identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0382470CFFCAFFFE7316FE329124FAF0.text	0382470CFFCAFFFE7316FE329124FAF0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Lophioneuridae Tillyard 1921	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Family  Lophioneuridae Tillyard, 1921</p>
            <p> Subfamily  Iotacyphinae Shcherbakov, Bashkuev et Shmakov ,  subfam.n.</p>
            <p>  TYPE GENUS.  Iotacypha Shcherbakov, Bashkuev et Shmakov ,  gen.n . </p>
            <p> DIAGNOSIS. Distinct from remaining  Lophioneuridae in elytrized, convex, finely areolate forewings and а very small ovipositor. In other lophioneurids the forewings are flat, somewhat more sclerotized than the hind wings, lacking areolate microsculpture, and the ovipositor is well developed, with cutting valvulae. </p>
            <p> COMPOSITION. Type genus and  Burmacypha Zherikhin, 2000 , both from mid-Cretaceous Burmese amber. </p>
            <p> REMARKS. The families  Lophioneuridae and  Cyphoneuridae were described from the Permian as Homoptera by Tillyard [1921] and Carpenter [1932], respectively. Tillyard [1935] merged these families, described the allied Permian family  Zoropsocidae and placed it along with  Lophioneuridae in Psocoptera. Becker-Migdisova described the genus  Lophioneurodes Becker-Midgisova, 1953 , bridging the gap between  Lophioneuridae and  Zoropsocidae , and united these two families [Becker-Midgisova, 1961]. Vishniakova [1981] retained Lophioneurinae and Zoropsocinae as subfamilies and for the first time described Mesozoic  Lophioneuridae , including two species from Late Cretaceous Taimyr amber. Jell and Duncan [1986] described the monotypic family  Edgariekiidae from the Lower Cretaceous in Psocoptera, but Ansorge [1996] synonymized it under  Lophioneuridae . Our observations confirm that there is no reason to distinguish between Lophioneurinae and Zoropsocinae even at the subfamily level. </p>
            <p> Genus  Iotacypha Shcherbakov, Bashkuev et Shmakov ,  gen.n.</p>
            <p>  TYPE SPECIES.  Iotacypha zherikhini Shcherbakov, Bashkuev et Shmakov ,  sp.n . </p>
            <p>DIAGNOSIS. Minute subbrachypterous lophioneurids. Forewings weakly elongated, strongly sclerotized, deeply convex; veins carinate, beset with strong curved setae; membrane finely areolate; Sc absent; RA long sigmoidal, leaving short R+M stem before separation of M (short RP+M stem developed); arculus (basal m-cu crossvein) between R+M and CuA; veins at wing apex Y-shaped (apical cell petiolate); discal cell bordered by 4 cells; clavus short and broad. Hind wings narrow, with long R+M+CuA stem, R and M separating distally; R, M and CuA subtransverse; clavus areolate. Antennae 7-segmented, at most 1/2 as long as forewing, scape and pedicel stout, flagellum moderately slender. Ocelli far apart, lateral ocelli close to eyes, median ocellus near clypeus. Pronotum short, saddle-shaped. Hind coxae enlarged. Tibiae with rows of setae. Tarsi 2-segmented; two slender curved claws. Genitalia very small.</p>
            <p>COMPOSITION. Monobasic.</p>
            <p> REMARKS. Similar to  Burmacypha Zherikhin, 2000 in very small size, convex areolate forewings and the hindwing venation, but in the latter genus the veins at the forewing apex are H-shaped (apical cell is not petiolate), the discal cell is bordered by 7 cells, and the antennae are at least 2/3 forewing length with a filiform flagellum. Similar to  Jantardachus Vishniakova, 1981 from Late Cretaceous Taimyr amber in very small size, 7-segmented antennae, absence of Sc and short CuA in forewings, but in the latter genus forewings are non-areolate, RA leaves the R stem after separation of M, tarsi and claws are shorter. </p>
            <p> ETYMOLOGY. From Greek iota (smallest letter of Greek alphabet; anything very small) and genus  Burmacypha ; gender feminine. </p>
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	https://treatment.plazi.org/id/0382470CFFCAFFFE7316FE329124FAF0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Shcherbakov, D. E.;Bashkuev, A. S.;Shmakov, A. S.	Shcherbakov, D. E., Bashkuev, A. S., Shmakov, A. S. (2024): A new genus of Lophioneuridae (Thysanoptera s. l.) with elytrized forewings from mid-Cretaceous Burmese amber. Russian Entomological Journal 33 (3): 272-275, DOI: 10.15298/rusentj.33.3.02, URL: https://doi.org/10.15298/rusentj.33.3.02
