identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
038987E8FFF16E13FF461092FC6EFD35.text	038987E8FFF16E13FF461092FC6EFD35.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus Genier & Saxton 2025	<div><p>Rhytomus Génier &amp; Saxton, new genus</p><p>urn:lsid:zoobank.org:act: 3800DB4F-9D65-4AD7-9A13-8809F0F0C3F0</p><p>Types species. Tesserodon elongatum van Lansberge, 1885 .</p><p>Diagnosis. The presence of a trochanterofemoral pit combined with a pseudepipleuron, the presence of a posterior hypomeral carina and well-defined elytral striae 9 and 10 will separate Rhytomus from all the other genera in New Guinea. Within the Australasian endemic clade, Rhytomus can be distinguished from other genera by those previous characters listed in conjunction with having 9 elytral striae, and wide eyes with an interocular space 3–6 times the eye width.</p><p>Description. Body. Total length 3.0– 6.3 mm. Tegument hard, brittle. Color dark reddish brown to black (teneral specimens lighter in color), ventral surface and legs usually lighter in color, lacking metallic sheen. Head. Anterior edge of clypeus with two clypeal teeth, with or without emargination on external side of each tooth. Dorsal part of eyes wide (Fig. 4A,B), inner edge straight basally, outer edge arcuate, with at least 12 facets along its widest portion. Ocular canthus incomplete. Occiput of head simply rounded (Fig. 4C,D). Antennae nine segmented, with 3-antennomere club, club antennomeres approximately equal width. Antennomeres 7 and 8 lacking median cavity.</p><p>Pronotum. Basal edge of pronotum broadly arcuate. Posterior hypomeral carina present, almost as long as half lateral pronotal edge. Anterior edge lacking membranous imbrication, anterior marginal bead very narrow. Elytra. Elytra with 9 striae, 7 on disc and 2 on the pseudepipleuron adjacent to the epipleural edge (Fig. 4D). Stria 8 missing. Stria 9 effaced anteriorly, fused or very closely set with tenth stria past basal third to posterior declivity where it is distinct again. With a well-defined pseudepipleural fold from base to apical declivity outside stria 7. Laterobasal elytral pit present or absent (Fig. 4A). Hind wings. Present, fully developed. Pygidium. Deeply and narrowly sulcate across the base, sulcus usually wider and deeper laterally (Fig. 8). Pterothoracic ventrite. Mesepimera with coarse semicircular ocellate punctures. Abdominal ventrite. Ventrite 6 approximately as long as 1–5 combined along midline. Legs. Protibiae (Fig. 4E) of male abruptly internally expanded apically, distal edge of internal angle with a row of contiguous spiniform setae flanked posteriorly by a row of long squamiform setae, with more or less elongated triangular spur. Protibiae distally straight with 3 teeth on outer edge, each tooth with two groups of setae dorsally. Prothoracic legs with trochanterofemoral anterior pit (Fig. 4C), trochanter excavated dorsally for reception of inner apical edge of protibia. Protarsi with 5 tarsomeres, tarsomere 1 and 5 longer. Long, setae on all tarsomeres or tarsomere 1 lacking setae. Male mesotibiae with dense setae on ventral edge. Metatibiae of males with well-developed tooth on inner apical angle. Mesotarsi with 5 tarsomeres, tarsomere 2 about same length as tarsomere 5. Metatarsomere 1 widest, metatarsomere 2 equal to metatarsomere 5 in length, all tarsomeres with long setae ventrally. Claws simple or slightly angulate at base. Metacoxal connecting membrane apodeme present, oval (internal). Genitalia. Aedeagus with parameres asymmetrical, right paramere apically bent downwards and expanded, left paramere more or less bent downward apically or simply rounded (Figs. 9, 10). Spermatheca Vshaped, with or without apodeme distally and basally.</p><p>Etymology. Based on the combination of two Greek words. The feminine noun ῥῠτῐ́ς (rhytís) meaning wrinkle, crease and the masculine noun ὦμος (omos) meaning shoulder, the terminal component transliterated in Latin with the masculine ending -us (ICZN art. 30.1.2). The name refers to the presence of a crease on the humeral surface of the elytron.</p><p>Distribution. Papua New Guinea and Indonesia (Seram Island, Waigeo Island and Papua). Specimens collected at elevations from 3 m to 2050 m (Fig. 5).</p><p>Remarks. Tesserodon seramicum Krikken &amp; Huijbregts, 2009 (described from Seram Island, Indonesia) and Tesserodon waigeoensis Ochi, Kon &amp; Hartini, 2017 (described from Waigeo Island, West Papua) are from outside the study area. However, examination of the illustrations provided in the original descriptions and images of a specimen (RMNH.INS.1488474) substantiate their transfer to the genus Rhytomus and should be combined as follows: Rhytomus seramicus (Krikken &amp; Huijbregts, 2009), new combination and Rhytomus waigeoensis (Ochi, Kon &amp; Hartini, 2017), new combination . Spermathecal morphology was studied only for species where enough material was available. Males of R. heurni Paulian, 1937 new combination and R. setulosus (Balthasar, 1965) new combination are unknown; the online identification key will be updated when additional material becomes available.</p><p>Species checklist</p><p>Rhytomus elongatus (van Lansberge, 1885), new combination (type species) Rhytomus fervidus Saxton &amp; Génier, new species</p><p>Rhytomus heurni (Paulian, 1937), new combination</p><p>Rhytomus howdeni (Paulian, 1985), new combination</p><p>Rhytomus larseni Saxton &amp; Génier, new species</p><p>Rhytomus latidentatus Saxton &amp; Génier, new species</p><p>Rhytomus papuanus (Paulian, 1985), new combination</p><p>Rhytomus seramicus (Krikken &amp; Huijbregts, 2009), new combination Rhytomus setulosus (Balthasar, 1965), new combination</p><p>Rhytomus sodalis Saxton &amp; Génier, new species</p><p>Rhytomus vanus Saxton &amp; Génier, new species</p><p>Rhytomus waigeoensis (Ochi, Kon &amp; Hartini, 2017), new combination</p></div>	https://treatment.plazi.org/id/038987E8FFF16E13FF461092FC6EFD35	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFF6E12FF4612CAFDC2FC31.text	038987E8FFFF6E12FF4612CAFDC2FC31.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus elongatus (van Lansberge 1885) Génier & Gunter & Saxton 2025	<div><p>Rhytomus elongatus (van Lansberge, 1885), new combination</p><p>Figs 6, 9A</p><p>Tesserodon elongatum van Lansberge, 1885: 376 (original description)— Paulian 1934: 287 (identification key, distribution); Paulian 1985: 221 (identification key, distribution)</p><p>Tesserodon elongatus — Gillet 1911: 40 (catalogue); Gunter et al. 2019: 453 (distribution)</p><p>Name-bearing type data. Lectotype ♂ (MSNG) by present designation: || Nuova Guinea | Fly River | L.M.D Albertis 1876-77 ||; || TYPUS || [bordered and printed in red]; || Tesserodon | elongatum | Lansberge Type || [handwritten]; || elongatum | Lansb. || [handwritten with a border]; || SYNTYPUS | Tesserodon | elongatum | Lansberge, 1885 || [partly handwritten]; || WORLD SCARAB. | DATABASE | WSD00028173 || [QR code label]; || LECTOTYPE ♂ | Tesserodon | elongatum | van Lansberge, 1885 | des. Génier &amp; Larsen, 2015 || [red card]. Studied.</p><p>Material examined (1♂, 1♀). PAPUA NEW GUINEA: WESTERN PROVINCE, Fly River, [08°00’ S, 142°00’E], 1876–1877, L.M. D’Albertis — ♂ lectotype, 1 ♀ paralectotype (MHNG) .</p><p>Redescription. Measurements. Body length 6.0 mm, maximum body width 2.9 mm. Body. Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third. Head. Clypeal edge lateral to clypeal teeth lacking emargination (Fig. 6B), head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.3 times eye width. Pronotum. Pronotal width to length along midline ratio 1.7, lateral pronotal edge sharply defined throughout, pronotal pubescence long and conspicuous, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures (Fig. 6A), basal elytral pit wide and deep, basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3–5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures. Legs. Profemoral anterior surface deeply furrowed, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (e.g. Fig. 12D). Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex. Aedeagus. Left paramere apex produced into a rounded oblique projection (Fig. 9A).</p><p>Remarks. This species was described based on two syntypes. In 2015, Génier and Larsen labeled one male from this series as the lectotype.</p></div>	https://treatment.plazi.org/id/038987E8FFFF6E12FF4612CAFDC2FC31	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFE6E12FF4613CBFBADF801.text	038987E8FFFE6E12FF4613CBFBADF801.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus fervidus Saxton & Genier 2025	<div><p>Rhytomus fervidus Saxton &amp; Génier, new species</p><p>urn:lsid:zoobank.org:act: 27C09598-572B-4AF0-904F-59067A688C08</p><p>Figs. 4B, 7A, 8A, 9B, 11A, 11D, 11E, 12B</p><p>Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA | Western highlands, Mt. | Hagen, 6000’ | 5-8.VII.1974, S.Peck | oak forest dung tps. 24-25 ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049283 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | fervidus | Des. N. Saxton &amp; F. Génier, 2024 || [red card] .</p><p>Material examined (20 ♂♂, 9 ♀♀). PAPUA NEW GUINEA: WESTERN HIGHLANDS PROVINCE, Mount Hagen, 1829 m, [05°52’ S, 144°14’E], 5–8.vii.1974, oak forest, dung traps, S. Peck, (dung traps 24-25)— ♂ holotype, ♀ allotype, 12 ♂♂, 6 ♀♀ paratypes (CMNC), 6 ♂♂, 1 ♀ paratypes (MNHN), 1 ♂, 1 ♀ paratypes (QM) .</p><p>Description. Measurements. Body length 4.2–5.0 mm, maximum body width 2.6–3.1 mm. Holotype: body length 4.7 mm, maximum body width 3.0 mm. Body. Overall shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination (Fig. 4C), head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.9 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures (Fig. 12B), basal elytral pit slightly wider than stria 10 and shallow (Fig. 11D), basal surface of pseudepipleuron with irregularly distributed more or less defined punctures (Fig. 11E), interstriae 3–5 on basal fifth with scattered unaligned punctures. Legs. Profemoral anterior surface flat, male metatibial apicointernal tooth approximately in line with posterior tibial edge. Pygidium. Pygidial surface along midline with punctures absent or distinctly more scattered than rest of discs, at least basally, pygidial surface along basal groove smooth between punctures (Fig. 8A), pygidium of female depressed apically. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 9B). Female genitalia. Spermatheca with two apodemes basally.</p><p>Etymology. The specific epithet for this species is the masculine nominative Latin adjective meaning “boiling hot” or “fiery” in reference to the type locality, which is near an old composite volcano.</p></div>	https://treatment.plazi.org/id/038987E8FFFE6E12FF4613CBFBADF801	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFD6E10FF46173AFC56FCED.text	038987E8FFFD6E10FF46173AFC56FCED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus heurni (Paulian 1937) Génier & Gunter & Saxton 2025	<div><p>Rhytomus heurni (Paulian, 1937) new combination</p><p>Fig. 7B, 8B, 11F</p><p>Tesserodon heurni Paulian, 1937: 122 (original description)— Paulian 1985: 221 (diagnosis, distribution, comment); Gunter et al. 2019: 453 (distribution)</p><p>Name-bearing type data. Holotype ♀ (SDEI): || N. Guinea Exped. | W.C. van Heurn | 1410 m. Okt. 1920 | Dormanpadbivak ||; || Holotypus || [red card]; || DEI Müncheberg | Col-05059 || [blue card]; || WORLD SCARAB. | DATABASE | WSD00028176 || [QR code label]; || Tesserodon | Heurni n. sp. Type | det. R. Paulian 1937 || [partly handwritten]. Studied.</p><p>Redescription. Measurements. Body length 4.3 mm, maximum body width 3.1 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 4.1 times eye width. Pronotum. Pronotal width to length along midline ratio 1.8, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with sharply defined ocellate punctures (Fig. 11F), interstriae 3–5 on basal fifth with scattered unaligned punctures. Legs. Profemoral anterior surface flat. Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex (Fig. 8B).</p><p>Remarks. The original description of this species is based on a single female specimen collected during W. C. van Heurn New Guinea expedition in October 1920. The type locality “Dormanpadbivak” roughly translates to “Doorman path bivouac.” According to Holthuis and Husson (1973) this locality was approximating 3º24’S 138º38’E at 1410-1450m, which is about 25km S. of Prauwenbivak.</p></div>	https://treatment.plazi.org/id/038987E8FFFD6E10FF46173AFC56FCED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFC6E10FF4616ECFEBAF898.text	038987E8FFFC6E10FF4616ECFEBAF898.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus howdeni (Paulian 1985) Génier & Gunter & Saxton 2025	<div><p>Rhytomus howdeni (Paulian, 1985), new combination</p><p>Fig. 7C, 4A, 8C, 9C</p></div>	https://treatment.plazi.org/id/038987E8FFFC6E10FF4616ECFEBAF898	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFC6E16FF461757FAABFB10.text	038987E8FFFC6E16FF461757FAABFB10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tesserodon howdeni Paulian 1985	<div><p>Tesserodon howdeni Paulian, 1985: 222 (original description)</p><p>Name-bearing type data. Holotype ♂ (CMNC): || Papua N.Guinea | Brown R. 30mi. N | Fort Moresby | 15- 16.VII.1974 ||; || S. Peck | collector ||; || H. &amp; A. Howden | collection ||; || HOLOTYPE || [red card]; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00011854 ||; || Tesserodon | howdeni | n. sp. | R. Paulian det. || [partly handwritten]. Note. Previously stated as: “Nouvelle-Guinée (Papouasie): Brown River, 20 miles N. Port Moresby, 14/ 16-VII-1974 (S. Peck), piège à excréments, en forêt” by Paulian. Studied .</p><p>Material examined (5 ♂♂, 2 ♀♀). PAPUA NEW GUINEA: CENTRAL PROVINCE, 18 mi. N port Moresby, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.2875&amp;materialsCitation.latitude=-9.204166" title="Search Plazi for locations around (long 147.2875/lat -9.204166)">Brown River</a>, 3 m, [09°12’15’’S, 147°17’15’’E], 14–18.vii.1974, forest, dung traps, S. Peck, (dung traps 39-42)— 1 ♂, 1 ♀ (CMNC) ; 20 mi. N Port Moresby, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.31667&amp;materialsCitation.latitude=-9.191667" title="Search Plazi for locations around (long 147.31667/lat -9.191667)">Brown River</a>, 3 m, [09°11’30’’S, 147°19’00’’E], 16–18.vii.1974, dung traps, S. Peck, (dung traps 39-41)— 1 ♀ (CMNC) ; same locality, 16–18.vii.1974, dung trap, S. Peck, (dung traps 39-41)— 2 ♂♂ paratypes (MNHN); 30 mi. N Port Moresby, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.34166&amp;materialsCitation.latitude=-9.183333" title="Search Plazi for locations around (long 147.34166/lat -9.183333)">Brown River</a>, [09°11’S, 147°20’30’’E], 15–16.vii.1974, S. Peck — ♂ holotype , 1 ♂ paratype (CMNC) .</p><p>Redescription. Measurements. Body length 3.6–4.1 mm, maximum body width 2.3–2.7 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth emarginate (Fig. 4D), head margin at clypeogenal suture of females moderately protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.6 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3–5 on basal fifth with scattered unaligned punctures. Legs. Profemoral anterior surface flat or might appear furrowed in some individuals, male metatibial apicointernal tooth approximately in line with posterior tibial edge. Pygidium. Pygidial groove transverse, narrowed medially, and terminating in a pit at each end (Fig. 8C), pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 9C).</p><p>Remarks. Examination of the allotype revealed that the specimen labeled by Paulian is actually a male and is here considered a male paratype in the material examined. Additionally, the date ranges provided by Paulian (1985) in the original description do not match any label data from the specimens studied. Recent examination of the specimens at the MNHN (by FG) confirmed that both are males, indicating that the female specimen described by Paulian is either missing or that Paulian misidentified the gender of the allotype. Two female specimens in the CMNC are not labeled as paratypes. While one of these could theoretically be the allotype, it is unclear if Henry Howden ever sent them to Paulian for study, as he often retained some specimens as a precaution against parcels being lost in the mail. Consequently, these specimens cannot be definitively considered part of the type series.</p></div>	https://treatment.plazi.org/id/038987E8FFFC6E16FF461757FAABFB10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFA6E15FF4614EBFDA4FE61.text	038987E8FFFA6E15FF4614EBFDA4FE61.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus larseni Saxton & Genier 2025	<div><p>Rhytomus larseni Saxton &amp; Génier, new species</p><p>urn:lsid:zoobank.org:act: 277D719A-381F-4447-8665-B42E8617FA3A</p><p>Fig. 7D, 8D, 9D, 11B, 11C, 12D</p><p>Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA: Western | Prov., P’nvang, CI RAP Survey | Camp 1. 05º29.5S 141º32.6E | 573 m. May 2013. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049307 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | larseni | Des. N. Saxton &amp; F. Génier, 2024 || [red card ].</p><p>Material examined (163 ♂♂, 90 ♀♀). PAPUA NEW GUINEA: WESTERN PROVINCE, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.53334&amp;materialsCitation.latitude=-5.483333" title="Search Plazi for locations around (long 141.53334/lat -5.483333)">CI RAP Survey Camp 1</a>, P’nvang, 575 m, 05°29’S, 141°32’E, v.2013, T. Larsen — 2 ♂♂, 2 ♀♀ paratypes (ANIC), 2 ♂♂, 2 ♀♀ paratypes (NHMUK), ♂ holotype, ♀ allotype, 101 ♂♂, 54 ♀♀ paratypes (CMNC), 2 ♂♂, 2 ♀♀ paratypes (MNHN), 2 ♂♂, 2 ♀♀ paratypes (NMPC), 2 ♂♂, 2 ♀♀ paratypes (USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.53334&amp;materialsCitation.latitude=-5.483333" title="Search Plazi for locations around (long 141.53334/lat -5.483333)">CI RAP Survey Camp 2</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.53334&amp;materialsCitation.latitude=-5.483333" title="Search Plazi for locations around (long 141.53334/lat -5.483333)">P’nvang</a>, 800 m, 05°30’S, 141°34’E, v.2013, T. Larsen — 49 ♂♂, 23 ♀♀ paratypes (CMNC), 2 ♂♂, 2 ♀♀ paratypes (QM) .</p><p>Description. Measurements. Body length 4.1–5.4 mm, maximum body width 2.7–3.1 mm. Holotype: body length 4.7 mm, maximum body width 3.0 mm. Body. Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.5 times eye width. Pronotum. Pronotal width to length along midline ratio 1.8, lateral pronotal edge sharply defined throughout, pronotal pubescence long and conspicuous, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures with distinct microsculptures (Fig. 11B), basal elytral pit wide and deep (Fig. 11C), basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3–5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures in some individuals. Legs. Profemoral anterior surface deeply furrowed, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (Fig. 12D). Pygidium. Pygidial groove terminating in a pit at each end, pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures (Fig. 8D), pygidium of female evenly convex. Aedeagus. Left paramere apex produced into a rounded oblique projection (Fig. 9D). Female genitalia. Spermatheca lacking apodemes.</p><p>Etymology. This specific epithet of this species is the Latinized genitive form of “Larsen”, in honor of Trond H. Larsen who collected all the specimens in the type series and contributed greatly to the number of Rhytomus specimens available for study from PNG.</p></div>	https://treatment.plazi.org/id/038987E8FFFA6E15FF4614EBFDA4FE61	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF86E0BFF4616DCFD52FBA0.text	038987E8FFF86E0BFF4616DCFD52FBA0.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus latidentatus Saxton & Genier 2025	<div><p>Rhytomus latidentatus Saxton &amp; Génier, new species</p><p>urn:lsid:zoobank.org:act: 8C418C61-9081-48AD-B9BB-6CBC74D23206</p><p>Figs. 4C, 4E, 7E, 8E, 10A, 12A</p><p>Name-bearing type data. Holotype ♂ (CMNC). || PAPUA NEW GUINEA: Western | Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.54333&amp;materialsCitation.latitude=-5.491667" title="Search Plazi for locations around (long 141.54333/lat -5.491667)">P’nvang</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.54333&amp;materialsCitation.latitude=-5.491667" title="Search Plazi for locations around (long 141.54333/lat -5.491667)">CI RAP Survey</a> | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.54333&amp;materialsCitation.latitude=-5.491667" title="Search Plazi for locations around (long 141.54333/lat -5.491667)">Camp 1.</a> 05º29.5S 141º32.6E | 573 m. May 2013. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049540 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | latidentatus | Des. N. Saxton &amp; F. Génier, 2024 || [red card] .</p><p>Material examined (22 ♂♂, 12 ♀♀). PAPUA NEW GUINEA: CENTRAL PROVINCE, Kokoda Trak, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.625&amp;materialsCitation.latitude=-9.191667" title="Search Plazi for locations around (long 147.625/lat -9.191667)">Kauai River</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.625&amp;materialsCitation.latitude=-9.191667" title="Search Plazi for locations around (long 147.625/lat -9.191667)">Menari</a>, 700 m, [09°11’30’’S, 147°37’30’’E], 12–14.viii.1976, rainforest, dung, O. Kukal — 3 ♂♂, 1 ♀ paratypes (CMNC), 2 ♂♂ paratypes (MNHN); WESTERN PROVINCE, CI RAP Survey <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.625&amp;materialsCitation.latitude=-9.191667" title="Search Plazi for locations around (long 147.625/lat -9.191667)">Camp 1</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=147.625&amp;materialsCitation.latitude=-9.191667" title="Search Plazi for locations around (long 147.625/lat -9.191667)">P’nvang</a>, 575 m, 05°29’S, 141°32’E, v.2013, T. Larsen — 1 ♂, 1 ♀ paratypes (ANIC), ♂ holotype, ♀ allotype, 14 ♂♂, 8 ♀♀ paratypes (CMNC), 1 ♂, 1 ♀ paratypes (QM) .</p><p>Description. Measurements. Body length 4.5–6.3 mm, maximum body width 3.1–4.2 mm. Holotype: length 5.5 mm, maximum body width 3.9 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, clypeal teeth wide, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.1 times eye width. Pronotum. Pronotal width to length along midline ratio 1.6, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline with fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures with distinct microsculptures (Fig. 12A), basal elytral pit absent, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3–5 on basal fifth with two rows of punctures along striae with the odd unaligned punctures on interstriae 3 and 5 and scattered unaligned punctures on interstria 4. Legs. Profemoral anterior surface flat, male metatibial apicointernal teeth approximately in line with posterior tibial edge. Pygidium. Pygidial groove transverse, narrowed medially, and terminating in a pit at each end (Fig. 8E), surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex. Aedeagus. Left paramere apex rounded in a dorsoventral plane (Fig. 10A). Female genitalia. Spermatheca with two apodemes basally and one apodeme apically.</p><p>Etymology. The specific epithet of this species is a combination of the Latin adjective “ latus ” meaning wide and the adjective “ dentatus ” meaning dentate in reference to this species’ characteristically wide clypeal teeth.</p><p>Remarks. Paulian (1985) studied four specimens of this species which he identified as “ Tesserodon heurni ” in his work on the Canthonines of New Guinea.</p></div>	https://treatment.plazi.org/id/038987E8FFF86E0BFF4616DCFD52FBA0	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE76E0BFF461467FE8FFB10.text	038987E8FFE76E0BFF461467FE8FFB10.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus papuanus (Paulian 1985) Génier & Gunter & Saxton 2025	<div><p>Rhytomus papuanus (Paulian, 1985), new combination</p><p>Fig. 7F, 8F, 10B</p></div>	https://treatment.plazi.org/id/038987E8FFE76E0BFF461467FE8FFB10	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE76E0AFF4614CFFE52FE1D.text	038987E8FFE76E0AFF4614CFFE52FE1D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Tesserodon papuanum Paulian 1985	<div><p>Tesserodon papuanum Paulian, 1985: 221 (original description)</p><p>Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA | Wau, 1400 m. | 29.VI-2.VII.1974. S. Peck | carrion traps 18-19 ||; || PAPUA NEW GUINEA | Wau, 1400 m. | 29.vi-2.vii.74 | S. Peck, carT18-19 || [photocopy of handwritten label]; || H. &amp; A.Howden | Collection ||; || HOLOTYPE || [red card]; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00011856 || [QR code label]; || Tesserodon | papuanum n.sp. | R. Paulian det. || [partly handwritten]. Studied .</p><p>Material examined (21 ♂♂, 7 ♀♀). PAPUA NEW GUINEA: MOROBE PROVINCE, Aseki Road, 56 km W Bululo, 1750 m, 07°19’33’’S, 146°20’49’’E, 8.ii.2000, mixed montane forest litter, R.S. Anderson, (RSA2000- 039a)— 2 ♂♂ (CMNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.71666&amp;materialsCitation.latitude=-7.3333335" title="Search Plazi for locations around (long 146.71666/lat -7.3333335)">Gumi L.A. Bulolo</a>, 2134 m, [07°11’S, 146°25’E], 18.xi.1984, infly trap, H. Roberts, (C.I.E. A16609)— 1 ♂ (NHMUK); Mount Kaindi, 9 km NNW Wau, 2050 m, 07°20’24’’S, 146°40’01’’E, 6.ii.2000, montane forest litter, R.S. Anderson, (RSA2000-035b)— 1 ♂ (CMNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.71666&amp;materialsCitation.latitude=-7.3333335" title="Search Plazi for locations around (long 146.71666/lat -7.3333335)">Wau</a>, 1219 m, [07°20’S, 146°43’E], 1–3.vii.1974, forest litter, berlese, S. Peck — 1 ♀ (CMNC); same locality, 1–3.vii.1974, H. Howden — 1 ♂, 1 ♀ (MNHN); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=146.71666&amp;materialsCitation.latitude=-7.3333335" title="Search Plazi for locations around (long 146.71666/lat -7.3333335)">Wau</a>, 1400 m, [07°20’S, 146°43’E], 29.vi.–2.vii.1974, carrion traps, S. Peck, (carrion traps 18-19)— ♂ holotype, 12 ♂♂, 3 ♀♀ paratypes (CMNC), 2 ♂♂, 1 ♀ paratypes (MNHN); same locality, 29.vi.–2.vii.1974, carrion traps, S. Peck, (carrion trap 20)— ♀ allotype (CMNC), 1 ♂ paratype (MNHN) .</p><p>Redescription. Measurements. Body length 4.1–5.3 mm, maximum body width 2.7–3.1 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.9 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge rounded, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3–5 on basal fifth with scattered unaligned punctures. Legs. Profemoral anterior surface flat, male metatibial apicointernal tooth approximately in line with posterior tibial edge. Pygidium. Pygidial surface along midline with punctures absent or distinctly more scattered than rest of discs, at least basally, pygidial surface along basal groove smooth between punctures (Fig. 8F), pygidium of female evenly convex. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 10B). Female genitalia. Spermatheca with two apodemes basally and one apodeme apically.</p></div>	https://treatment.plazi.org/id/038987E8FFE76E0AFF4614CFFE52FE1D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E0AFF4611FBFCE4FAC3.text	038987E8FFE66E0AFF4611FBFCE4FAC3.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus setulosus (Balthasar 1965) Génier & Gunter & Saxton 2025	<div><p>Rhytomus setulosus (Balthasar, 1965), new combination</p><p>Fig. 7G, 8G</p><p>Tesserodon setulosum Balthasar, 1965: 17 (original description)— Paulian 1985: 221 (diagnosis, distribution, comment)— Bezděk &amp; Hájek 2011: 374 (type data, comment); Gunter et al. 2019: 453 (distribution)</p><p>Name-bearing type data. Holotype ♀ (NMPC): || N. Guinea | Biró 1899 ||; || Sattelberg | Huon-Golf ||; || 1899 | IV.1-15 || [handwritten on verso of previous label]; || Tesserodon | setulosum | n.sp. Balth. | 63 Holotypus || [partly handwritten]. Studied.</p><p>Material examined (2 ♀♀). PAPUA NEW GUINEA: MOROBE PROVINCE, Sattelberg, [06°29’S, 147°46’E], 1–15.iv.1899, Biró —♀ holotype, 1 ♀ paratype (NMPC) (female paratype examined via image courtesy of Jiří Hájek).</p><p>Redescription. Measurements. Body length 3.2 mm, maximum body width 2.3 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females moderately protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.3 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline strongly convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit wide and deep, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3–5 on basal fifth with scattered unaligned punctures on interstriae 3 and 5 and two rows of punctures along striae with the odd unaligned punctures on interstria 4. Legs. Profemoral anterior surface flat. Pygidium. Pygidial surface along midline with punctures absent or distinctly more scattered than rest of discs, at least basally, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex (Fig. 8G).</p><p>Remarks. No male specimens known for this species.</p></div>	https://treatment.plazi.org/id/038987E8FFE66E0AFF4611FBFCE4FAC3	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E08FF4615C0FDEAFD7D.text	038987E8FFE66E08FF4615C0FDEAFD7D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus sodalis Saxton & Genier 2025	<div><p>Rhytomus sodalis Saxton &amp; Génier, new species</p><p>urn:lsid:zoobank.org:act: 7168813D-24AF-4CCA-8B46-B5168014A9E3</p><p>Figs. 7H, 8H, 10C, 12C</p><p>Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA: Western | Prov., <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.54333&amp;materialsCitation.latitude=-5.491667" title="Search Plazi for locations around (long 141.54333/lat -5.491667)">P’nvang</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.54333&amp;materialsCitation.latitude=-5.491667" title="Search Plazi for locations around (long 141.54333/lat -5.491667)">CI RAP Survey</a> | <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.54333&amp;materialsCitation.latitude=-5.491667" title="Search Plazi for locations around (long 141.54333/lat -5.491667)">Camp 1.</a> 05º29.5S 141º32.6E | 573 m. May 2013. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049583 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | sodalis | Des. N. Saxton &amp; F. Génier, 2024 || [red card] .</p><p>Material examined (69 ♂♂, 24 ♀♀). PAPUA NEW GUINEA: WESTERN PROVINCE, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.53334&amp;materialsCitation.latitude=-5.483333" title="Search Plazi for locations around (long 141.53334/lat -5.483333)">CI RAP Survey Camp</a> 1, P’nvang, 575 m, 05°29’S, 141°32’E, v.2013, T. Larsen — 2 ♂♂, 1 ♀ paratypes (ANIC), 2 ♂♂, 1 ♀ paratypes (NHMUK), ♂ holotype, ♀ allotype, 23 ♂♂, 12 ♀♀ paratypes (CMNC), 2 ♂♂, 1 ♀ paratypes (MNHN), 2 ♂♂, 1 ♀ paratypes (NMPC), 1 ♂, 1 ♀ paratypes (USNM); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.55&amp;materialsCitation.latitude=-5.5333333" title="Search Plazi for locations around (long 141.55/lat -5.5333333)">CI RAP Survey Camp 2</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.55&amp;materialsCitation.latitude=-5.5333333" title="Search Plazi for locations around (long 141.55/lat -5.5333333)">P’nvang</a>, 800 m, 05°30’S, 141°34’E, v.2013, T. Larsen — 15 ♂♂, 3 ♀♀ paratypes (CMNC); <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.55&amp;materialsCitation.latitude=-5.5333333" title="Search Plazi for locations around (long 141.55/lat -5.5333333)">CI RAP Survey Camp 3</a>, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=141.55&amp;materialsCitation.latitude=-5.5333333" title="Search Plazi for locations around (long 141.55/lat -5.5333333)">P’nvang</a>, 1110 m, 05°32’S, 141°33’E, v.2013, T. Larsen — 19 ♂♂, 1 ♀ paratypes (CMNC), 2 ♂♂, 2 ♀♀ paratypes (QM) .</p><p>Description. Measurements. Body length 3.0– 4.6 mm, maximum body width 2.1–2.7 mm. Holotype: body length 3.8 mm, maximum body width 2.5 mm. Body. Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.2 times eye width. Pronotum. Pronotal width to length along midline ratio 1.7, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit wide and deep, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3–5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures in some individuals. Legs. Profemoral anterior surface flat or might appear furrowed in some individuals, male metatibial apicointernal tooth approximately in line with posterior tibial edge (Fig. 12C). Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove with coarse transverse microsculptures (Fig. 8H), pygidium of female depressed basally. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 10C). Female genitalia. Spermatheca with two apodemes basally and one apodeme apically.</p><p>Etymology. The specific epithet of this species is a masculine noun in apposition “sŏdālis ” (companion, mate, fellow) relating to the fact that it was collected along with R. larseni and R. latidentatus .</p><p>Remarks. One female specimen (excluded from type series) was discovered from the same collecting event (i.e., Camp 1, 575 m) that had coarser pronotal punctation and a barely indicated meso-metaventrite suture. This specimen likely represents a fourth species. However, because no male specimens could be found at the time of this study, we refrain from describing it.</p></div>	https://treatment.plazi.org/id/038987E8FFE66E08FF4615C0FDEAFD7D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE46E08FF46131BFD16F899.text	038987E8FFE46E08FF46131BFD16F899.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus vanus Saxton & Genier 2025	<div><p>Rhytomus vanus Saxton &amp; Génier, new species</p><p>urn:lsid:zoobank.org:act: CD0A0711-1AC6-4D0B-8CE9-C54C31A001EA</p><p>Figs. 4D, 7I, 8I, 10D</p><p>Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA: Madang | Baitabag (8 km N.W.) | 100m, 30.I.2000 | 5°08’19”S 145°46’34”E | R.S.Anderson, rainforest litter | RSA2000-027 ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049581 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | vanus | Des. N. Saxton &amp; F. Génier, 2024 || [red card] .</p><p>Material examined (4♂♂, 1 ♀). PAPUA NEW GUINEA: MADANG PROVINCE, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=145.77611&amp;materialsCitation.latitude=-5.138611" title="Search Plazi for locations around (long 145.77611/lat -5.138611)">Baitabag</a>, 8 km NW Madang, 100 m, 05°08’19’’S, 145°46’34’’E, 30.i.2000, rainforest litter, R.S. Anderson, (RSA2000-027) — ♂ holotype, ♀ allotype, 3 ♂♂ paratypes (CMNC) .</p><p>Description. Measurements. Body length 3.3–3.6 mm, maximum body width 2.1–2.3 mm. Holotype: body length 3.6 mm, maximum body width 2.3 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females moderately protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 4.8 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures with distinct microsculptures, basal elytral pit wide and deep (Fig. 4A), basal surface of pseudepipleuron with irregularly distributed more or less defined punctures or with sharply defined simple punctures (by misinterpretation), interstriae 3–5 on basal fifth with scattered unaligned punctures on interstriae 3 and 5 and two rows of punctures along striae with the odd unaligned punctures on interstria 4. Legs. Profemoral anterior surface flat or deeply furrowed (by misinterpretation), male metatibial apicointernal tooth approximately in line with posterior tibial edge.</p><p>Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures (Fig. 8I), pygidium of female depressed apically. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 10D).</p><p>Etymology. Vānus is a Latin adjective. It is used here in the sense of “suspicious, doubtful” which pertain to the previously unrecognized status of this new species.</p></div>	https://treatment.plazi.org/id/038987E8FFE46E08FF46131BFD16F899	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE36E0FFF461092FA5FFAAF.text	038987E8FFE36E0FFF461092FA5FFAAF.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Rhytomus	<div><p>Key to adult mainland New Guinea Rhytomus species</p><p>Access to online key: https://keys.lucidcentral.org/keys/v4/rhytomus/ (click on the magic wand icon to select the best character to use first)</p><p>1 Interstriae 3–5 on basal fifth all having unaligned scattered punctures (Fig. 11A); basal pseudepipleural pit present, shallow and slightly larger than stria 10 (Fig. 11D).................................................................... 2</p><p>- Interstriae 3–5 with at least one interval having two distinct rows of punctures on basal fifth (11B); basal pseudepipleural pit deep and distinctly larger than stria 10 (Fig. 11C) OR pseudepipleural pit is absent................................. 5</p><p>2 Elytral interval 1 with basal punctures much smaller in size to those in intervals 2–3; pronotal lateral edge obsolete or sharply defined............................................................................................. 3</p><p>- Elytral interval 1 with basal punctures similar in size and coarseness to those in intervals 2–3; pronotal lateral edge always sharply defined...................................................................................... 4</p><p>3 Pronotal lateral edge obsolete; male clypeal edge without emargination lateral to clypeal teeth (Fig. 4B); Body length 4–5.3 mm .................................................................... Rhytomus papuanus (Paulian, 1985)</p><p>- Pronotal lateral edge sharply defined; male clypeal edge with emargination lateral to clypeal teeth (Fig. 4A); Body length 3.6–4.1 mm .............................................................. Rhytomus howdeni (Paulian, 1985)</p><p>4 Basal pseudepipleural surface with irregularly distributed more or less defined punctures (Fig. 11E); female pygidium depressed apically...................................................... Rhytomus fervidus Saxton &amp; Génier, new species</p><p>- Basal pseudepipleural surface with sharply defined ocellate punctures (Fig. 11F); female pygidium evenly convex.................................................................................. Rhytomus heurni (Paulian, 1937)</p><p>5 Base of pronotum lacking fine longitudinal depression medially; basal pseudepipleural pit deep and distinctly larger than stria 10 (Fig. 11C); male pygidial groove not distinctly narrowed medially (Figs. 8D, H, I)............................... 6</p><p>- Base of pronotum with fine longitudinal depression medially (Fig. E); basal pseudepipleural pit absent; male pygidial groove distinctly narrowed medially and terminating in pits (Fig. 8E)........ Rhytomus latidentatus Saxton &amp; Génier, new species</p><p>6 Interstriae 3, 4 and 5 having two distinct rows of punctures; elytral pubescence long and conspicuous (Fig. 11B)......... 7</p><p>- Interstriae 3 and 5 having unaligned and scattered punctures, interstriae 4 having two distinct rows of punctures; elytral pubescence short and fine (as in Fig. 11A)................................................................. 9</p><p>7 Male metatibial apicointernal tooth approximately in line with posterior tibial edge (Fig. 12C); pygidium of female depressed basally; body length 3.0– 4.6 mm .................................. Rhytomus sodalis Saxton &amp; Génier, new species</p><p>- Male metatibial apicointernal tooth set anteriorly to posterior tibial edge (Fig. 12D); pygidium of female evenly convex: body length 4.1–6.0 mm.................................................................................... 8</p><p>8 Elytral surface between punctures dull, with distinct microsculptures (Fig. 7D, 11B); Body length 4.1–5.4 mm ......................................................................... Rhytomus larseni Saxton &amp; Génier, new species</p><p>- Elytral surface between punctures appearing glossy, at most with ill-defined microsculptures (Fig. 6A); Body length 6mm ................................................................... Rhytomus elongatus (van Lansberge, 1885)</p><p>9 Elytral surface appearing glossy, at most with ill-defined microsculpture (as in Fig. 12B); Elytra strongly convex in lateral view (Fig. 12E).............................................................. Rhytomus setulosus (Balthasar, 1965)</p><p>- Elytral surface dull, with distinct microsculpture (Fig. 12A); Elytra moderately convex in lateral view (Fig. 12F)........................................................................ Rhytomus vanus Saxton &amp; Génier, new species</p></div>	https://treatment.plazi.org/id/038987E8FFE36E0FFF461092FA5FFAAF	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Génier, François;Gunter, Nicole L.;Saxton, Natalie A.	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
