identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
039787E5C374FF9F6E7CFECEFC6A58AD.text	039787E5C374FF9F6E7CFECEFC6A58AD.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heuchera tuckasegeensis Pantinople 2024	<div><p>Heuchera tuckasegeensis Pantinople,  sp. nov.</p><p>TYPE:   USA. Transylvania Co.,  above Cold Mountain Road several miles from Toxaway, 4 October 2013, R.A. Folk 230 (holotype: OS!)  .</p><p>Heuchera tuckasegeensis resembles  Heuchera parviflora var. parviflora but is distinguished by its sparsely puberulent petioles (vs. densely villous), deeper leaf sinuses (0.9–1.3 vs. 0.6–0.7 per unit leaf length), and shorter styles (1.4–1.6 mm vs. 2.2–2.3 mm) and stamens (1.3–1.6 mm vs. 1.9–2.1 mm).</p><p>Perennial herb, acaulescent. Petioles sparsely puberulent to nearly glabrous (trichomes 0.25–0.35[–0.65] mm long), 8.86–17.51 cm long. Leaf blades 4.3–8.7 cm wide, 3.1–5.3(–7.9) cm long (from convergence of veins to apex), reniform, shallowly to deeply cordate, relatively deeply divided into 5 primary lobes; sinuses between leaf lobes 3.88–5.21 mm deep; teeth obtusely triangular to crenate, 1.79–2.5 mm long, 5.53–9.22 mm wide; abaxial lamina remotely viscid, strigose; adaxial lamina similarly vestitured (trichomes 0.13–0.45 mm long); leaf margin obscurely ciliate. Inflorescences thyrses (racemes of scorpioid cymes), 20.84–27.07 cm long, diffuse, interrupted, with 1–3 leaf-like bracts along the primary axis of the thyrse below the most proximal cymule, and a reduced bract at each cymule and flower node, 2.68–4.83 mm long, primary axis puberulent; cymules 2–7 flowered, 2.11–3.34 cm long, diffuse (pedicels of lower cymule first order flower 5.06–8.52 mm long), lower cymules dichasial but distal-most cymules becoming monochasial. Flowers 1.82–2.68 mm long, (1.37–) 1.50–1.81 mm wide (at the bottom of hypanthium), white, campanulate, external surface villous, orientation pendent; sepals triangular, 0.74–1.82 mm long; petals narrowly spatulate with a long claw, recurved, 1.92–2.66 mm long; stamens 1.28–1.55 (2.33) mm long; styles 1.39–1.62 (2.01) mm long, stigmas capitate. Capsules 4.22–5.47 mm long, 2–2.58 mm wide, ovoid. Seeds (0.39–) 0.45–0.62 mm long, widely elliptic, black, almost smooth with testa cells arranged in low ridges, the margins obscurely undulate. Figures 6, 7.</p><p>Phenology —Flowers from June to October and fruits from June (no specimens were available to confirm the end of the fruiting period).</p><p>Distribution —Southwestern tip of North Carolina, USA, in Jackson County along the east fork of the Tuckasegee River, and in Transylvania County around Panthertown Valley near Cold Mountain and Lake Toxaway (Fig. 8).</p><p>Habitat —Rockhouses behind waterfalls, shaded areas, rock outcrops, crevices, and ledges on cliffs, on highly acidic gneiss, (805–) 1073–1280 m.</p><p>Conservation —Currently only known from seven localities across a narrow range of approximately 10 km; a species of interest for conservation listing given its rarity, regional endemism, and genetic distinctness, but still warrants further field data.</p><p>Associates —  Leucothoe axillaris D.Don,  Parnassia asarifolia Vent.,  Houstonia serpyllifolia Graham,  Oxypolis rigidior (L.) Raf.,  Thalictrum clavatum Hook,  Eurybia divaricata (L.) G.L. Nesom,  Luzula echinata (Small) F.J.Herm.,  Kalmia latifolia L.,  Viburnum nudum var. cassinoides (L.) Torr. &amp; A.Gray,  Asplenium montanum Willd.,  Vittaria appalachiana Farrar &amp; Mickel,  Heuchera villosa Michx., and species in the liverwort order  Jungermanniales .</p><p>Etymology —Anglicized Cherokee (“turtle place”), named for the Tuckasegee River, corresponding to the primary distribution of this taxon. Epithet to be pronounced in six syllables (i.e. “tuckasege€ ensis”).</p><p>Other Specimens Examined —   USA. — NORTH CAROLINA: Jackson Co., Maidenhair Falls above  Lake Fairfield, 9 August 1951, R.K. Godfrey 51818 (DUKE) ;   Jackson Co., Bonas’ Defeat,  East fork of  Tuckasegee River, 23 August 1951, R.K. Godfrey 52017 (NY) ;   Jackson Co., Tuckasegee River gorge,  Bonas’ Defeat cliffs, 20 July 1972, D. Pitillo 9361 (WCUH) ;   Jackson Co., Bonas’ Defeat cliffs,  Tuckasegee River gorge, 7 July 1973, D. Pitillo 4591 (WCUH) ;   Jackson Co., Wolf Creek gorge of  Canada Community, 805 m, 6 April 1974, D. Pitillo 6543 (WCUH) ;   Jackson Co., near Bonas’ Defeat,  East fork of  Tuckasegee River, 15 September 1974, E.F. Wells 3279 (NCU) ;   Jackson Co., South slopes of  Shelton Pisgah Mountain along  Little Green Creek, north of  Cold Mountain Gap, 1159 m, 25 November 1989, D. Pitillo 10538 (WCUH) ;   Jackson Co., Panthertown Valley on Shelton Pisgah Mountain,  Northwest end above  Little Greenland Creek, 1128 m, 1 August 1990, D. Pitillo 10659 ;   Jackson Co., North of Camp Merrie Woode at  Fairfield Lake,  Twin Falls grotto on  Trays Island Creek, 1073 m, 17 May 1991, D. Pitillo 10753 (WCUH) ;   Jackson Co.,  Wolf Creek Falls below dam, 854 m, 23 June 1994, D. Pitillo 11691 (WCUH) ;   Transylvania Co., near  Camp Toxaway on Cold Mountain Road, 7 August 1949, A.E. Radford 4819 (NCU) ;   Transylvania Co.,  Showerbath Falls East of Cold Mountain Gap, 1098 m, 15 July 1988, D. Pitillo 9984 (WCUH) ;   Transylvania Co., Panthertown Valley,  Cold Mt., 25 May 1990, S. Wiser 90212 (NCU) ;   Transylvania Co., Panthertown Valley,  Cold Mountain, 1189 m, 10 June 1993, D. Pitillo 11255 (WCUH) ;   Transylvania Co.,  Shower Falls on Cold Mountain Road, 23 April 2006, E. Schwartzman s. n. (UNCA)  .</p></div>	https://treatment.plazi.org/id/039787E5C374FF9F6E7CFECEFC6A58AD	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pantinople, Dexcem J.;Engle-Wrye, Nicholas J.;Folk, Ryan A.	Pantinople, Dexcem J., Engle-Wrye, Nicholas J., Folk, Ryan A. (2024): Heuchera tuckasegeensis sp. nov. (Saxifragaceae), a New Species from Western North Carolina. Systematic Botany (Basel, Switzerland) 49 (1): 37-47, DOI: 10.1600/036364424X17110456120677, URL: https://doi.org/10.1600/036364424x17110456120677
039787E5C370FF9F6C8AF9A4FEC95AED.text	039787E5C370FF9F6C8AF9A4FEC95AED.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Heuchera parviflora Bartl.	<div><p>KEY TO THE SPECIES AND VARIETIES OF THE  HEUCHERA PARVIFLORA GROUP</p><p>(partly based on Folk and Freudenstein 2015)</p><p>1. Petioles puberulent.................................................................................................................... 2</p><p>1. Petioles villous........................................................................................................................ 3</p><p>2. Stamens exserted. 1.7 mm, styles exserted. 1.7 mm, laminar hair length. 0.5 mm, leaf coriaceous, growing on calcareous alkaline substrates (dolomite or limestone)............................................................................................  H. puberula</p><p>2. Stamens exserted almost always, 1.7 mm, styles exserted almost always, 1.7 mm, laminar hair length, 0.6 mm, leaf membraneous, growing on highly acidic substrates (gneiss)................................................................................  H. tuckasegeensis</p><p>3. Flower orientation subhorizontal.................................................................................  H. missouriensis</p><p>3. Flower orientation pendent................................................................................................... 4</p><p>4. Leaf suede-like, not viscid at maturity, growing almost exclusively on sandstone substrates..............  H. parviflora var. parviflora</p><p>4. Leaf viscid at maturity, growing exclusively on quartzite substrates....................................  H. parviflora var. saurensis</p><p>Note: Flower orientation is recommended to be measured in life as it can be prone to ambiguous determination using herbarium specimens.</p></div>	https://treatment.plazi.org/id/039787E5C370FF9F6C8AF9A4FEC95AED	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		Plazi	Pantinople, Dexcem J.;Engle-Wrye, Nicholas J.;Folk, Ryan A.	Pantinople, Dexcem J., Engle-Wrye, Nicholas J., Folk, Ryan A. (2024): Heuchera tuckasegeensis sp. nov. (Saxifragaceae), a New Species from Western North Carolina. Systematic Botany (Basel, Switzerland) 49 (1): 37-47, DOI: 10.1600/036364424X17110456120677, URL: https://doi.org/10.1600/036364424x17110456120677
