identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03817A535842FFECFFA00345FAF7F9DE.text	03817A535842FFECFFA00345FAF7F9DE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplopseustis Meyrick 1884	<div><p>Diplopseustis Meyrick, 1884: 284 - 285 .</p><p>Type species:  Cymoriza minima Butler, 1881, by subsequent designation by Aurivillius, 1885, Zool. Jahresbericht, II. Abt. (1884), Arthropoda: 526</p><p>Diagnosis. The genus can be separated from  Sufetula by characters from the forewings and male and female genitalia. The forewing of  Diplopseustis moths displays small dots interspaced between the three larger spots along the costa, extended between the antemedial and postmedial lines, and the antemedian and postmedian lines are very thin. The hindwing lacks the antemedian and median markings, and the transverse line runs in the postmedian or subterminal area rather than in the median area as in  Sufetula . Vein Rs 4 of the forewing diverges from the stalk of Rs 2 and Rs 3 at least 1/4 along its distance from the cell. Veins M 2 and M 3 are stalked in both the forewing and the hindwing. Posterior of the sacci tympani are puteoli that are separated from the posterior part of sternum II by a low, narrow ridge. In the male genitalia, the uncus is very thin and elongated. The valva has various shapes but is generally longer than wide. In the female genitalia, the corpus bursae is ovoid to oblong, without a signum.</p><p>Distribution and diversity.  Diplopseustis is currently known from the Oriental, Australasian (Australia, New Guinea) and Afrotropical regions. The genus currently includes seven species:  Diplopseustis brunnealis (Hampson, 1917) comb. nov.,  D. constellata Warren, 1896,  D. nigerialis Hampson, 1906,  D. pallidalis Warren, 1896,  D. perieresalis (Walker, 1859),  D. prophetica Meyrick, 1887, and  D. selenalis Hampson, 1906 .  Diplopseustis brunnealis is the only species currently known from the Philippines.</p><p>Phylogenetic relationships.  Diplopseustis was recovered as monophyletic together with  Sufetula in Léger et al. (2021). The two  Sufetula species S. diminutalis (Walker, 1866) from Central America and  S. sythof fi (Snellen, 1899) from South-East Asia included in the phylogenetic analysis of Léger et al. (2021) cluster together.</p></div>	https://treatment.plazi.org/id/03817A535842FFECFFA00345FAF7F9DE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535842FFEBFCDD0011FF73F9FE.text	03817A535842FFEBFCDD0011FF73F9FE.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Diplopseustis brunnealis (Hampson 1917) Müller & Hayden & Lees & Léger 2025	<div><p>Diplopseustis brunnealis (Hampson,1917),  comb. nov. (Figures 3, 19, 32, 42)</p><p>Sufetula brunnealis Hampson, 1917: 373</p><p>Type locality. Philippines, Manila</p><p>Material examined.   Holotype. ♀ (specimen identifier: NHMUK013699637): “Type H.T.” [red circled label] | “ Philippines near Manila. E.W. Ledyard 1912 - 97” | “  Sufetula brunnealis type ♀ Hampsn”;  3 ♂ (specimen identifiers coll.mfn-berlin.de_u_2a6f6a, 2a6f68, 2a6f69), 1 ♀ (specimen identifier coll.mfn-berlin.- de_u_2a6f6b). Detailed information in Table S1 .</p><p>Similar species.  Diplopseustis selenalis Hampson, 1906</p><p>Diagnosis.  Diplopseustis brunnealis comb. nov. (Figure 3) displays several characters that are characteristic of  Diplopseustis: 4 - 5 smaller costal spots are inserted interstitially between the three primary costal spots; the ante- and postmedian lines are thin, cream to white; the postmedian line is abruptly shifted inwards at 2/3 length, then running almost straight to the dorsum; and the hindwing has a thin, white, jagged postmedian line. In the male genitalia (Figure 32), the uncus is very thin and long, with the apex split in two. The valva is twice as long as wide. In the female genitalia (Figure 42),  D. brunnealis comb. nov. can be separated from the similar looking species  D. perieresalis (type locality: Malaysia, Sarawak) and  D. selenalis (type locality: New Guinea, Fergusson Island) by the shape of the antrum and of the corpus bursae: the antrum forms two semi-rounded sclerites and the corpus bursae is oblong in  D. brunnealis comb. nov., while it forms a sclerotised ring in  D. selenalis and in  D. perieresalis, and the corpus bursae is globular in  D. perieresalis (Speidel et al. 2007) .</p><p>Redescription. Wings (Figure 3). Forewing length: 6.5 - 7mm; background colour dark brown. Antemedian line jagged, cream-coloured, distally edged with dark brown. Discoidal spot oblong, cream, basally edged with dark brown. Costa with three distinctive subtriangular cream spots, in-between chequered with dark brown and cream. Postmedian line thin, cream-coloured, slightly arched outwards on first half, then disrupted inwards on 3/4, last 1/4 running vertically toward dorsum. Tympanal organs (Figure 19). ♂ ♀. Bullae tympani 1/4 to 1/3 recessed laterad of venula prima, not extended posterior of ramus tympani. Tympanic pockets small, more or less marked. Venulae secundae long, straight. Male genitalia (Figure 32). Uncus roughly as long as tegumen arms, conspicuously narrow, apex split in two tips. Valva twice as long as wide, oval shaped, ventral edge conspicuously convex, dorsal edge straight, apex rounded. Juxta with V-shaped basal margin, deeply split in middle. Manica with two combs of dense sclerotised setae. Phallus short, bulky; cornuti absent. Female genitalia (Figure 42). Posterior apophyses ca 2.3 × tergite VIII length. Anterior apophyses ca 1.9 × tergite VIII length. Antrum with two semi-rounded sclerites surrounding opening. Ductus bursae roughly straight, in distal half covered with minute spicules. Corpus bursae ca 1.5 × ductus length, oblong-shaped, kinked near opening, membranous, devoid of signa.</p><p>Distribution. Known from Luzon (Manilla, Mt Makiling), Negros (Patag) at elevations between 200 m and 750 m (Figure 55).</p><p>DNA barcoding. The specimen PYRALPHIL09-F11 from Negros shows a p-distance of 2.0% with the specimen MFNLEP937 (Negros, same locality) and the specimen MFNLEP949 from Luzon (Laguna, Mt Makiling).</p></div>	https://treatment.plazi.org/id/03817A535842FFEBFCDD0011FF73F9FE	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535845FFEAFF8900F1FE1FFB2D.text	03817A535845FFEAFF8900F1FE1FFB2D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula Walker 1859	<div><p>Sufetula Walker, 1859</p><p>Type species:  Sufetula sunidesalis Walker, 1859, by monotypy</p><p>-  Loetrina Walker, 1863: 132 . Type species:  Loetrina flexalis Walker, 1863, by monotypy. Hampson 1896: 252 (syn.)</p><p>-  Mirobriga Walker, 1863: 131-132 . Type species:  Mirobriga albicans Walker, 1863, by monotypy. Hampson 1896: 252 (syn.)</p><p>-  Nannomorpha Turner, 1908: 80 . Type species:  Nannomorpha alychnopa Turner, 1908, by monotypy. Munroe 1995 a: 10, 76 (syn.)</p><p>-  Perforadix Seín, 1930: 168 . Type species:  Perforadix sacchari Seín, 1930, by original designation. Solis &amp; Shaffer 1999: 7 (syn.)</p><p>-  Pseudochoreutes Snellen, 1880: 202 . Type species:  Pseudochoreutes choreutalis Snellen, 1880, by monotypy. Hampson 1896: 252 (syn.)</p><p>-  Pseudochoreutis Hampson, 1899: 613</p><p>-  Safetula Hampson, 1893: 44, 161</p><p>Diagnosis.  Sufetula displays the following characters: the labial palpi are more upwardly curved than those of other  Lathrotelinae, with a second segment that is ascendant and a third segment in the same arc, whereas other lathrotelines have a straight, porrect second segment and the third segment turned up at an obtuse angle. The postmedian line of the wings is very curved, with a distinctive curve outward near the costa and often a darker area at the apex (Figure 1). All have a dark discoidal spot and two to four spots on the medial area of the costa between the antemedian and the postmedial line (Figure 1). In the forewing, Rs 4 either diverges from the cell at the same point as the stalk of Rs 2+ 3 or shares a very short stalk with Rs 2+ 3, diverging less than 1/10 the length of Rs 2+ 3 (Figure 2). In some but not all species of  Sufetula, the forewing has a tubular CuP vein. In the tympanal organs, the bullae tympani protrude into the body cavity below the level of the adjacent sternite. The male eighth sternum is medially membranous. The male genitalia have a manica with two distinctive hairy lobes and a valva with a more or less pronounced demarcated margin, like a membranous bumper. The uncus is present and variably reduced, not long and thin nor completely absent. The caecum penis is more than half the length of the phallus, with the ductus ejaculatorius inserted near the distal apex of the phallus. The female genitalia have papillae anales fused in a narrow triangle, and the Asian species of  Sufetula have an ovate signum in the corpus bursae consisting of erect papillae.</p><p>Description. Wings. Forewing: Sc, R 1, Rs 1 separate; Rs 2 and Rs 3 long-stalked; Rs 4 connate with Rs 2+ 3 or very short-stalked with it; M 1 separate, M 2 and M 3 connate, not stalked; CuA 1 and CuA 2 separate; CuP present in some species, extended from the margin 1/ 3 to 1/2 toward the base of the wing before attenuating; 1A+2A present, not forked at base; 3A separate, not looped with 1A+2A. Hind wing: Rs 1 anastomosed briefly with Sc+R 1; M 1 connate with base of Rs 1 at corner of cell; M 2 and M 3 connate, not stalked; CuA 1 and CuA 2 present, separate; CuP, 1A+2A, and 3A present, separate. Tympanal organs. Bullae tympani recessed into body cavity, not lying flush with surface; saccus tympani reduced, broader than wide, in some species sexually dimorphic, males having a rounder, deeper, and more clearly defined depression than females. Pons tympani deeply indented in females, indentation fused in males. Male genitalia. Uncus present, rather simple, narrow, apex pointed or rounded, some species with few sparse setae. Gnathos missing. Juxta elongate, slender, deeply indented. Manica of two combs of densely sclerotised chaetae. Phallus long, slender, distally covered with tiny spicules, cornuti absent to multiple, distal half with marked Y-shaped sclerotisation; caecum penis longer than half the length of phallus, ductus ejaculatorius inserted around apical 1/4. Female genitalia. Papillae anales fused in acute triangle, longer than wide; ostium not armed; colliculum present, simple or with pair of low lobes; ductus bursae without distinct swellings or diverticula, without sclerotisation or with fine granules not aggregated; corpus bursae spheroidal, reticulate, without signum (Old- and New-World species) or with round or oblong signum (Old-World species).</p><p>Distribution and diversity.  Sufetula includes 30 described species distributed over the whole tropical region. Species reported from Europe are the result of opportunistic introduction with ornamental plant trade.</p><p>Phylogenetic relationships.  Sufetula is recovered as sister to  Diplopseustis in Léger et al. (2019).</p></div>	https://treatment.plazi.org/id/03817A535845FFEAFF8900F1FE1FFB2D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535844FFEAFFA002A3FC13F894.text	03817A535844FFEAFFA002A3FC13F894.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula paula (West 1931) Müller & Hayden & Lees & Léger 2025	<div><p>Sufetula paula (West, 1931), comb. nov.</p><p>(Figures 4, 20, 33, 43)</p><p>Orthoraphis paula West, 1931</p><p>Type locality. Philippines Is., Luzon I., subprov. Benguet Pauai, Haight’ s Place, 7000 f[ee]t.</p><p>Material examined.   Holotype. ♀ (specimen identifier NHMUK013699662): “Type” [red circled] |  “Haight’ s Place, Pauai, subprov. Benguet, Luzon, 7,000 ft. 27. Vi. 1913 A.E. Wileman ” | “  Orthoraphis paula West Holotype ♀” | “Wileman Coll. B.M. 1929 - 261”;  3 ♂ (specimen identifiers coll.mfn-berlin.de_u_2a6f67, 2a6f65, d7744e), 3 ♀ (specimen identifiers coll.mfn-berlin.- de_u_2a6f66, 2a6f64, a64977). Detailed information in Table S1 .</p><p>Diagnosis.  Sufetula paula comb. nov. (Figure 4) is a rather large species (forewing length 10 - 13 mm), with forewing much wider than observed in other species of  Sufetula . The jagged antemedian line, the conspicuous discoidal black spot surrounded with white, the four conspicuous cream-coloured costal spots and the only faintly curved postmedian line separate this species from other  Lathrotelinae species from the Philippines. In the male genitalia (Figure 33), the shape of the uncus and the valva separate  Sufetula paula comb. nov. from other  Sufetula species. The female genitalia (Figure 43) have a comparatively small corpus bursae to the rest of the species and are without erect papillae.</p><p>Redescription. Wings (Figure 4). Forewing length: 10 - 13 mm; forewing ground colour tawny to brown. Basal spot round, dark brown. Antemedian line jagged, white with distal edge dark brown. Discoidal spot black, surrounded with white. Costa with four pale yellow median spots; distal one abutted to postmedian line. Postmedian line thin, white, faintly edged with dark brown, waved, running vertically down to half length, then bent inwards. Fringes basally dark brown, distally stripped with dark brown and white. Hindwings dirty white with faintly marked median dark brown line, subterminal area speckled with brown scales. Tympanal organs (Figure 20). ♂ ♀. Bullae tympani slightly recessed laterad, recessed posteriad by 1/6 beyond ramus tympani. Posterior depressions as wide as bullae, nearly square (as long as wide), not sexually dimorphic. Venulae secundae long, straight. Male genitalia (Figure 33). Uncus short, narrowing at basal 1/4, apex with rounded tip. Valva elliptical, dorsal margin conspicuously convex on basal half. Juxta forming two bumps on basal edge, distal half narrowing, apex rounded. Manica with two combs of dense sclerotised setae. Phallus elongated, lightly sclerotised, with sclerotised dent at apex. Female genitalia (Figure 43). Papillae anales fused, narrowing to tip. Posterior apophyses thin, ca 1.5 × tergite VIII length, apex spatulate. Anterior apophyses roughly as long as tergite VIII length, apex spatulate. Colliculum forming short sclerotised ring. Ductus bursae very long, thin, covered with minute spicules, distal 1/3 widening toward corpus opening. Corpus bursae globular, roughly 1/3 of ductus length, with large oval patch of short thorns surrounded with thin sclerotised spines.</p><p>Distribution. Known from Mount Tabayoc on Luzon at an elevation of 2350 m and Pauai, Haight’ s Place, Benguet, Luzon at ca. 2100 m as well as N. Luzon ca. 1500 - 1800 m (Figure 55).</p><p>DNA barcoding. An intraspecific divergence of 0.3 % was found among two specimens from the same locality (Luzon, Benguet, Mt Tabayoc).</p><p>Remarks. The species was identified with the holotype specimen from the NHMUK (Figure 17). This species is transferred to  Sufetula on the basis of maculation and venation.  Orthoraphis obfuscata, the type species of  Orthoraphis, has numerous intercalary spots along the forewing costa (shared with  Diplopseustis); one radial vein of the forewing is absent; and M 2 and M 3 are stalked in the forewing and hindwing. In contrast, species of  Sufetula have no extra spots along the forewing costa (only two or three large lunules), all five radial veins are present, and M 2 and M 3 are separate in both wings.  Sufetula paula comb. nov. likewise lacks intercalary spots, has five distinct radial veins, and M 2 and M 3 are separate in the forewing. The stalked M 2 and M 3 in the hindwing is explained as an autapomorphy independently derived from  Orthoraphis and  Diplopseustis .</p><p>In  Sufetula, Rs 4 is connate with the base of Rs 2 +Rs 3 or short-stalked about 1/12 to 1/10 of their length, whereas in  Diplopseustis these veins are stalked to about 1/3 of their length (Meyrick, 1884; Hampson, 1896). In all the specimens investigated in this study, Rs 4 is short-stalked with the long-stalked Rs 2 +Rs 3 (Figure 1). This is consistent with the observations of  Sufetula sacchari (Seín),  S. carbonalis, and  S. anania (Seín 1930, Hayden, 2013; Solis et al. 2019). Seín (1930) illustrated the intraspecific variation in the length of the short stalk of Rs 4 (p. 167 and Figure 20). Lastly, obvious differences of the wing pattern are found along the costa with the costal median spots. Nearly all the species show two to three relatively large costal median spots, suggesting another possible apomorphy for  Sufetula .</p></div>	https://treatment.plazi.org/id/03817A535844FFEAFFA002A3FC13F894	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535844FFF5FCDD015BFACFFEA8.text	03817A535844FFF5FCDD015BFACFFEA8.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula choreutalis (Snellen 1880)	<div><p>Sufetula choreutalis (Snellen, 1880)</p><p>(Figures 5, 21, 34, 45)</p><p>Pseudochoreutes choreutalis Snellen, 1880: 202 - 203 . Type locality: Indonesia, Sulawesi [- Celebes]: Makassar, Saleyer</p><p>-  Sufetula chloreutalis Whalley, 1962: 108</p><p>Material examined.  7 ♂ (specimen identifiers coll.mfn-berlin.- de_u_de00bb, b4b0fe, 2a6f59, 2a6f5a, 2a6f5b, 2a6f5c, e80bee), 3 ♀ (specimen identifiers coll.mfn-berlin.de_u_2a6f58, 6d7f3a, 430abf), 2 unsexed (specimen identifiers coll.mfn-berlin.de_u_87a5c0, 73609d). Detailed information in Table S1.</p><p>Similar species.  Sufetula sunidesalis Walker, 1859,  Sufetula gigantea sp. nov.</p><p>Diagnosis. This species (Figure 5) is superficially similar to  Sufetula gigantea sp. nov., but its wing length of 5 - 6 mm is conspicuously smaller than in  S. gigantea sp. nov. (wing length - 9.5 - 12 mm). The brown nuclei of the costal spots of the forewing as well as the continuous submarginal lines on the fore- and hindwing further separates  S. choreutalis from  S. gigantea sp. nov. In the male genitalia, the short, tapering uncus and the quadrangular shape of the valva with a pointed apex separates this species from other  Sufetula species. In the female genitalia (Figure 45), the bulge at the posterior end of the ductus bursae with irregular sclerotisation is unique to this species.</p><p>Description. Wings (Figure 5). Forewing length: 9.5 - 12.3 mm; forewing ground colour brown. Antemedian area with two oblique cream stripes. Antemedian line cream with distal edge dark brown, slightly arched outwards, forming three spikes pointing distally. Costa with three conspicuous cream spots with innerside dark brown, the third one fused with postmedian line. Discoidal spot rather large, round, black. Postmedian line cream, curved outwards on basal 1/3, abruptly angled at 1/3, running inwardly; third 1/4 running downwards towards dorsum, abruptly angled at 3/4, running outwardly, then down to dorsum. Subterminal area with dark brown patch at apex, with thin, dark brown submarginal line. Wing margin deeply notched at M 1 -M 2. Fringes basally cream, distally brown. Hindwing basally cream speckled with dark brown, with antemedian patch of dark brown. Median line jagged, cream-coloured. Distal part light to dark brown, with thin, dark brown submarginal line. Forewing CuP vein present. Tympanal organs (Figure 21). ♂ ♀. Bullae tympani with lateral fourth recessed lateral of venula prima. Saccus tympani small, wider than long, deeper and more demarcated in male, opened ventrad. Venulae secundae short, kinked twice, extending to midlength of sternum II. Male genitalia (Figure 34). Uncus ca 2/3 of tegumen arm length, slender, slightly enlarging toward middle, with scattered hairs on lateral edges, tip pointed. Valva quadrangular, innerside of ventral margin with tuft of dense hairs; ventral part protruding distally into rounded projection. Phallus with one small cornutus, phallus apex sclerotised, notched. Female genitalia (Figure 45). Posterior apophyses ca 2.25 × tergite VIII length. Anterior apophyses ca 2 × tergite VIII length, at basal 1/5 with both ventral and dorsal short bump. Colliculum ca 1/4 of tergite VIII length, ring-shaped. Ductus bursae with small, round, irregular sclerite anterior of colliculum and ductus seminalis, posterior 3/4 of ductus bursae membranous, anterior 1/4 lined with spicules near corpus opening. Corpus bursae subequal to length of ductus, oval-shaped, with posterior area covered with short erected papillae; anterior area covered with erected acanthae. Signum ca half the corpus length, forming an oblong patch of short, sclerotised, dent-shaped thorns.</p><p>Distribution. INDONESIA: Sulawesi, Selayer [Salayer], 1,600 ft.; PAPUA NEW GUINEA: Rossel I.; Astrolabe Bay; Alor; Talaut; Sariba I.; Witu - French Is.; Hydrographer Mts, 2000 ft.; St Matthias I.; Fergusson Is.; Milne Bay; Neu Pommern [New Britain]; Feni I, E. of New Ireland; Key I.; Little Kei; Squally Is.; Rossel Is; Sangir; INDONESIA: Tenimber [Tanimbar}; SOLOMONS: Rennel Is.; Bellona I, Matahenua, Hutuna; Vella Lavella; Nissan I.; S. side of Choiseul Is.; Bougainville; PHILIPPINES. Leyte, Luzon (Quezon), Mindanao (90 - 216m); Los Banos; Kolambugan, Lano, plains, Mindanao (Figure 55); SAMOA: Apia, Upolu.</p><p>DNA barcoding. An intraspecific variation of 0.3 % was found between the specimen MFNLEP970 from Leyte and the BOLD record LNAUW 3158-17 from Luzon (Laguna, Mt Makiling).</p><p>Remarks. This species is sexually dimorphic, with females exhibiting wing length 1 - 2 mm longer than males.</p></div>	https://treatment.plazi.org/id/03817A535844FFF5FCDD015BFACFFEA8	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A53585BFFF4FCC7072CFE64FE5F.text	03817A53585BFFF4FCC7072CFE64FE5F.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula gigantea Muller, Hayden & Leger 2025	<div><p>Sufetula gigantea Müller, Hayden &amp; Léger,  sp. nov.</p><p>LSID urn:lsid:zoobank.org:act: 1C91E835-808D-478A-B0C5 - 0206262B1ADB</p><p>(Figures 6, 22, 44)</p><p>Material examined.   Holotype. ♀. PHILIPPINES: Negros,  Patag,  Lake Danao, 9.35 ° N 123.18 ° E; 1400m; 21 May 1996; W. Mey leg.; DNA voucher MFNLEP-PYRALPHIL09-B10; genitalia slide MfN gen. prep. 302F; specimen identifier coll.mfn-berlin.de_u_2a6f56 (MfN).  Paratypes. 6 ♀; specimen identifiers coll.mfn-berlin.de_u_2a6f55, 2a6f57, 7816dc, 5a289e, 74ed33, d7611e; 2 unsexed; specimen identifiers coll.mfn-berlin.de_u_339649, e554a3; 2 ♀; specimen identifiers NHMUK015666225, NHMUK015111017. Detailed information in Table S1 .</p><p>Similar species.  Sufetula choreutalis,  S. sythof fi.</p><p>Diagnosis. This species (Figure 6) is the largest  Lathrotelinae species found on the Philippines. It is superficially similar to  Sufetula choreutalis, but its wing length of 9.5 - 12.5 mm is conspicuously longer than that of  S. choreutalis (wing length - 5 - 6 mm). The uniformly coloured cream costal spots without dark centers and the dashed marginal lines of the fore- and hindwing further separates this species from  Sufetula choreutalis . In the female genitalia (Figure 45), the light sclerotisation on the posterior 1/3 of the ductus bursae separates this species from  S. choreutalis . Male specimens are unknown.</p><p>Description. Wings (Figure 6). Forewing length: 9.5 - 12.5 mm; ground colour brown. Antemedian line pale yellow, thinly edged with dark brown distally; slightly arched outwards, conspicuously indented outwards twice at 1/3 and 2/3. Costa with two conspicuous cream spots. Discoidal spot rather large, round, black. Postmedian line cream, curved outwards around apical spot around medial veins, then straight basad along M 2 -M 3; perpendicular towards dorsum from M 3 to CuP; straight distad on CuP; perpendicular to dorsum between CuP and 1A+2A, then obliquely basad to dorsum. Subterminal area suffused with round, dark brown spot at apex and indent of postmedian line between M 3 and CuP, otherwise cream; submarginal line thin, dashed with dark brown. Fringes pale brown. Hindwing cream, faintly speckled with dark brown. Median line slightly waved, cream-coloured with basal edge dark brown. Postmedian dark brown patch at apex. Submarginal line thin, dashed with dark brown. Fringes pale brown. Forewing CuP vein present. Tympanal organs (Figure 22). ♀. Bullae tympani with lateral fourth recessed lateral of venula prima. Saccus tympani shallow, small, less than half width of bulla, not observed in male. Venulae secundae incurved, reaching posterior 1/3 of sternum. Male genitalia. Not known. Female Genitalia (Figure 44). Posterior and anterior apophyses very similar to  Sufetula choreutalis . Colliculum cylindrical, sclerotised, at anterior edge with transverse ridges. Ductus bursae straight, with light sclerotisation on posterior 1/3, covered with spicules near corpus opening. Corpus bursae ca 1.4 × ductus length, oval, reticulated with erected papillae at anterior fourth. Signum 3/5 of corpus length, forming oblong patch of short, tooth-shaped thorns.</p><p>Distribution. Known from Negros (Patag), Mindoro (Mt Halcon, Mt Dulangan) and Mindanao (Davao Oriental, Mt Agtuuganon) at an elevation between 1050 m and 1675 m (Figure 55).</p><p>Etymology. From the Latin giganteus, a, um, “from the Giants”, referring to the large size of the moths.</p><p>DNA barcoding. The specimen with DNA voucher MFNLEP-PYR-ALPHIL-C10 from Mindanao differs from the other two specimens from Negros by a p-distance of 3.5%.</p><p>Remarks. The unique specimen on hand from Mindanao (coll.mfn-berlin.de_u_2a6f57) shows substantially smaller forewings (9.5 mm), a less marked postmedian line and slightly different colour when compared to the specimen of Negros. The female genitalia show minor differences. Investigation of additional material will be essential to decide whether or not the Mindanao specimen represents a different species.</p></div>	https://treatment.plazi.org/id/03817A53585BFFF4FCC7072CFE64FE5F	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A53585AFFF4FFA00790FC07FE3B.text	03817A53585AFFF4FFA00790FC07FE3B.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula fulgurata Muller, Hayden & Leger 2025	<div><p>Sufetula fulgurata Müller, Hayden &amp; Léger,  sp. nov.</p><p>LSID urn:lsid:zoobank.org:act: 8DFC4324-BFA0-49AE-BEFE-03D6A6 29713A</p><p>(Figures 7, 8, 23, 35, 46)</p><p>Material examined.   Holotype. ♂. PHILIPPINES: Luzon, Zambales, Zambales Mountains,  Coto; 15.46 ° N 120.05 ° E; 150m; 09 - 10 Nov. 1998; W. Mey leg.; DNA voucher MFNLEP943; genitalia slide MfN gen. prep. 334M; specimen identifier coll.mfn-berlin.de_u_b416f 6 (MfN).  Paratypes. 16 ♂; specimen identifiers coll.mfn-berlin.- de_u_58ebd 1, 25d228, d1cfde, 6d0a7e, a70683, 2db249, 869146, 5738f7, 62093c, 15b3d2, a31b7e, f971ee, 5708ef, 1f1859, 2a6f6d, 2a6f6e; 4 ♀; specimen identifiers coll.mfn-berlin.de_u_18dc19, 1ff9c0, 9d8864, 205b59. Detailed information in Table S1. Other specimens examined. 5 unsexed (specimen identifiers coll.mfn-berlin.de_u_e89e14, 901ad4, b6c087, 18dc18, 56cc79). Detailed information in Table S1 .</p><p>Similar species.  Sufetula elfridea sp. nov.,  Sufetula serrata sp. nov.</p><p>Diagnosis.  Sufetula fulgurata sp. nov. (Figures 7, 8) is distinguished from  S. elfridea sp. nov. by the postmedian line jagged on the anal fold (straight in  S. elfridea sp. nov.). From  Sufetula serrata sp. nov., it is separated by examination of genitalia: the narrow uncus has a pointed apex, and the valva is tapered to a pointed apex (Figure 35), and in the female genitalia (Figure 46), the colliculum forms a narrow sclerotised ring not reaching the posterior margin.</p><p>Description. Wings (Figures 7, 8). Male forewing length: 4.5 mm (Figure 7); colour generally pale yellow to fuscous. Antemedian area dark brown. Antemedian line thick, white, thinly edged with dark brown distally, conspicuously curved outwards on costal 1/3, then jagged toward dorsum. Costa with three oblique spots pointing towards margin, white, thinly edged with dark brown. Discoidal spot oval, dark brown. Thick, white postmedian line curved from costa to acute point between veins M 1 and M 2, then bent inwards, with marked indentation on anal fold. Subterminal area with subapical dark brown patch. Fringes pale yellow and brown. Hindwing pale yellow, thinly speckled with dark brown towards dorsum. Median line thick, white, fading toward costa, shifted inwardly at 2A vein. Postmedian area with prominent dark brown area in dorsal half. Female forewing length: 7 mm (Figure 8); as male, but markedly suffused with dark brown. Tympanal organs (Figure 23). ♂. Bullae tympani slightly recessed laterad, deeply recessed posteriad, extended by 2/5 posterior of ramus tympani. Saccus tympani small, very shallow, poorly differentiated. Venulae secundae faintly marked, medially incurved, running down to anterior 1/3 of sternum II. Male genitalia (Figure 35). Uncus ca half of tegumen arm length, narrow, triangular, apex pointed. Valva ovate, apex narrowed into a pointed tip. Phallus long, slender, with minute denticles distally; cornuti absent. Female genitalia (Figure 46). Posterior apophyses ca 2.5 × tergite VIII length. Anterior apophyses 2 - 2.6 × tergite VIII length; slightly thickened at posterior 1/3 of both apophyses. Colliculum forming narrow sclerotised ring of ca 1/10 of tergite VIII length, not reaching antrum opening. Ductus bursae ca 2.1 × corpus bursae length, narrow, straight. Corpus bursae globular with some short bristles in posterior area. Signum 1/5 of corpus length, round, with short, tooth-shaped thorns.</p><p>Distribution. Known from Luzon (Zambales, Benguet) at an elevation between 150 and 750 m (Figure 55).</p><p>Etymology. From the Latin fulguratus, a, um “struck by lightning”, referring to the zigzagging postmedian line of the forewing.</p><p>DNA barcoding. An intraspecific variation of 0.3% was found among specimens from the same locality (Zambales, Coto, at 150 m elevation).</p></div>	https://treatment.plazi.org/id/03817A53585AFFF4FFA00790FC07FE3B	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A53585AFFF7FCDD07BCFC98FE67.text	03817A53585AFFF7FCDD07BCFC98FE67.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula elfridea Muller, Hayden & Leger 2025	<div><p>Sufetula elfridea Müller, Hayden &amp; Léger,  sp. nov.</p><p>LSID urn:lsid:zoobank.org:act: 8DCE7F7E-32A0-412 B-B 40C-E77F0 A806664</p><p>(Figures 9, 24, 36, 47)</p><p>Material examined.   Holotype. ♀. PHILIPPINES: Luzon, Nueva Vizcaya, Bald Mountains,  Santa Fe; 16.18 ° N 120.86 ° E; 1150m; 11-13 Nov.1997; K. Ebert, W. Mey, M. Nuss leg.; DNA voucher MFNLEP942; genitalia slide MfN gen. prep. 333F; coll.mfn-berlin.de_u_2a6f6f (MfN).  Paratypes. 3 ♂; specimen identifiers coll.mfn-berlin.- de_u_2a6f70, dc8da3, f0e838. Detailed information in Table S1 .</p><p>Similar species.  Sufetula fulgurata sp. nov.,  Sufetula serrata sp. nov.</p><p>Diagnosis. The wings of  Sufetula elfridea sp. nov. (Figure 9) are very similar to  Sufetula fulgurata sp. nov. In the forewing, the postmedian line is projected outwards into a tip between M 1 and M 2, and then runs inwards into a slightly wavy line to the dorsum, lacking the indentation observed in  S. fulgurata sp. nov. near the dorsum. In the male genitalia (Figure 36), the valva apex is rounded, while it is pointed in  Sufetula fulgurata sp. nov. In the female genitalia (Figure 47), the anterior apophyses are slightly longer than in  Sufetula fulgurata sp. nov. and  Sufetula serrata sp. nov., and the colliculum forms a sclerotised ring that reaches the posterior edge of the antrum, while it does not reach the posterior edge in  S. fulgurata sp. nov.</p><p>Description</p><p>Wings (Figure 9).</p><p>Male forewing length: 9 mm; colour generally pale yellow to fuscous. Antemedian area dark brown. Antemedian line thick, white, thinly edged with dark brown distally, gently arched outwards on costal 1/3, then waving toward dorsum. Costa with three oblique triangular spots pointing towards margin, white, thinly edged with dark brown. Discoidal spot oval, dark brown. Postmedian line white, curved from costa to acute point between veins M 1 and M 2, then bent inwards, indentation on anal fold faintly marked. Subterminal area with subapical dark brown patch. Fringes pale yellow and brown. Hindwing pale yellow, thinly speckled with dark brown towards dorsum. Median line thick, white, fading toward costa, shifted inwardly at 2A vein. Postmedian area with prominent dark brown area in dorsal half.</p><p>Tympanal organs (Figure 24). ♂. Bullae tympani slightly recessed lateral of venula prima, deeply recessed by 1/4 to 1/3 posterior of ramus tympani. Venulae secundae faintly marked, running down to anterior 1/3 of sternum II. ♀. Posterior depressions almost as wide as bullae tympani, shallow, not sexually dimorphic. Bullae tympani completely opened ventrally in female. Venulae secundae conspicuously marked, running down to tergite II anterior margin.</p><p>Male genitalia (Figure 36). Uncus ca half of tegumen arm length, narrow, with rounded apex. Valva subquandrangular, apex with a dorsal bump. Phallus long, slender, with minute denticles distally, without cornuti.</p><p>Female genitalia (Figure 47). Posterior apophyses ca 3 × tergite VIII length. Anterior apophyses 2.4 × tergite VIII length; slightly thickened at posterior 1/3 of both apophyses. Colliculum forming narrow sclerotised ring of ca 1/10 of tergite VIII length, reaching the posterior edge of the antrum. Ductus bursae ca 2.9 × corpus bursae length, narrow, straight. Corpus bursae globular with some short bristles posterior. Signum 1/4 of corpus length, rounded, with short, tooth-shaped thorns.</p><p>Etymology. This species is dedicated to Elfriede Anni Wollschläger, the grandmother of the first author.</p><p>Distribution. Known from Luzon (Ifugao, Nueva Vizcaya) at an elevation between 1150 and 1700 m (Figure 55).</p><p>DNA barcoding. The three specimens investigated from Nueva Vizcaya display identical DNA barcodes. The nearest non-conspecific neighbour is  Sufetula fulgurata sp. nov. with a p-distance of 3%.</p></div>	https://treatment.plazi.org/id/03817A53585AFFF7FCDD07BCFC98FE67	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535859FFF7FF890469FBB1FE43.text	03817A535859FFF7FF890469FBB1FE43.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula serrata Muller, Hayden & Leger 2025	<div><p>Sufetula serrata Müller, Hayden &amp; Léger,  sp. nov.</p><p>LSID urn:lsid:zoobank.org:act: D3B46F08-E10E-49F8 - 9CC5-6BAA1 27DB4CB</p><p>(Figures 10, 11, 25, 37, 48)</p><p>Material examined.   Holotype. ♂. PHILIPPINES: Luzon, Benguet,  Adunot-river Unterlauf; 16.41 ° N 120.74 ° E; 750 m; 19-21 Nov. 1997; K. Ebert, W. Mey, M. Nuss leg.; DNA voucher MFNLEP981; genitalia slide MfN gen. prep. 357M; specimen voucher coll.mfn-berlin.- de_u_47117f (MfN).  Paratypes. 5 ♀; specimen identifiers coll.mfn-berlin.de_u_2a6f6c, b09940, d0aa91, 2e049a, 780f46; 3 ♂; specimen identifiers coll.mfn-berlin.de_u_9179ce, 37d979, 651f8a. Detailed information in Table S1 .</p><p>Similar species.  Sufetula fulgurata sp. nov.,  Sufetula elfridea sp. nov.</p><p>Diagnosis. In the forewing (Figures 10, 11), the antemedian line is more conspicuously jagged than in  Sufetula fulgurata sp. nov. and  S. elfridea sp. nov. In the male genitalia, the valva apex is evenly rounded, while it is slightly pointed in  Sufetula fulgurata sp. nov. In the female genitalia (Figure 48), the colliculum opens posteriad directly outwards and is square in ventral aspect, while the sclerotised ring observed in  Sufetula fulgurata sp. nov. is narrower and does not reach the posterior margin.</p><p>Description. Wings (Figures 10, 11). Forewing length: 4.5 - 6.5 mm in both sexes. Male forewing (Figure 11) colour pale yellow speckled with brown. Antemedian line white, edged with dark brown on both sides, jagged in shape, forming two teeth pointing distad. Costa with pale orange streak. Two white patches in median area close to antemedial line: one between teeth of line, other occupying most of dorsal area. Costa with three white triangular spots obliquely pointed towards margin, basally edged with dark brown. Small, round discal spot. Postmedian line white, bordered with dark brown, enlarged at costa; first curved outwards, acutely pointed toward distal margin along medial fold, then mostly straight and oblique to dorsum at 2/3 from base of wing, with one jag directed basad; curved outwards at termen. Forewing female (Figure 11) pale brown, with antemedian, postmedian and subterminal area brown coloured. Marked orange suffusion at costa. Costal spots with a fourth spot near postmedian line. Postmedian line narrower at costa than in male specimens. Forewing CuP vein absent. Tympanal organs (Figure 25). ♀. Bullae tympani with posterior fourth recessed posterior of ramus tympani. Posterior depression shallow, as wide as bulla tympani, with posterior edge not clearly demarcated. Venulae secundae conspicuously marked, slightly incurved, running down to anterior edge of sternum II. Male genitalia (Figure 37). Uncus ca 1/3 of tegumen arm length, narrowed at basal 1/3, apex spatulate. Valva ovate, with apex evenly rounded. Female genitalia (Figure 48). Posterior apophyses 2.5 - 2.7 × tergite VIII length. Anterior apophyses 1.8 - 2 × tergite VIII length, at basal 1/4 with rounded ventral bump. Colliculum opening directly posteriad, tubular, short. Ductus bursae 2.2 - 2.7 × corpus length, slender, straight. Corpus bursae medium-sized, globular; posterior half with erected acanthae. Signum 1/3 of corpus length, forming oval patch of small, sclerotised, tooth-shaped thorns.</p><p>Distribution. Known from Luzon (Benguet) at an elevation of 750 m (Figure 55).</p><p>Etymology. From the Latin serratus, a, um, “serrate, saw-toothed”, referring to the serrate antemedian line on the forewing.</p><p>DNA barcoding. An intraspecific variation of 0.7% is observed between specimen MFNLEP962 from Zambales (Luzon) and specimen MFNLEP963 from Benguet (Luzon). The nearest nonconspecific neighbour is  Sufetula choreutalis (specimen MFNLEP970) from Leyte.</p></div>	https://treatment.plazi.org/id/03817A535859FFF7FF890469FBB1FE43	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535859FFF6FCC70474FDD2FDD3.text	03817A535859FFF6FCC70474FDD2FDD3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula monticola Muller, Hayden & Leger 2025	<div><p>Sufetula monticola Müller, Hayden &amp; Léger,  sp. nov.</p><p>LSID urn:lsid:zoobank.org:act: 685C97C9-AA41-4AEE-B5DE-BE59D A55AF1C</p><p>(Figures 12, 13, 26, 38, 49)</p><p>Material examined.   Holotype. ♂. PHILIPPINES: Luzon, Nueva Vizcaya,  Bald Mountains,  Santa Fe; 16.18 ° N 120.86 ° E; 1150m; 11-13 Nov.1997; K. Ebert, W. Mey, M. Nuss leg.; DNA voucher MFNLEP941; genitalia slide MfN gen. prep. 332M; specimen identifier coll.mfn-berlin.de_u_2a6f5f (MfN).  Paratypes. 4 ♂; specimen identifiers coll.mfn-berlin.de_u_2a6f5e, 96e51d, 884347, 2a6f5d; 7 ♀; specimen identifiers coll.mfn-berlin.de_u_f3238a, b03275, 18dc1a, 410f79, 2a6f61, 2a6f60, 2265d4. Detailed information on Table S1 .</p><p>Diagnosis.  Sufetula monticola sp. nov. is best separated from other  Sufetula species by examination of the forewings: the antemedian line forms one conspicuous, narrow jag projected toward the discoidal spot, followed by two well-marked curves, and the costal spots are semi-circular, pale yellow edged with dark brown (Figures 12, 13). In the male genitalia (Figure 38), the uncus is slightly shorter than in other  Sufetula species, being ca 1/4 of the length of the tegumen. In the female genitalia (Figure 49), the anterior apophyses are comparatively short, and the corpus bursae bears a large signum about half of the corpus length.</p><p>Description. Wings (Figures 12,13). Forewing length: 7 - 8.5 mm in both sexes; forewing ground colour greyish brown. Small basal dark brown dot. Antemedian line thin, white, edged with dark brown, forming one conspicuous tooth pointing distad toward discal spot, then arced toward dorsum with two shallow curves. Median discoidal spot oblong, black. Three costal pale yellow semicircles innerly light brown. Postmedian line forming triangular blotch at costa, slightly incurved on radial veins, roundly curved distad on medial fold, then bent inwards toward dorsum, slightly thickened around anal fold. Margin chequered with dark brown and pale yellow. Fringes chequered with brown and pale yellow. Hindwing cream coloured with faintly marked white median line shifted inwards at 2A vein. Dark brown dashed line bordering margin; margin forming thin, cream-coloured line. Forewing CuP vein absent. Tympanal organs (Figure 26). ♂ ♀. Bullae tympani posterior fourth recessed. Posterior depressions shallow, large, half width of bulla. Venulae secundae faintly marked, incurved medially, running down to anterior 1/3 of sternum II. Pons tympani fused anteriorly, forming rounded opening posteriad. Male genitalia (Figure 38). Uncus ca 1/4 of tegumen arm length, triangular, apex bluntly pointed. Valva suboval, ventral margin straight, dorsal margin conspicuously rounded at base, slightly convex otherwise; apex rounded. Female genitalia (Figure 49). Posterior apophyses 2.3 × tergite VIII length. Anterior apophyses 1.9 × tergite VIII length, thicker than posterior apophyses. Ductus bursae ca twice the length of corpus, straight, covered with minute spicules. Corpus bursae large, globular, covered with acanthae. Signum ca half the length of corpus, forming rounded patch of short, tooth-shaped, sclerotised thorns.</p><p>Distribution. Known from Luzon (Benguet, Mountain Province) at an elevation between 600 and 2350 m (Figure 55).</p><p>Etymology. From the Latin  monticola, ae, f, “mountain dwelling”, referring to the medium elevation distribution of the species.</p><p>DNA barcoding. The species delimitation analysis recovers three different MOTUs differing by 4.7 - 6.2%. Two MOTUs are recovered within the specimens from the Cordillera Central on Luzon, one including the specimens MFNLEP939, MFNLEP940 and MFNLEP-PYRALPHIL09-G11 collected at elevations between 1650 and 2350 m, while the 11 specimens of the other MOTUs were collected at elevations between 750 and 1150 m. The third MOTU represents the single specimen MFNLEP-PYRALPHIL09-H10 from Mindoro.</p><p>Remarks. The second MOTU represented by three specimens collected at higher elevation could represent a distinct species. In the male genitalia, the uncus is shorter than in other specimens. The restriction to higher elevation of this MOTU suggests possible allopatric divergence through elevational isolation.</p></div>	https://treatment.plazi.org/id/03817A535859FFF6FCC70474FDD2FDD3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535858FFF6FFA00405FA21FF60.text	03817A535858FFF6FFA00405FA21FF60.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula falcata Muller, Hayden & Leger 2025	<div><p>Sufetula falcata Müller, Hayden &amp; Léger,  sp. nov.</p><p>LSID urn:lsid:zoobank.org:act: 31A9F139 - 796F-4D06-B14 C-B 58F9 AB447B8</p><p>(Figures 14, 27, 39, 50)</p><p>Material examined.   Holotype. ♂. PHILIPPINES: Luzon, Zambales, Zambales Mountains,  Coto; 15.46 ° N 120.05 ° E; 150m; 09-10 Nov. 1998; W. Mey, W. Speidel leg.; DNA voucher MFNLEP977; MfN gen. prep. 400M; specimen voucher coll.mfn-berlin.de_u_5f078e (MfN).  Paratypes. 2 ♀; specimen identifiers coll.mfn-berlin.de_u_961e1d, 2af8cb; 1 unsexed; specimen identifier coll.mfn-berlin.- de_u_c7387e. Detailed information in Table S1 .</p><p>Diagnosis.  Sufetula falcata sp. nov. exhibits short and slender forewings (Figure 14) with a nearly straight postmedian line. In the male genitalia (Figure 39), the oblong shape of the valva separates this species from other  Sufetula species. In the female genitalia (Figure 50), the clearly demarcated C-shaped signum bearing sclerotised spines separates this species from its congeners.</p><p>Description. Wings (Figure 14). Forewing length: 4.3 mm (n -1); narrow, ground colour greyish brown to dirty white. Antemedian line white, thin, broadly curved outwards to CuP vein, running straight to dorsum along anal area. Costa with three rounded, whitish spots. Median discoidal spot small, rounded, black. Postmedian line white, running roughly straight toward dorsum up to vein M 2, dorsal half slightly incurved inwards. Margin dashed with dark brown. Wing outer margin projected outwards at apex. Fringe chequered dirty white and brown. Tympanal organs (Figure 27). ♂ ♀. Bullae tympani 1/6 recessed posterior of ramus tympani. Posterior depressions shallow, half as wide as bulla, length about 1/3 width, apparently not sexually dimorphic. Venulae secundae faintly marked, incurved medially, running down to anterior 1/3 of sternum II. Pons tympani fused throughout. Male genitalia (Figure 39). Uncus ca 2/5 of tegumen arm length, slender, with constricted base, apex blunt with bristles. Tegumen bridge broad, roughly as long as uncus. Valva with oblong, elongate shape, about twice as long as wide, distally slightly curved upwards, apex truncate. Phallus with two patches of minute sclerotised spines at anterior 1/4. Female genitalia (Figure 50). Posterior apophyses ca 2.7 × tergite VIII length. Anterior apophyses ca 2 × tergite VIII length. Colliculum short, forming sclerotised ring. Ductus bursae ca 2.3 × corpus length, straight, looped twice on anterior 1/4. Corpus bursae ovoid. Signum ca 1/4 × corpus length, C-shaped with concave side facing posteriad, bearing sclerotised spines.</p><p>Distribution. Known from Luzon (Zambales), Palawan and Panay an elevation between 150 and 900 m (Figure 55).</p><p>Etymology. From the Latin falcatus, a, um, “bent, scythe-shaped”, referring to the C-shaped signum in the corpus bursae of the female genitalia.</p><p>DNA barcoding. The maximum observed intraspecific variation is of 0.3% between the specimen MFNLEP-PYRALPHIL11-D07 from Palawan and the specimen MFNLEP977 from Zambales (Luzon).</p></div>	https://treatment.plazi.org/id/03817A535858FFF6FFA00405FA21FF60	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535858FFF6FCDD0754FABDF9A7.text	03817A535858FFF6FCDD0754FABDF9A7.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula minuta Muller, Hayden & Leger 2025	<div><p>Sufetula minuta Müller, Hayden &amp; Léger,  sp. nov.</p><p>LSID urn:lsid:zoobank.org:act: 66E6A5C3-6202-4762-AF7B-8724A 2E4872</p><p>(Figures 15, 28, 51)</p><p>Material examined.   Holotype. ♀. PHILIPPINES: Samar, Borongan,  San Mateo; 11.67 ° N 125.35 ° E; 40m; 26 Apr. 1997; W. Mey, W. Speidel leg.; DNA voucher MFNLEP967; genitalia slide MfN gen. prep. 396F; specimen voucher coll.mfn-berlin.de_u_c135d6 (MfN).  Paratype. 2 ♂; specimen identifiers coll.mfn-berlin.de_u_065c76, 065c40; 1 ♀; specimen identifier coll.mfn-berlin.de_u_065b67; 1 unsexed; specimen identifier coll.mfn-berlin.de_u_752be2. Detailed information in Table S1 .</p><p>Diagnosis.  Sufetula minuta sp. nov. (Figure 15) is very small (forewing length - 3.6 mm). The costal median spots are very small, flat semicircles accompanied with a dark brown streak. The female genitalia (Figure 51) are unmistakable because of the lack of a discrete signum, the looped and sclerotised ductus bursae, and the remarkably long papillae anales.</p><p>Description. Wings (Figure 15). Forewing length: 3.6 mm; narrow, ground colour cream, speckled with dark brown. Antemedian area dark brown. Antemedian line cream, jagged. Costa with three small cream median spots. Median area dark brown on basal half. Median discoidal spot oblong, dark brown. Postmedian line cream, basally edged with dark brown, slightly incurved from costa to medial veins, bent outwards toward marginal emargination, then markedly arched basad from M 3 to dorsum. Subterminal area dark brown. Margin cream. Fringes chequered cream and dark brown. Hindwing brown, medially with two broad cream bands, postmedian area with cream patches. Tympanal organs (Figure 28). ♀. Bullae tympani posterior fourth recessed. Venulae secundae faintly marked, running nearly straight down to tergite II anterior edge. Tympanic depression small, about 1/4 width of bulla, a welldemarcated saccus tympani. Male genitalia. Not known. Female genitalia (Figure 51). Posterior apophyses ca 3.1 × tergite VIII length. Anterior apophyses ca 2.5 × tergite VIII length. Colliculum tubular, weakly sclerotised, ca 1/5 × tergite VIII length. Ductus bursae ca 3.3 × corpus length, anterior 1/3 forming sclerotised loop. Corpus bursae globular, covered with acanthae, anterior half with V-shaped patch of densely packed acanthae, proper signum absent.</p><p>Distribution. Known from the Philippines (Palawan, Samar), at elevation between 40 and 390 m (Figure 55).</p><p>Etymology. From the Latin minutus, a, um, “short”, referring to the short size of the species.</p><p>DNA barcoding. A maximum intra-specific variation of 7.3% is observed between the specimen MFNLEP-PYRALPHIL11-E07 from Palawan and the specimen MFNLEP967 from Samar.</p></div>	https://treatment.plazi.org/id/03817A535858FFF6FCDD0754FABDF9A7	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A535858FFF1FCDD0029FF54F95A.text	03817A535858FFF1FCDD0029FF54F95A.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula metallias (Meyrick 1897) Müller & Hayden & Lees & Léger 2025	<div><p>Sufetula metallias (Meyrick, 1897),  comb. nov.</p><p>(Figures 16, 29, 40, 52)</p><p>Diplopseustis metallias Meyrick, 1897: 90-91 . Type locality: Indonesia, Sangir [Sangihe] Is[land].</p><p>Material examined.  Syntypes: ♂ (specimen ID NHMUK013699638):  “ LECTO-TYPE ” [dark blue circled] | “Type” [red circled] | “ Sangir. Doherty. Feb.–Mar. 1892 ” |“  Diplopseustis |  metallias Meyr. ”  “metallias Meyr ”; | 97-97; ♂ (specimen ID NHMUK013699636): “   PARA-LECTO-TYPE ” [light blue circled] | “ Sangir. Doherty. Feb.–Mar.1892 ” | “  Diplopseustis metallias Meyr. 1/ 1 Meyrick E. Meyrick det. In Meyrick coll.”. Deposited in NHMUK  .</p><p>Other specimens examined. 15 ♂, 13 ♀, 2 unsexed. Detailed information in Table S1.</p><p>Similar species.  Sufetula cyanolepis Hampson, 1912 .</p><p>Diagnosis. In the forewing (Figure 16), the shiny silver markings, the tawny yellow background colour and the two costal spots are shared with  Sufetula cyanolepis (type locality: South Sulawesi). The absence of a broad brownish suffusion in the median area of the fore- and hindwings separates this species from  S. cyanolepis . In  S. metallias comb. nov. the tawny yellow background predominates not only in the median area but in the pre-terminal area of both wing pairs, so that the species appears contrastingly marked with brown. In addition, in  S. cyanolepis there is a premedian and pre-terminal narrow whitish band in the forewing and a postmedian one in the hindwing that are not evident in  S. metallias comb. nov. In the male genitalia (Figure 40), the uncus is narrowed at basal 1/3, and the apex is spatulate. The female genitalia (Figure 52) are identical to those of  Sufetula elfridea sp. nov. and  S. serrata sp. nov.</p><p>Redescription. Wings (Figure 16). Forewing ground colour tawny to yellow. Antemedian area with thick dark brown patch at costa, on its middle with faintly marked shiny silver patch. Antemedian line thick, white speckled with silver iridescent scales, thinly edged with dark brown; arched distad across discal cell to anal fold, then running straight and perpendicular to dorsum from anal fold to edge. Costa with two oblique, tooth-shaped median spots, white with iridescent scales. Discal spot oblong, parallel to costa, white with iridescent scales, dorso-distally flanked with dark brown patch. Postmedian line thick, white, near costa with iridescent scales, thinly edged with dark brown; running perpendicular to costa to medial veins, bent obliquely basad on median fold, conspicuously indented inwards on CuA 2. Subterminal area speckled with dark brown, along margin with two patches of iridescent scales. Margin dark brown. Fringes chequered tawny yellow and brown. Hindwing pale yellow. Postmedian line white with iridescent scales, faded at costa, shifted inwards on CuP. Subterminal area dark brown on dorsal half. Tympanal organs (Figure 29). ♂ ♀. Bullae tympani 1/4 recessed posterior of ramus tympani. Posterior depressions large, almost as wide as bulla, length 1/3 width, not dimorphic. Venulae secundae faintly marked, kinked twice (♂) or running nearly straight (♀), running down to anterior 1/3 of sternum II. Pons tympani fused throughout. Male genitalia (Figure 40). Uncus ca 1/3 of tegumen arm length, narrowed at basal 1/4, apex spatulate. Valva subovate, apex rounded. Phallus slender, devoid of denticles apically, cornuti absent. Female genitalia (Figure 52). Posterior apophyses ca 2.2 - 2.5 × tergite VIII length. Anterior apophyses ca 1.7 - 2 × tergite VIII length. Colliculum short, tubular, sclerotised. Ductus bursae ca 2.1 × corpus length, slender, straight. Corpus bursae medium-sized, globular; posterior half with erected acanthae. Signum 1/4 of corpus length, forming an oval patch of small, sclerotised, tooth-shaped thorns.</p><p>Distribution. Known from Indonesia, Sulawesi (Sangihe island, South Sulawesi) and from the Philippines (Leyte), at an elevation between 120 and 650 m (Figure 55).</p><p>DNA barcoding. An intraspecific distance of 3.7% is observed between the specimens MFNLEP968 from Libas (Southern Leyte, 120 m elevation) and MFNLEP974 from Lake Danao (Leyte, 650 m elevation).</p></div>	https://treatment.plazi.org/id/03817A535858FFF1FCDD0029FF54F95A	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A53585FFFF1FF89009DFB3AF983.text	03817A53585FFFF1FF89009DFB3AF983.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula polystrialis (Hampson 1912)	<div><p>Sufetula polystrialis (Hampson, 1912)</p><p>(Figures 17, 30, 41)</p><p>Sufetula polystrialis Hampson, 1912 . Type locality: “Louisiades, St. Aignan Island [Misima Island, Papua New Guinea]]”.</p><p>Material examined.   Syntypes: 1 ♂ (specimen identifier NHMUK 013699689; left wings missing): “Type” [red circled] | “St. Aignan I. Louisiade. A.S. Meek, 99 - 32” | “  Sufetula polystrialis type ♂. Hmpsn”;   1 ♀ (specimen identifier NHMUK 013699634); “Type” [red circled]; “  St. Aignan Nov. 1897 (Meek)” [black squared] | “  Sufetula polystrialis type ♀. Hmpsn.” | “99 - 49”; Deposited in NHMUK  .</p><p>Other specimens examined.  ♀ (probably not syntypic; specimen identifier NHMUK015666552): Milne Bay A.S. Meek | 1900 - 45; 1 ♂ (specimen identifier coll.mfn-berlin.de_u_7b23a3). Detailed information in Table S1.</p><p>Diagnosis. This species can be best recognized by the white striated postmedian area and the two flattened lunule-shaped costal spots on the forewing (Figure 17), as well as the broad postmedian line running roughly straight, slightly disrupted near the dorsum on the hindwing. In the male genitalia (Figure 41), the evenly tongue-shaped uncus separates this species from other  Sufetula species. The female genitalia are not known.</p><p>Redescription. Wings (Figure 17). Forewing length: 7 - 9 mm; ground colour light brown. Antemedian line forming short tip projecting distad on radial vein, bent outwards in discoidal cell, then bent inwards in cubital area, running to basal 1/4 of dorsum. Median and postmedian area with large patch of dark brown striated with white stripes on veins. Costa with two large, flattened rounded cream spots innerly cream or light brown. Postmedian line cream, arched outwards to M 3, then abruptly bent inwards toward base, jagged, directed vertically towards dorsum in cubital area. Submarginal line dark brown. Fringe basally cream, distally greyish brown. Hindwing ground colour gray and white. Antemedian area with dark brown transverse band. Postmedian line white, basally edged with dark brown, rather wide, running straight, slightly disrupted inwards near dorsum. Submarginal area with broad dark brown band. Marginal area whitish gray. Subterminal line thin, dark brown, sharply contrasted from marginal white. Fringe light brown. Tympanal organs (Figure 30). ♂. Bullae tympani 1/4 recessed lateral of venula prima. Saccus tympani present, laterally demarcated, showing typical shape associated with sexually dimorphism, almost 1/2 width of bulla. Venulae secundae barely marked, fading posteriad from saccus tympani. Male genitalia (Figure 41). Uncus ca 2/5 of tegumen arm length, moderately large, with linguiform apex. Valva ca 1.6 × longer than wide, subrectangular, ventral margin concave on basal 1/3, then straight; angled at 2/3, then running upwards; apex forming slightly demarcated rounded protrusion. Female genitalia. Not known.</p><p>Distribution. PHILIPPINES: Luzon (Quezon) (Figure 55). INDONESIA: Irian Jaya (Birdshead Peninsula). PAPUA NEW GUINEA: Milne Bay (Misima Island); Hydrographer Mts 2500 ft.; Mt. Riu, Sudest I., 2000 ft.; Dampier Isl.; Ishurava, 3,000 ft; Upper Aroa R.; Sariba Isl.; New Ireland; MALUKU: Batian [Batjan]; SOLOMONS: Arawa, N. Bougainville; Rendova. Collected at an elevation of 120 m on Luzon; ca. 600 - 900 m elsewhere.</p><p>DNA barcoding. The only specimen subjected to molecular analyses did not yield any DNA barcode.</p></div>	https://treatment.plazi.org/id/03817A53585FFFF1FF89009DFB3AF983	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
03817A53585FFFF0FCC70035FDA3FB09.text	03817A53585FFFF0FCC70035FDA3FB09.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Sufetula oculalis Muller, Hayden & Leger 2025	<div><p>Sufetula oculalis Müller, Hayden &amp; Léger,  sp. nov.</p><p>LSID urn:lsid:zoobank.org:act: FDD6D9D5-3DE6-4051-AECD-ACDB F3CED9D8</p><p>(Figures 18, 31, 53)</p><p>Material examined.   Holotype. ♀. PHILIPPINES: Mindoro, Oriental Mindoro, Mount Halcon,  Mount Halcon; 13.32 ° N 120.90 ° E; 1300m; 15-17 Jan. 1998; W. Mey, W. Speidel leg.; DNA voucher MFNLEP-PYRALPHIL11-D08; genitalia slide TL1420F; specimen identifier coll.mfn-berlin.de_u_24e965 (MfN).  Paratype. 1 ♀; specimen identifier NHMUK013699660. Detailed information in Table S1 .</p><p>Diagnosis. The conspicuous ocellus-like median discoidal spot of the forewing (Figure 18) unambiguously separates this species from its congeners. In the female genitalia (Figure 53), the long tubular colliculum and the short ductus with sclerotised spicules in the distal half differentiate this species from other  Lathrotelinae .</p><p>Description. Wings (Figure 18). Forewing ground colour brown. Antemedian area with three white blotches. Antemedian line wide, white, waved on radial and cubital veins. Median area with three white spots: one in proximal half of discal cell, one proximally between Cu and anal fold, and one distally between Cu and fold. Distal discoidal patch forming white ocellus with dark brown center. Costa with three median white lunule-shaped spots. Postmedian line wide, J-shaped in costal half, abutted inwardly with white blotch; extended from M 2 and M 3 veins, roughly straight to dorsum, with tooth-shaped projection on or about anal fold directed toward distal white spot. Subterminal area dark brown, distally light brown. Hindwing dirty white; broad postmedian band thickly edged with dark brown, shifted inwards near dorsum. Subterminal line faded white, running parallel to median line. Tympanal organs (Figure 31). ♀. Bullae tympani 1/4 recessed posterior of ramus tympani. Saccus tympani small, 1/2 width of bulla, widest in mesal corner between bulla and pons tympani. Venulae secundae strongly bent inwards at base, then running straight towards sternum II posterior margin, nearly reaching it. Transverse ridge medially marked, straight. Male genitalia. Not known. Female genitalia (Figure 53). Posterior apophyses ca 2.3 × tergite VIII length. Anterior apophyses 2 × tergite VIII length. Antrum membranous, elongated. Colliculum four times longer than wide, tubular, sclerotised. Ductus bursae ca 2/3 × length of corpus bursae, rather straight, anterior half broader, covered with sclerotised spicules. Corpus bursae globular, posterior half covered with short, erect papillae. Signum 3/5 × corpus length, forming large patch of short, sclerotised, tooth-shaped thorns.</p><p>Distribution. Known from Mindoro at an elevation of 1300 m (Figure 55).</p><p>Etymology. From the Latin ocularis, e, ”related to the eye“, referring to the ocelli-like distal discoidal spot of the forewing.</p><p>DNA barcoding. A 313bp DNA barcode could be obtained for one single specimen (MFNLEP-PYRALPHIL11-D08).</p></div>	https://treatment.plazi.org/id/03817A53585FFFF0FCC70035FDA3FB09	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Müller, Anne;Hayden, James;Lees, David C.;Léger, Théo	Müller, Anne, Hayden, James, Lees, David C., Léger, Théo (2025): Assessment of species diversity of the Lathrotelinae (Lepidoptera: Crambidae) from the Philippines using morphology and DNA barcoding reveals eight new species. Journal of Asia-Pacific Biodiversity CLXVI (CLXVI): 1-24, DOI: 10.1016/j.japb.2025.01.012, URL: https://doi.org/10.1016/j.japb.2025.01.012
