identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
0389433CFFFCAE58FF27E33D7F068FEB.text	0389433CFFFCAE58FF27E33D7F068FEB.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microhyla nanapollexa Bain & Nguyen 2004	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Microhyla nanapollexa Bain &amp; Nguyen, 2004</p>
            <p>(Figs. 2, 3; Tables 2)</p>
            <p>
                 Specimens examined. (n = 2),  one adult male IEBR A.6269 (Field number 04287) and  one adult female IEBR A.6270 (Field number KP2023.52), collected by C. V.  
                <a title="Search Plazi for locations around (long 108.26642/lat 14.664972)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=108.26642&amp;materialsCitation.latitude=14.664972">Hoang</a>
                 et al. in April 2023 in the forest of Kon Plong District, Kon Tum Province, Vietnam (14°39'53.9"N 108°15'59.1"E, at an elevation of 1260 m a.s.l.)  . 
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            <p>Description of specimens examined. Morphological characters of the specimens from Kon Tum Province, Vietnam, match the description of Bain &amp; Nguyen (2004): Size small (SVL 15.43 mm in male; 16.02 mm in female); habitus slender, head longer than wide (HL/HW 1.10 in male, 1.15 in female); snout round, slightly protruding beyond margin of lower jaw, longer than diameter of eye (SL/ED 1.56 in male; 1.65 in female); eyes protuberant, pupil round; dorsal surface of head flat; canthus rostralis round, distinct; loreal region oblique, flat; nostril round, lateral, on canthus rostralis, closer to tip of snout than to eye (NED/SL 0.60 in male, 0.56 in female); interorbital distance wider than internarial distance (IOD/IND 1.61 in male, 1.62 in female) and wider than upper eyelid width (IOD/UEW 1.85 in male and female); tympanum hidden; vomerine teeth absent, tongue round posteriorly, free for approximately two thirds its length; slit like openings to a median vocal sac.</p>
            <p>Forelimbs relatively short, shorter than snout vent length (FLL/SVL 0.74 in male, 0.78 in female); hand shorter than half of forelimb length (HAL/FLL 0.40 in male, 0.38 in female); fingers slender, free of webbing, dorsoventrally flattened, fingers with weak skin fringes on finger II and III; nuptial pad absent on male finger I; partially free nub just proximal to finger II (1FLO/2FLO 0.20 in male, 0.21 in female); finger II slightly longer than finger IV length (2FLI/4FLI 1.06 in male, 1.04 in female), and much shorter than the third (2FLI/3FLI 0.60 in male and female); relative finger lengths: I&lt;II≤IV&lt;III; tip of finger I round, not enlarged; narrow peripheral grooves present on fingers II, III and IV; grooves on dorsal surface of disks present in fingers II, III, IV, absent in finger I; relative finger disk widths II&lt;IV&lt;III; subarticular tubercles on fingers distinct, round, formula 1:1:2:1 (given for fingers I:II:III:IV, respectively); two round, flat palmar tubercle.</p>
            <p>Hindlimbs slender, twice as long as snout-vent length (HLL/SVL 1.92 in male, 2.07 in female), tibia longer than half of snout-vent length (TL/SVL 0.63 in male, 0.68 in female); tibiotarsal articulation reaching beyond snout when limb adpressed along body; foot longer than tibia (FL/TL 1.43 in male, 1.38 in female); relative toe lengths: I&lt;II&lt;V&lt;III&lt;IV; tarsus smooth, inner tarsal fold absent; tips of all toes slightly dilated, forming truncated disks, disks of all toes with peripheral grooves, dorsal surface of toe disks II–V with short median longitudinal grooves; relative toe disk widths I&lt;V&lt;II&lt;III=IV; webbing formula I1–2II1–2½III2½–2½IV2½–1V; subarticular tubercles on toes distinct, round, formula 1:1:2:3:2 (for toes I:II:III:IV:V, respectively); inner metatarsal tubercle oval; outer metatarsal tubercle absent.</p>
            <p>Skin on dorsum relatively smooth with low, round tubercles concentrated on the outer edge, flanks, below and posterior to eye; dorsolateral folds distinct in adult male; cloaca unmodified, directed posteriorly, at upper level of thighs; supratympanic fold indistinct; lateral side of head smooth; ventral surface of body and limbs smooth, vent area smooth.</p>
            <p>Coloration in life. Dorsal surface greyish brown to light brown with a dark brown or brown spear-shape marking, running from the posterior eye to the groin; black spear-shape marking border, or just two black spots on either side of the spear-shape marking; a black broken streak below dorsolateral folds; black streak-shape marking present on the groin region in adult male; a light brown streak from posterior corner of eye to arm insertion; dorsolateral surfaces of arm and fingers dark brown with a small black bar near the wrist; dorsal surfaces of thigh, tibia, tarsus and toes brown with dark brown cross bars on each hindlimb; throat and chest dark grey with intense small white mottling, belly and limbs ventrally lighter with white marbling mixed with black spots; pupil black, iris copper (Fig. 2).</p>
            <p>Coloration in preservative. In ethanol, dorsal coloration changed to dark greyish brown, ventral surface of chest, belly, and limbs greyish beige; dorsal pattern, dark spots on dorsum and stripes on dorsal surface of limbs unchanged, dark brown pattern changed to dark grey; pupil white, iris completely black (Fig. 3).</p>
            <p> Extended diagnosis.  Microhyla nanapollexa is characterized by having a small body size (SVL 15.43 mm, n = 1 male; 16.02–16.63 mm, n= 2 females); body habitus slender; snout round; skin on dorsum relatively smooth with low, round tubercles concentrated on the outer edge, flanks, below and posterior to eye; venter smooth; vertebral skin ridge and dorsomedial stripe absent; superciliary tubercles absent; supratympanic fold indistinct; fingers slender, free of webbing, with weak dermal fringes on fingers II and III; nuptial pad absent on male finger I; partially free nub just proximal to finger II; grooves on dorsal surface of disks present in fingers II – IV; two round, flat palmar tubercles; hindlimbs long, tibiotarsal articulation of adpressed limb reaching well beyond snout; toe webbing formula I1–2 II1–2 ½ III2 ½–2½ IV 2 ½–1 V; inner metatarsal tubercle oval; outer metatarsal tubercle absent; dorsal surface greyish brown to light brown with a dark brown or brown spear shape marking, running from the posterior eye to the groin; a black broken streak below dorsolateral folds; black streak shape marking absent or present on the groin region; belly and limbs ventrally lighter with white marble mixed with black spots; pupil black, iris copper in life. </p>
            <p> Distribution.  Microhyla nanapollexa is currently known from the type locality in Tra My District, Quang Nam Province and Kon Plong District, Kon Tum Province, Vietnam at elevations between 1260 and 1480 m a.s.l. </p>
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	https://treatment.plazi.org/id/0389433CFFFCAE58FF27E33D7F068FEB	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hoang, Chung Van;Pham, Cuong The;Nguyen, Truong Quang;Phan, Tien Quang;Ziegler, Thomas;Orlov, Nikolai;Jiang, Jianping;Do, Dang Trong	Hoang, Chung Van, Pham, Cuong The, Nguyen, Truong Quang, Phan, Tien Quang, Ziegler, Thomas, Orlov, Nikolai, Jiang, Jianping, Do, Dang Trong (2025): A new species of Microhyla (Amphibia: Anura: Microhylidae), with an extended description of Microhyla nanapollexa from Vietnam. Zootaxa 5566 (2): 347-369, DOI: 10.11646/zootaxa.5566.2.6, URL: https://doi.org/10.11646/zootaxa.5566.2.6
0389433CFFF1AE51FF27E6C478EA88D1.text	0389433CFFF1AE51FF27E6C478EA88D1.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Microhyla roedderi Hoang & Pham & Nguyen & Phan & Ziegler & Orlov & Jiang & Do 2025	<html xmlns:mods="http://www.loc.gov/mods/v3">
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            <p> Microhyla roedderi sp. nov.</p>
            <p>LSID: urn:lsid:zoobank.org:act: 0299FDAF-0B60-4FF6-A2AC-F34550C27F42</p>
            <p>(Figs. 4, 5; Table 2, Table supplement 2)</p>
            <p> Holotype. Adult male IEBR A.6271 (Field number KH2022.26), collected by D.T. Do et al. on 8 April 2022 in Van Ninh District, Khanh Hoa Province, Vietnam (12°49'49.0"N 109°11'55.8"E, at an elevation of 784 m a.s.l.).</p>
            <p>
                 Paratypes (n= 12). Nine adult males IEBR A.6272–6280 (Field numbers KH2022.25, KH2022.28, KH2022.29– 35) and two adult females IEBR A.6281,.6282 (Field numbers KH2022.24, KH2022.27), collection data the same as the holotype ;   one adult female IEBR A.6283 (Field number PY2022.67), collected in  
                <a title="Search Plazi for locations around (long 109.21533/lat 12.858611)" href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=109.21533&amp;materialsCitation.latitude=12.858611">Tay Hoa District</a>
                 , Phu Yen Province, Vietnam (12°51'31.0"N 109°12'55.2"E, at an elevation of 433 m a.s.l., collected by D.T. Do et al. on 5 April 2022. 
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            <p> Diagnosis.  Microhyla roedderi sp. nov. is distinguished from other species of the subgenus  Nanohyla by a combination of the following characters: Size medium (SVL 15.88–17.00 mm, n= 10 in males; 20.74–21.33 mm, n= 3 in females); body habitus slender; snout bluntly round; tympanum hidden; canthus rostralis round, distinct; loreal region oblique, weakly concave; dorsal skin slightly bumpy with low tubercles, slightly concentrated on dosolateral area, scattered over dorsal surface of limbs and upper eyelid; ventral surface smooth; mid vertebral skin ridge and dorsomedial stripe absent; superciliary tubercles absent; supratympanic fold weakly developed; fingers slender, free of webbing, with weak skin fringes on all fingers; finger I reduced slightly, less than half of finger II in length, males without nuptial pad; grooves on dorsal surface of disks present in fingers II–IV; two round, flat palmar tubercles; hindlimbs long, tibiotarsal articulation of adpressed limb reaching well beyond snout; toe webbing formula I1–2II1–2III1–2IV2–1V; inner metatarsal tubercle oval; outer metatarsal tubercle absent; dorsal surface of head and trunk greyish brown to light brown with dark brown markings, a distinct light brown interorbital bar between eyelids, a&gt;–&lt;shape marking running from the eyelids to the groin; few small black spots on the flank and in the inguinal region; two small black spots in the groin region; belly and limbs ventrally lighter with white marbling; pupil black, iris copper in life. </p>
            <p>Description of holotype. Size medium (SVL 17.0 mm); habitus slender, head longer than wide (HL/HW 1.01); snout bluntly rounded, slightly protruding beyond margin of lower jaw, longer than diameter of eye (SL/ED 1.03); eyes protuberant, pupil round; dorsal surface of head flat; canthus rostralis round, distinct; loreal region oblique, weakly concave; nostril round, lateral, on canthus rostralis, closer to tip of snout than to eye (NED/SL 0.56); interorbital distance wider than internarial distance (IOD/IND 1.50), twice broader than upper eyelid (IOD/UEW 2.16); tympanum hidden, supratympanic fold weakly developed, from posterior corner of eye to arm insertion; vomerine teeth absent, tongue round posteriorly, free at the rear half of its length; slit like openings to a median vocal sac.</p>
            <p>Forelimbs shorter than snout-vent length (FLL/SVL 0.85); hand shorter than a half of forelimb length (HAL/ FLL 0.43); fingers slender, free of webbing, with weak dermal fringes on all fingers; finger I reduced notably, less than half the length of finger II (1FLO/2FLO 0.29); nuptial pad absent; finger II slightly longer than finger IV (2FLI/4FLI 1.04), much shorter than finger III (2FLI/3FLI 0.69); relative finger lengths: I&lt;IV&lt;II&lt;III; tip of finger I round, not enlarged; narrow peripheral grooves absent on all fingers; grooves on dorsal surface of disks present in fingers II, III, IV, absent in finger I; relative finger disk widths IV&lt;II&lt;III; subarticular tubercles on fingers distinct, round, formula 1:1:2:1 (given for fingers I:II:III:IV, respectively); two round, flat palmar tubercles.</p>
            <p>Hindlimbs slender, almost twice as long as snout-vent length (HLL/SVL 1.98), tibia longer than half of snout-vent length (TL/SVL 0.65); tibiotarsal articulation reaching beyond snout when limb adpressed along body; foot longer than tibia (FL/TL 1.39); relative toe lengths: I&lt;II&lt;V&lt;III&lt;IV; tarsus smooth, inner tarsal fold absent; tips of all toes slightly dilated, forming truncated disks, disks of all toes with peripheral grooves, dorsal surface of toe disks II, V with short median longitudinal grooves; relative toe disk widths: I&lt;IV&lt;V&lt;II&lt;III; webbing formula I1–2II1– 2III1–2IV2–1V; subarticular tubercles on toes distinct, round, formula 1:1:2:3:2 (for toes I:II:III:IV:V, respectively); inner metatarsal tubercle oval; outer metatarsal tubercle absent.</p>
            <p>Dorsal skin bumpy with low tubercles, slightly concentrated on the dorsolateral area, scattered over dorsal surface of limbs and upper eyelid; supratympanic fold weakly developed, from posterior corner of eye to arm insertion; lateral side of head and flank smooth; ventral side of body and limbs smooth, vent area smooth.</p>
            <p>Coloration in life. Dorsal surface of head and trunk greyish brown to light brown with dark brown markings, a distinct light brown interorbital bar between eyelids, a&gt;–&lt;shape marking running from the eyelids to the groin; few small black spots on the flank and in the inguinal region; two small black spots in the groin region; supratympanic fold light brown; dorsolateral surface of arm and fingers dark brown with a small black bar near the wrist; dorsal surface of thigh, tibia, tarsus and toes brown with dark brown cross bars; throat and chest brownish grey with intense small white mottling, belly and limbs ventrally lighter with white marbling; pupil black, iris copper (Fig. 4).</p>
            <p>Coloration in preservative. In ethanol, dorsal coloration changed to dark greyish brown, ventral surface of chest, belly, and limbs greyish beige; dorsal pattern, dark spots on dorsum and stripes on dorsal surfaces of limbs unchanged, dark brown pattern changed to dark grey; iris completely black (Fig. 5).</p>
            <p>Variation. Type specimens vary in body size. Three female paratypes have a larger body size than those of ten males (SVL 20.95 ± 0.33 mm [20.74–21.33 mm; n=3] vs. 16.43± 0.35 mm [15.88–17.00 mm; n=10] in males). Dark brown marking on dorsum is more distinctly lobed in two paratypes (IEBR A.6281, A.6283). Black spots in the flank and inguinal region are various in size (smaller in IEBR A.6283). For more details see Table 2 and Fig. 6.</p>
            <p> Natural history. Specimens were collected at night, from 19:00 to 23:00 h, on the ground nearby streams in evergreen forest. The surrounding habitat was evergreen forest with large, medium and small hardwoods mixed with shrubs and vines (Fig. 7), at elevations between 433 and 784 m a.s.l. Air temperatures at the sites ranged from 20.5–28.5 oC and relative humidity was 60–80%. Other amphibian species found at the sites included  Limnonectes limborgi (Sclater) ,  Limnonectes poilani (Bourret) ,  Hylarana sp. ,  Odorrana cf. banaorum (Bain, Lathrop, Murphy, Orlov &amp; Ho) ,  Kurixalus sp. , and  Hylarana attigua (Inger, Orlov &amp; Darevsky) . Larval stages and eggs of the new species are unknown. </p>
            <p> Comparison.  Microhyla roedderi sp. nov. most closely resembles  M. nanapollexa in morphology, the close relationship being supported by molecular phylogenetic position (Fig. 1, Table 3). However, the new species differs from the latter by having a larger body size (SVL 15.98–17.00 mm in males, 20.74–21.33 mm in females vs. SVL 15.43 mm in male, 16.02–16.60 mm in females of  M. nanapollexa ); dorsal skin bumpy with low tubercles, concentrated on the dorsolateral area, scattered over the dorsal surface of limbs and upper eyelid (Figs. 4A; 6A; 6B) (vs. skin on dorsum relatively smooth with low, round tubercles concentrated on the outer edge, flanks, below and posterior to eye in  M. nanapollexa (Fig. 2A); finger I reduced notably, less than half the length of finger II (Fig. 8B) (vs. partially free nub just proximal to finger II in  M. nanapollexa (Fig. 8A); different webbing formula I1–2II1–2III1–2IV2–1V (vs. I1–2II1–2½III2½–2½IV 2½–1V in  M. nanapollexa ) (Table S1); dorsal surface of head and trunk greyish brown to light brown with dark brown markings, a distinct light brown interorbital bar between eyelids, a&gt;–&lt;shape marking running from the eyelids to the groin (vs. dorsal surface of greyish brown to light brown with a dark brown or brown spear shape marking, running from the posterior eyes to the groin in  M. nanapollexa ); and few small black spots on the flank and inguinal region (vs. a black broken streak below dorsolateral folds in  M. nanapollexa ). </p>
            <p>......continued on the next page</p>
            <p> Microhyla roedderi sp. nov. differs from other known species in the subgenus  Nanohyla by having a larger body size (SVL 15.88–17.00 mm in males, 20.74–21.33 mm in females vs. SVL 13.6–14.7 mm in males, 18.3–18.63 mm in females of  M. hongiaoensis ; vs. SVL 14.4–15.6 mm in males, 18.2–18.4 mm in females of  M. annectens ; vs. SVL 13.2–15.0 mm in males, 15.9–17.0 mm in females of  M. arboricola ; vs. SVL 10.5–11.9 mm in males, 12.4–14.5 mm in females of  M. perparva ; vs. SVL 13.9–16.2 mm in males, 15.1–17.8 mm in females of  M. petrigena ); by having a smaller body size (SVL 15.9–17.0 mm in males, 20.7–21.3 mm in females vs. SVL 18.2–20.2 mm in males of  M. albopunctata ; vs. SVL 18.8–21.5 mm in males, 21.1–23.2 mm in females of  M. marmorata ); by having snout bluntly round (vs. pointed in  M. arboricola ; obtusely pointed in  M. perparva ,  M. petrigena ); by having dorsum skin slightly bumpy (vs. smooth in  M. annectens ,  M. perparva ,  M. pulchella ); by having FI &lt;½ FII (vs. nub or bulge in  M. perparva ,  M. petrigena ); by having dorsal median longitudinal grooves on finger disks present (vs. absent in  M. hongiaoensis ); by having one metatarsal tubercle (vs. two metatarsal tubercles in  M. albopunctata ,  M. annamensis ,  M. hongiaoensis ,  M. marmorata ); by superciliary tubercles absent (vs. superciliary tubercles present in  M. perparva ). </p>
            <p> Distribution.  Microhyla roedderi sp. nov. is currently known only from the type locality in Deo Ca Mountain, Khanh Hoa and Phu Yen provinces, Vietnam, at elevations between 433 and 784 m a.s.l. </p>
            <p> Conservation status.  Microhyla roedderi sp. nov. is likely to be endemic to Deo Ca Mountain, central Vietnam. However, the extent of its actual distribution range should be confirmed in further studies. Given the available information, we suggest this species be considered as Data Deficient following IUCN’s Red List categories (IUCN 2001). </p>
            <p>Etymology. The new species is named after Ass. Prof. Dr. Dennis Rödder, Leibniz Institut zur Analyse des Biodiversitätswandels (LIB) / Zoologisches Forschungsmuseum Alexander Koenig (ZFMK), Bonn, to honour his long-term support of conservation-based research including the support of student work and providing his expertise in species distribution modelling. We recommend “Roedder’s Narrow-mouth Frog” as the common English name and “Nhái bầu roedder” as the Vietnamese name.</p>
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	https://treatment.plazi.org/id/0389433CFFF1AE51FF27E6C478EA88D1	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Hoang, Chung Van;Pham, Cuong The;Nguyen, Truong Quang;Phan, Tien Quang;Ziegler, Thomas;Orlov, Nikolai;Jiang, Jianping;Do, Dang Trong	Hoang, Chung Van, Pham, Cuong The, Nguyen, Truong Quang, Phan, Tien Quang, Ziegler, Thomas, Orlov, Nikolai, Jiang, Jianping, Do, Dang Trong (2025): A new species of Microhyla (Amphibia: Anura: Microhylidae), with an extended description of Microhyla nanapollexa from Vietnam. Zootaxa 5566 (2): 347-369, DOI: 10.11646/zootaxa.5566.2.6, URL: https://doi.org/10.11646/zootaxa.5566.2.6
