taxonID	type	description	language	source
038987E8FFF16E13FF461092FC6EFD35.taxon	description	urn: lsid: zoobank. org: act: 3800 DB 4 F- 9 D 65 - 4 AD 7 - 9 A 13 - 8809 F 0 F 0 C 3 F 0 Types species. Tesserodon elongatum van Lansberge, 1885.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF16E13FF461092FC6EFD35.taxon	diagnosis	Diagnosis. The presence of a trochanterofemoral pit combined with a pseudepipleuron, the presence of a posterior hypomeral carina and well-defined elytral striae 9 and 10 will separate Rhytomus from all the other genera in New Guinea. Within the Australasian endemic clade, Rhytomus can be distinguished from other genera by those previous characters listed in conjunction with having 9 elytral striae, and wide eyes with an interocular space 3 – 6 times the eye width.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF16E13FF461092FC6EFD35.taxon	description	Description. Body. Total length 3.0 – 6.3 mm. Tegument hard, brittle. Color dark reddish brown to black (teneral specimens lighter in color), ventral surface and legs usually lighter in color, lacking metallic sheen. Head. Anterior edge of clypeus with two clypeal teeth, with or without emargination on external side of each tooth. Dorsal part of eyes wide (Fig. 4 A, B), inner edge straight basally, outer edge arcuate, with at least 12 facets along its widest portion. Ocular canthus incomplete. Occiput of head simply rounded (Fig. 4 C, D). Antennae nine segmented, with 3 - antennomere club, club antennomeres approximately equal width. Antennomeres 7 and 8 lacking median cavity. Pronotum. Basal edge of pronotum broadly arcuate. Posterior hypomeral carina present, almost as long as half lateral pronotal edge. Anterior edge lacking membranous imbrication, anterior marginal bead very narrow. Elytra. Elytra with 9 striae, 7 on disc and 2 on the pseudepipleuron adjacent to the epipleural edge (Fig. 4 D). Stria 8 missing. Stria 9 effaced anteriorly, fused or very closely set with tenth stria past basal third to posterior declivity where it is distinct again. With a well-defined pseudepipleural fold from base to apical declivity outside stria 7. Laterobasal elytral pit present or absent (Fig. 4 A). Hind wings. Present, fully developed. Pygidium. Deeply and narrowly sulcate across the base, sulcus usually wider and deeper laterally (Fig. 8). Pterothoracic ventrite. Mesepimera with coarse semicircular ocellate punctures. Abdominal ventrite. Ventrite 6 approximately as long as 1 – 5 combined along midline. Legs. Protibiae (Fig. 4 E) of male abruptly internally expanded apically, distal edge of internal angle with a row of contiguous spiniform setae flanked posteriorly by a row of long squamiform setae, with more or less elongated triangular spur. Protibiae distally straight with 3 teeth on outer edge, each tooth with two groups of setae dorsally. Prothoracic legs with trochanterofemoral anterior pit (Fig. 4 C), trochanter excavated dorsally for reception of inner apical edge of protibia. Protarsi with 5 tarsomeres, tarsomere 1 and 5 longer. Long, setae on all tarsomeres or tarsomere 1 lacking setae. Male mesotibiae with dense setae on ventral edge. Metatibiae of males with well-developed tooth on inner apical angle. Mesotarsi with 5 tarsomeres, tarsomere 2 about same length as tarsomere 5. Metatarsomere 1 widest, metatarsomere 2 equal to metatarsomere 5 in length, all tarsomeres with long setae ventrally. Claws simple or slightly angulate at base. Metacoxal connecting membrane apodeme present, oval (internal). Genitalia. Aedeagus with parameres asymmetrical, right paramere apically bent downwards and expanded, left paramere more or less bent downward apically or simply rounded (Figs. 9, 10). Spermatheca Vshaped, with or without apodeme distally and basally.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF16E13FF461092FC6EFD35.taxon	etymology	Etymology. Based on the combination of two Greek words. The feminine noun ῥῠτῐ ́ ς (rhytís) meaning wrinkle, crease and the masculine noun ὦμος (omos) meaning shoulder, the terminal component transliterated in Latin with the masculine ending - us (ICZN art. 30.1.2). The name refers to the presence of a crease on the humeral surface of the elytron.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF16E13FF461092FC6EFD35.taxon	distribution	Distribution. Papua New Guinea and Indonesia (Seram Island, Waigeo Island and Papua). Specimens collected at elevations from 3 m to 2050 m (Fig. 5).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF16E13FF461092FC6EFD35.taxon	discussion	Remarks. Tesserodon seramicum Krikken & Huijbregts, 2009 (described from Seram Island, Indonesia) and Tesserodon waigeoensis Ochi, Kon & Hartini, 2017 (described from Waigeo Island, West Papua) are from outside the study area. However, examination of the illustrations provided in the original descriptions and images of a specimen (RMNH. INS. 1488474) substantiate their transfer to the genus Rhytomus and should be combined as follows: Rhytomus seramicus (Krikken & Huijbregts, 2009), new combination and Rhytomus waigeoensis (Ochi, Kon & Hartini, 2017), new combination. Spermathecal morphology was studied only for species where enough material was available. Males of R. heurni Paulian, 1937 new combination and R. setulosus (Balthasar, 1965) new combination are unknown; the online identification key will be updated when additional material becomes available.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFF6E12FF4612CAFDC2FC31.taxon	description	Figs 6, 9 A	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFF6E12FF4612CAFDC2FC31.taxon	materials_examined	Name-bearing type data. Lectotype ♂ (MSNG) by present designation: || Nuova Guinea | Fly River | L. M. D Albertis 1876 - 77 ||; || TYPUS || [bordered and printed in red]; || Tesserodon | elongatum | Lansberge Type || [handwritten]; || elongatum | Lansb. || [handwritten with a border]; || SYNTYPUS | Tesserodon | elongatum | Lansberge, 1885 || [partly handwritten]; || WORLD SCARAB. | DATABASE | WSD 00028173 || [QR code label]; || LECTOTYPE ♂ | Tesserodon | elongatum | van Lansberge, 1885 | des. Génier & Larsen, 2015 || [red card]. Studied. Material examined (1 ♂, 1 ♀). PAPUA NEW GUINEA: WESTERN PROVINCE, Fly River, [08 ° 00 ’ S, 142 ° 00 ’ E], 1876 – 1877, L. M. D’Albertis — ♂ lectotype, 1 ♀ paralectotype (MHNG).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFF6E12FF4612CAFDC2FC31.taxon	description	Redescription. Measurements. Body length 6.0 mm, maximum body width 2.9 mm. Body. Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third. Head. Clypeal edge lateral to clypeal teeth lacking emargination (Fig. 6 B), head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.3 times eye width. Pronotum. Pronotal width to length along midline ratio 1.7, lateral pronotal edge sharply defined throughout, pronotal pubescence long and conspicuous, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures (Fig. 6 A), basal elytral pit wide and deep, basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3 – 5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures. Legs. Profemoral anterior surface deeply furrowed, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (e. g. Fig. 12 D). Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex. Aedeagus. Left paramere apex produced into a rounded oblique projection (Fig. 9 A).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFF6E12FF4612CAFDC2FC31.taxon	discussion	Remarks. This species was described based on two syntypes. In 2015, Génier and Larsen labeled one male from this series as the lectotype.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFE6E12FF4613CBFBADF801.taxon	description	urn: lsid: zoobank. org: act: 27 C 09598 - 572 B- 4 AF 0 - 904 F- 59067 A 688 C 08 Figs. 4 B, 7 A, 8 A, 9 B, 11 A, 11 D, 11 E, 12 B	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFE6E12FF4613CBFBADF801.taxon	materials_examined	Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA | Western highlands, Mt. | Hagen, 6000 ’ | 5 - 8. VII. 1974, S. Peck | oak forest dung tps. 24 - 25 ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049283 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | fervidus | Des. N. Saxton & F. Génier, 2024 || [red card]. Material examined (20 ♂♂, 9 ♀♀). PAPUA NEW GUINEA: WESTERN HIGHLANDS PROVINCE, Mount Hagen, 1829 m, [05 ° 52 ’ S, 144 ° 14 ’ E], 5 – 8. vii. 1974, oak forest, dung traps, S. Peck, (dung traps 24 - 25) — ♂ holotype, ♀ allotype, 12 ♂♂, 6 ♀♀ paratypes (CMNC), 6 ♂♂, 1 ♀ paratypes (MNHN), 1 ♂, 1 ♀ paratypes (QM).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFE6E12FF4613CBFBADF801.taxon	description	Description. Measurements. Body length 4.2 – 5.0 mm, maximum body width 2.6 – 3.1 mm. Holotype: body length 4.7 mm, maximum body width 3.0 mm. Body. Overall shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination (Fig. 4 C), head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.9 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures (Fig. 12 B), basal elytral pit slightly wider than stria 10 and shallow (Fig. 11 D), basal surface of pseudepipleuron with irregularly distributed more or less defined punctures (Fig. 11 E), interstriae 3 – 5 on basal fifth with scattered unaligned punctures. Legs. Profemoral anterior surface flat, male metatibial apicointernal tooth approximately in line with posterior tibial edge. Pygidium. Pygidial surface along midline with punctures absent or distinctly more scattered than rest of discs, at least basally, pygidial surface along basal groove smooth between punctures (Fig. 8 A), pygidium of female depressed apically. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 9 B). Female genitalia. Spermatheca with two apodemes basally.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFE6E12FF4613CBFBADF801.taxon	etymology	Etymology. The specific epithet for this species is the masculine nominative Latin adjective meaning “ boiling hot ” or “ fiery ” in reference to the type locality, which is near an old composite volcano.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFD6E10FF46173AFC56FCED.taxon	description	Fig. 7 B, 8 B, 11 F	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFD6E10FF46173AFC56FCED.taxon	materials_examined	Name-bearing type data. Holotype ♀ (SDEI): || N. Guinea Exped. | W. C. van Heurn | 1410 m. Okt. 1920 | Dormanpadbivak ||; || Holotypus || [red card]; || DEI Müncheberg | Col- 05059 || [blue card]; || WORLD SCARAB. | DATABASE | WSD 00028176 || [QR code label]; || Tesserodon | Heurni n. sp. Type | det. R. Paulian 1937 || [partly handwritten]. Studied.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFD6E10FF46173AFC56FCED.taxon	description	Redescription. Measurements. Body length 4.3 mm, maximum body width 3.1 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 4.1 times eye width. Pronotum. Pronotal width to length along midline ratio 1.8, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with sharply defined ocellate punctures (Fig. 11 F), interstriae 3 – 5 on basal fifth with scattered unaligned punctures. Legs. Profemoral anterior surface flat. Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex (Fig. 8 B).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFD6E10FF46173AFC56FCED.taxon	discussion	Remarks. The original description of this species is based on a single female specimen collected during W. C. van Heurn New Guinea expedition in October 1920. The type locality “ Dormanpadbivak ” roughly translates to “ Doorman path bivouac. ” According to Holthuis and Husson (1973) this locality was approximating 3 º 24 ’ S 138 º 38 ’ E at 1410 - 1450 m, which is about 25 km S. of Prauwenbivak.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFC6E10FF4616ECFEBAF898.taxon	description	Fig. 7 C, 4 A, 8 C, 9 C	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFC6E16FF461757FAABFB10.taxon	materials_examined	Name-bearing type data. Holotype ♂ (CMNC): || Papua N. Guinea | Brown R. 30 mi. N | Fort Moresby | 15 - 16. VII. 1974 ||; || S. Peck | collector ||; || H. & A. Howden | collection ||; || HOLOTYPE || [red card]; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00011854 ||; || Tesserodon | howdeni | n. sp. | R. Paulian det. || [partly handwritten]. Note. Previously stated as: “ Nouvelle-Guinée (Papouasie): Brown River, 20 miles N. Port Moresby, 14 / 16 - VII- 1974 (S. Peck), piège à excréments, en forêt ” by Paulian. Studied. Material examined (5 ♂♂, 2 ♀♀). PAPUA NEW GUINEA: CENTRAL PROVINCE, 18 mi. N port Moresby, Brown River, 3 m, [09 ° 12 ’ 15 ’’ S, 147 ° 17 ’ 15 ’’ E], 14 – 18. vii. 1974, forest, dung traps, S. Peck, (dung traps 39 - 42) — 1 ♂, 1 ♀ (CMNC); 20 mi. N Port Moresby, Brown River, 3 m, [09 ° 11 ’ 30 ’’ S, 147 ° 19 ’ 00 ’’ E], 16 – 18. vii. 1974, dung traps, S. Peck, (dung traps 39 - 41) — 1 ♀ (CMNC); same locality, 16 – 18. vii. 1974, dung trap, S. Peck, (dung traps 39 - 41) — 2 ♂♂ paratypes (MNHN); 30 mi. N Port Moresby, Brown River, [09 ° 11 ’ S, 147 ° 20 ’ 30 ’’ E], 15 – 16. vii. 1974, S. Peck — ♂ holotype, 1 ♂ paratype (CMNC).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFC6E16FF461757FAABFB10.taxon	description	Redescription. Measurements. Body length 3.6 – 4.1 mm, maximum body width 2.3 – 2.7 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth emarginate (Fig. 4 D), head margin at clypeogenal suture of females moderately protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.6 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.1, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3 – 5 on basal fifth with scattered unaligned punctures. Legs. Profemoral anterior surface flat or might appear furrowed in some individuals, male metatibial apicointernal tooth approximately in line with posterior tibial edge. Pygidium. Pygidial groove transverse, narrowed medially, and terminating in a pit at each end (Fig. 8 C), pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 9 C).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFC6E16FF461757FAABFB10.taxon	discussion	Remarks. Examination of the allotype revealed that the specimen labeled by Paulian is actually a male and is here considered a male paratype in the material examined. Additionally, the date ranges provided by Paulian (1985) in the original description do not match any label data from the specimens studied. Recent examination of the specimens at the MNHN (by FG) confirmed that both are males, indicating that the female specimen described by Paulian is either missing or that Paulian misidentified the gender of the allotype. Two female specimens in the CMNC are not labeled as paratypes. While one of these could theoretically be the allotype, it is unclear if Henry Howden ever sent them to Paulian for study, as he often retained some specimens as a precaution against parcels being lost in the mail. Consequently, these specimens cannot be definitively considered part of the type series.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFA6E15FF4614EBFDA4FE61.taxon	description	urn: lsid: zoobank. org: act: 277 D 719 A- 381 F- 4447 - 8665 - B 42 E 8617 FA 3 A Fig. 7 D, 8 D, 9 D, 11 B, 11 C, 12 D	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFA6E15FF4614EBFDA4FE61.taxon	materials_examined	Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA: Western | Prov., P’nvang, CI RAP Survey | Camp 1. 05 º 29.5 S 141 º 32.6 E | 573 m. May 2013. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049307 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | larseni | Des. N. Saxton & F. Génier, 2024 || [red card]. Material examined (163 ♂♂, 90 ♀♀). PAPUA NEW GUINEA: WESTERN PROVINCE, CI RAP Survey Camp 1, P’nvang, 575 m, 05 ° 29 ’ S, 141 ° 32 ’ E, v. 2013, T. Larsen — 2 ♂♂, 2 ♀♀ paratypes (ANIC), 2 ♂♂, 2 ♀♀ paratypes (NHMUK), ♂ holotype, ♀ allotype, 101 ♂♂, 54 ♀♀ paratypes (CMNC), 2 ♂♂, 2 ♀♀ paratypes (MNHN), 2 ♂♂, 2 ♀♀ paratypes (NMPC), 2 ♂♂, 2 ♀♀ paratypes (USNM); CI RAP Survey Camp 2, P’nvang, 800 m, 05 ° 30 ’ S, 141 ° 34 ’ E, v. 2013, T. Larsen — 49 ♂♂, 23 ♀♀ paratypes (CMNC), 2 ♂♂, 2 ♀♀ paratypes (QM).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFA6E15FF4614EBFDA4FE61.taxon	description	Description. Measurements. Body length 4.1 – 5.4 mm, maximum body width 2.7 – 3.1 mm. Holotype: body length 4.7 mm, maximum body width 3.0 mm. Body. Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.5 times eye width. Pronotum. Pronotal width to length along midline ratio 1.8, lateral pronotal edge sharply defined throughout, pronotal pubescence long and conspicuous, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures with distinct microsculptures (Fig. 11 B), basal elytral pit wide and deep (Fig. 11 C), basal surface of pseudepipleuron with sharply defined simple punctures, interstriae 3 – 5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures in some individuals. Legs. Profemoral anterior surface deeply furrowed, male metatibial apicointernal tooth set anteriorly to posterior tibial edge (Fig. 12 D). Pygidium. Pygidial groove terminating in a pit at each end, pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures (Fig. 8 D), pygidium of female evenly convex. Aedeagus. Left paramere apex produced into a rounded oblique projection (Fig. 9 D). Female genitalia. Spermatheca lacking apodemes.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFFA6E15FF4614EBFDA4FE61.taxon	etymology	Etymology. This specific epithet of this species is the Latinized genitive form of “ Larsen ”, in honor of Trond H. Larsen who collected all the specimens in the type series and contributed greatly to the number of Rhytomus specimens available for study from PNG.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF86E0BFF4616DCFD52FBA0.taxon	description	urn: lsid: zoobank. org: act: 8 C 418 C 61 - 9081 - 48 AD-B 9 BB- 6 CBC 74 D 23206 Figs. 4 C, 4 E, 7 E, 8 E, 10 A, 12 A	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF86E0BFF4616DCFD52FBA0.taxon	materials_examined	Name-bearing type data. Holotype ♂ (CMNC). || PAPUA NEW GUINEA: Western | Prov., P’nvang, CI RAP Survey | Camp 1. 05 º 29.5 S 141 º 32.6 E | 573 m. May 2013. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049540 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | latidentatus | Des. N. Saxton & F. Génier, 2024 || [red card]. Material examined (22 ♂♂, 12 ♀♀). PAPUA NEW GUINEA: CENTRAL PROVINCE, Kokoda Trak, Kauai River, Menari, 700 m, [09 ° 11 ’ 30 ’’ S, 147 ° 37 ’ 30 ’’ E], 12 – 14. viii. 1976, rainforest, dung, O. Kukal — 3 ♂♂, 1 ♀ paratypes (CMNC), 2 ♂♂ paratypes (MNHN); WESTERN PROVINCE, CI RAP Survey Camp 1, P’nvang, 575 m, 05 ° 29 ’ S, 141 ° 32 ’ E, v. 2013, T. Larsen — 1 ♂, 1 ♀ paratypes (ANIC), ♂ holotype, ♀ allotype, 14 ♂♂, 8 ♀♀ paratypes (CMNC), 1 ♂, 1 ♀ paratypes (QM).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF86E0BFF4616DCFD52FBA0.taxon	description	Description. Measurements. Body length 4.5 – 6.3 mm, maximum body width 3.1 – 4.2 mm. Holotype: length 5.5 mm, maximum body width 3.9 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, clypeal teeth wide, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.1 times eye width. Pronotum. Pronotal width to length along midline ratio 1.6, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline with fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures with distinct microsculptures (Fig. 12 A), basal elytral pit absent, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3 – 5 on basal fifth with two rows of punctures along striae with the odd unaligned punctures on interstriae 3 and 5 and scattered unaligned punctures on interstria 4. Legs. Profemoral anterior surface flat, male metatibial apicointernal teeth approximately in line with posterior tibial edge. Pygidium. Pygidial groove transverse, narrowed medially, and terminating in a pit at each end (Fig. 8 E), surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex. Aedeagus. Left paramere apex rounded in a dorsoventral plane (Fig. 10 A). Female genitalia. Spermatheca with two apodemes basally and one apodeme apically.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF86E0BFF4616DCFD52FBA0.taxon	etymology	Etymology. The specific epithet of this species is a combination of the Latin adjective “ latus ” meaning wide and the adjective “ dentatus ” meaning dentate in reference to this species’ characteristically wide clypeal teeth.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFF86E0BFF4616DCFD52FBA0.taxon	discussion	Remarks. Paulian (1985) studied four specimens of this species which he identified as “ Tesserodon heurni ” in his work on the Canthonines of New Guinea.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE76E0BFF461467FE8FFB10.taxon	description	Fig. 7 F, 8 F, 10 B	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE76E0AFF4614CFFE52FE1D.taxon	materials_examined	Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA | Wau, 1400 m. | 29. VI- 2. VII. 1974. S. Peck | carrion traps 18 - 19 ||; || PAPUA NEW GUINEA | Wau, 1400 m. | 29. vi- 2. vii. 74 | S. Peck, carT 18 - 19 || [photocopy of handwritten label]; || H. & A. Howden | Collection ||; || HOLOTYPE || [red card]; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00011856 || [QR code label]; || Tesserodon | papuanum n. sp. | R. Paulian det. || [partly handwritten]. Studied. Material examined (21 ♂♂, 7 ♀♀). PAPUA NEW GUINEA: MOROBE PROVINCE, Aseki Road, 56 km W Bululo, 1750 m, 07 ° 19 ’ 33 ’’ S, 146 ° 20 ’ 49 ’’ E, 8. ii. 2000, mixed montane forest litter, R. S. Anderson, (RSA 2000 - 039 a) — 2 ♂♂ (CMNC); Gumi L. A. Bulolo, 2134 m, [07 ° 11 ’ S, 146 ° 25 ’ E], 18. xi. 1984, infly trap, H. Roberts, (C. I. E. A 16609) — 1 ♂ (NHMUK); Mount Kaindi, 9 km NNW Wau, 2050 m, 07 ° 20 ’ 24 ’’ S, 146 ° 40 ’ 01 ’’ E, 6. ii. 2000, montane forest litter, R. S. Anderson, (RSA 2000 - 035 b) — 1 ♂ (CMNC); Wau, 1219 m, [07 ° 20 ’ S, 146 ° 43 ’ E], 1 – 3. vii. 1974, forest litter, berlese, S. Peck — 1 ♀ (CMNC); same locality, 1 – 3. vii. 1974, H. Howden — 1 ♂, 1 ♀ (MNHN); Wau, 1400 m, [07 ° 20 ’ S, 146 ° 43 ’ E], 29. vi. – 2. vii. 1974, carrion traps, S. Peck, (carrion traps 18 - 19) — ♂ holotype, 12 ♂♂, 3 ♀♀ paratypes (CMNC), 2 ♂♂, 1 ♀ paratypes (MNHN); same locality, 29. vi. – 2. vii. 1974, carrion traps, S. Peck, (carrion trap 20) — ♀ allotype (CMNC), 1 ♂ paratype (MNHN).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE76E0AFF4614CFFE52FE1D.taxon	description	Redescription. Measurements. Body length 4.1 – 5.3 mm, maximum body width 2.7 – 3.1 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.9 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge rounded, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit slightly wider than stria 10 and shallow, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3 – 5 on basal fifth with scattered unaligned punctures. Legs. Profemoral anterior surface flat, male metatibial apicointernal tooth approximately in line with posterior tibial edge. Pygidium. Pygidial surface along midline with punctures absent or distinctly more scattered than rest of discs, at least basally, pygidial surface along basal groove smooth between punctures (Fig. 8 F), pygidium of female evenly convex. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 10 B). Female genitalia. Spermatheca with two apodemes basally and one apodeme apically.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E0AFF4611FBFCE4FAC3.taxon	description	Fig. 7 G, 8 G	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E0AFF4611FBFCE4FAC3.taxon	materials_examined	Name-bearing type data. Holotype ♀ (NMPC): || N. Guinea | Biró 1899 ||; || Sattelberg | Huon-Golf ||; || 1899 | IV. 1 - 15 || [handwritten on verso of previous label]; || Tesserodon | setulosum | n. sp. Balth. | 63 Holotypus || [partly handwritten]. Studied. Material examined (2 ♀♀). PAPUA NEW GUINEA: MOROBE PROVINCE, Sattelberg, [06 ° 29 ’ S, 147 ° 46 ’ E], 1 – 15. iv. 1899, Biró — ♀ holotype, 1 ♀ paratype (NMPC) (female paratype examined via image courtesy of Jiří Hájek).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E0AFF4611FBFCE4FAC3.taxon	description	Redescription. Measurements. Body length 3.2 mm, maximum body width 2.3 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females moderately protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 5.3 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline strongly convex in lateral view, elytral pubescence short and fine, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit wide and deep, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3 – 5 on basal fifth with scattered unaligned punctures on interstriae 3 and 5 and two rows of punctures along striae with the odd unaligned punctures on interstria 4. Legs. Profemoral anterior surface flat. Pygidium. Pygidial surface along midline with punctures absent or distinctly more scattered than rest of discs, at least basally, pygidial surface along basal groove smooth between punctures, pygidium of female evenly convex (Fig. 8 G).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E0AFF4611FBFCE4FAC3.taxon	discussion	Remarks. No male specimens known for this species.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E08FF4615C0FDEAFD7D.taxon	description	urn: lsid: zoobank. org: act: 7168813 D- 24 AF- 4 CCA- 8 B 46 - B 5168014 A 9 E 3 Figs. 7 H, 8 H, 10 C, 12 C	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E08FF4615C0FDEAFD7D.taxon	materials_examined	Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA: Western | Prov., P’nvang, CI RAP Survey | Camp 1. 05 º 29.5 S 141 º 32.6 E | 573 m. May 2013. T. Larsen ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049583 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | sodalis | Des. N. Saxton & F. Génier, 2024 || [red card]. Material examined (69 ♂♂, 24 ♀♀). PAPUA NEW GUINEA: WESTERN PROVINCE, CI RAP Survey Camp 1, P’nvang, 575 m, 05 ° 29 ’ S, 141 ° 32 ’ E, v. 2013, T. Larsen — 2 ♂♂, 1 ♀ paratypes (ANIC), 2 ♂♂, 1 ♀ paratypes (NHMUK), ♂ holotype, ♀ allotype, 23 ♂♂, 12 ♀♀ paratypes (CMNC), 2 ♂♂, 1 ♀ paratypes (MNHN), 2 ♂♂, 1 ♀ paratypes (NMPC), 1 ♂, 1 ♀ paratypes (USNM); CI RAP Survey Camp 2, P’nvang, 800 m, 05 ° 30 ’ S, 141 ° 34 ’ E, v. 2013, T. Larsen — 15 ♂♂, 3 ♀♀ paratypes (CMNC); CI RAP Survey Camp 3, P’nvang, 1110 m, 05 ° 32 ’ S, 141 ° 33 ’ E, v. 2013, T. Larsen — 19 ♂♂, 1 ♀ paratypes (CMNC), 2 ♂♂, 2 ♀♀ paratypes (QM).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E08FF4615C0FDEAFD7D.taxon	description	Description. Measurements. Body length 3.0 – 4.6 mm, maximum body width 2.1 – 2.7 mm. Holotype: body length 3.8 mm, maximum body width 2.5 mm. Body. Overall body shape in dorsal view elongate-oval, elytral edge slightly subangular at apical third. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females slightly protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 3.2 times eye width. Pronotum. Pronotal width to length along midline ratio 1.7, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence long and conspicuous, elytral surface between punctures appearing glossy, at most with ill-defined microsculptures, basal elytral pit wide and deep, basal surface of pseudepipleuron with irregularly distributed more or less defined punctures, interstriae 3 – 5 on basal fifth with two rows of punctures along striae, with at most a few scattered punctures in some individuals. Legs. Profemoral anterior surface flat or might appear furrowed in some individuals, male metatibial apicointernal tooth approximately in line with posterior tibial edge (Fig. 12 C). Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove with coarse transverse microsculptures (Fig. 8 H), pygidium of female depressed basally. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 10 C). Female genitalia. Spermatheca with two apodemes basally and one apodeme apically.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E08FF4615C0FDEAFD7D.taxon	etymology	Etymology. The specific epithet of this species is a masculine noun in apposition “ sŏdālis ” (companion, mate, fellow) relating to the fact that it was collected along with R. larseni and R. latidentatus.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE66E08FF4615C0FDEAFD7D.taxon	discussion	Remarks. One female specimen (excluded from type series) was discovered from the same collecting event (i. e., Camp 1, 575 m) that had coarser pronotal punctation and a barely indicated meso-metaventrite suture. This specimen likely represents a fourth species. However, because no male specimens could be found at the time of this study, we refrain from describing it.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE46E08FF46131BFD16F899.taxon	description	urn: lsid: zoobank. org: act: CD 0 A 0711 - 1 AC 6 - 4 D 0 B- 8 CE 9 - C 54 C 31 A 001 EA Figs. 4 D, 7 I, 8 I, 10 D	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE46E08FF46131BFD16F899.taxon	materials_examined	Name-bearing type data. Holotype ♂ (CMNC): || PAPUA NEW GUINEA: Madang | Baitabag (8 km N. W.) | 100 m, 30. I. 2000 | 5 ° 08 ’ 19 ” S 145 ° 46 ’ 34 ” E | R. S. Anderson, rainforest litter | RSA 2000 - 027 ||; || Canadian Museum of | Musée canadien de la | NATURE | CMNEN 00049581 || [QR code label]; || HOLOTYPE ♂ | Rhytomus | vanus | Des. N. Saxton & F. Génier, 2024 || [red card].	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE46E08FF46131BFD16F899.taxon	materials_examined	Material examined (4 ♂♂, 1 ♀). PAPUA NEW GUINEA: MADANG PROVINCE, Baitabag, 8 km NW Madang, 100 m, 05 ° 08 ’ 19 ’’ S, 145 ° 46 ’ 34 ’’ E, 30. i. 2000, rainforest litter, R. S. Anderson, (RSA 2000 - 027) — ♂ holotype, ♀ allotype, 3 ♂♂ paratypes (CMNC).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE46E08FF46131BFD16F899.taxon	description	Description. Measurements. Body length 3.3 – 3.6 mm, maximum body width 2.1 – 2.3 mm. Holotype: body length 3.6 mm, maximum body width 2.3 mm. Body. Overall body shape in dorsal view oval, elytral edge evenly rounded from base to apex. Head. Clypeal edge lateral to clypeal teeth lacking emargination, head margin at clypeogenal suture of females moderately protruding. Dorsal parts of eyes semi-ovoid, separated by an interocular space of about 4.8 times eye width. Pronotum. Pronotal width to length along midline ratio 1.9, lateral pronotal edge sharply defined throughout, pronotal pubescence short and fine, pronotal basal portion along midline lacking fine longitudinal depression. Elytra. Elytral maximum width to length along suture ratio 1.0, elytral outline moderately convex in lateral view, elytral pubescence short and fine, elytral surface between punctures with distinct microsculptures, basal elytral pit wide and deep (Fig. 4 A), basal surface of pseudepipleuron with irregularly distributed more or less defined punctures or with sharply defined simple punctures (by misinterpretation), interstriae 3 – 5 on basal fifth with scattered unaligned punctures on interstriae 3 and 5 and two rows of punctures along striae with the odd unaligned punctures on interstria 4. Legs. Profemoral anterior surface flat or deeply furrowed (by misinterpretation), male metatibial apicointernal tooth approximately in line with posterior tibial edge. Pygidium. Pygidial surface along midline with punctures similar in density to rest of discs, pygidial surface along basal groove smooth between punctures (Fig. 8 I), pygidium of female depressed apically. Aedeagus. Left paramere apex truncated apically, distinctly angular externally (Fig. 10 D).	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
038987E8FFE46E08FF46131BFD16F899.taxon	etymology	Etymology. Vānus is a Latin adjective. It is used here in the sense of “ suspicious, doubtful ” which pertain to the previously unrecognized status of this new species.	en	Génier, François, Gunter, Nicole L., Saxton, Natalie A. (2025): Rhytomus Génier & Saxton: A new dung beetle genus from New Guinea with five new species and phylogenetic insights (Coleoptera: Scarabaeidae: Scarabaeinae). Zootaxa 5575 (1): 87-110, DOI: 10.11646/zootaxa.5575.1.3, URL: https://doi.org/10.11646/zootaxa.5575.1.3
