″; femur III d; genu I – II d, l′, l ″; tibia I – II d, l ″; tibia III – IV d. All blunt setae appear smooth.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA032A7A1F8FCFD7BF9BA.taxon	description	Description Adult female. (6 measured) Dorsum. (Fig. 43) Body measurements: v 2 – h 1 195 – 221 [217], sc 2 – sc 2 111 – 128 [128], c 3 – c 3 118 – 130 [126], f 3 – f 3 75 – 80 [77]. Prodorsal and opisthosomal shields weakly developed; prodorsum mostly reticulate to rugose; anterior margin of prodorsum smoothly rounded without median forked projection or notch; with up to 3 pairs of minute pores sublaterally (can be difficult to see). Opisthosoma with rugose reticulation over large central region, becoming weaker posteriorly and mostly smooth at lateral margins; with 3 pairs of large pores — mesad c 3, mesad d 3, posteromesad d 3; with at least 2 pairs of minute pores (can be difficult to see). Dorsal setae all short, smooth, blunt. Setal measurements: v 2 4 – 6 [5 – 6], sc 1 5 – 6 [5 – 6], sc 2 4 – 6 [4 – 5], c 1 5 [5], c 3 4 – 5 [4 – 5], d 1 4 – 5 [4 – 5], d 3 4 – 5 [4 – 5], e 3 4 – 5 [4], f 3 4 – 5 [4 – 5], h 1 3 – 5 [4], h 2 4 – 5 [4 – 5]. Infracapitulum. (Figs 43, 44) Dorsal cuticle with fine longitudinal striae; ventral cuticle smooth centrally with fine oblique striae laterally; infracapitular setae m (10 – 13) [11]. Palps one segmented, with single distal seta (11 – 14) [12 – 14]. Venter. (Fig. 44) Ventral cuticle with various patterns of fine to strong plicae; with few fine longitudinal plicae between setae 1 b – 1 a and longitudinal to oblique plicae on coxae I – II; with widely spaced transverse plicae between setae 1 a to just posterior to 3 a; with small region broadly V-shaped plicae anterior to 4 a – 4 a; with fine arching plicae between 4 a – ag; genital region flanked by widely spaced longitudinal plicae. Setae g 1 inserted just posterior to g 2 on genital flap. Most ventral setae short, fine, apparently smooth, tapered; except setae 1 a elongate, fine (often broken), and seta l′ on tr III short, blunt. Setal measurements: 1 a 68 – 76 [68 – 70], 1 b 14 – 19 [19], 1 c 13 – 18 [16], 2 b 11 – 18 [14], 2 c 14 – 17 [14 – 15], 3 a 12 – 16 [15 – 16], 3 b 13 – 16 [16], 4 a 14 – 19 [14 – 15], 4 b 11 – 13 [11], ag 9 – 14 [13], g 1 9 – 11 [10 – 11], g 2 9 – 11 [10 – 11], ps 2 7 – 9 [8 – 9], ps 3 6 – 7 [6 – 7]. Spermathecal apparatus. (Fig. 45 B) A moderately short membranous duct terminating in small bulb; basal half much broader than distal half, with a slightly enlarged bulb midway, and granulate texture basally. Legs. (Fig. 45 A) Setal formulae for legs I – IV: cx 2 - 2 - 1 - 1, tr 1 - 1 - 2 - 1, fe 4 - 4 - 2 - 1, ge 3 - 3 - 0 - 0, ti 5 - 5 - 3 - 3, ta 9 (1) - 9 (1) - 5 - 5 respectively. Leg chaetotaxy for legs I – IV: cx I 1 b, 1 c; cx II 2 b, 2 c; cx III 3 c; cx IV 4 b; tr I – II IV v′, tr III l′, v′; fe I – II d, l′, v′, bvʺ; fe III d, ev′, fe IV ev′; ge I – II d, l′, lʺ, ge III – IV nude; ti I – II d, l′, lʺ, v′, vʺ; ti III – IV d, v′, vʺ; ta I – II ftʺ, tc′, p′ζ, pʺζ, u′, uʺ, ωʺ; ta III – IV ft′, tc′, tcʺ, u′, uʺ. Tarsi I and II each with one abaxial solenidion ωʺ (7 [7], 6 – 7 [7], respectively); barbed companion seta ftʺ subequal in length to solenidion (7 – 9 [7 – 8], 7 – 8 [7 – 8], respectively); pair of prorals p′ζ, pʺζ inserted on minute cuneiform tubercles (both 7 – 8). Solenidion and setae ft′, ftʺ inserted on small tubercles. Some leg setae blunt and of similar form to dorsal idiosomal setae (some may have 1 – 2 minute barbs) — seta l′ on trochanter III; seta d, l′ on femora I – II, d on femora III; all setae genua I – II; d, l ″ on tibiae I – II and d on tibiae III – IV.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA032A7A1F8FCFD7BF9BA.taxon	etymology	Etymology. This species is named after Mr John Jones, for his generous permission and assistance with collecting on his property.	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
03F8C565802DA032A7A1F8FCFD7BF9BA.taxon	discussion	Remarks. This species was collected at the same site and on the same host species as Goodeniacarus wondul sp. nov. (see Remarks for G. wondul sp. nov.).	en	Beard, Jennifer J., Seeman, Owen D. (2025): Two new genera and six new species of flat mites (Prostigmata: Tenuipalpidae) from Goodeniaceae (Asterales) in Australia. Zootaxa 5637 (2): 201-259, DOI: 10.11646/zootaxa.5637.2.1, URL: https://doi.org/10.11646/zootaxa.5637.2.1
