taxonID	type	description	language	source
039DBD0D7850FFACFF28FB62FDECF99A.taxon	diagnosis	Diagnosis (adapted from Rodríguez et al. 2012, modifications on bold): Enthemonae with basilar muscles and mesogleal marginal sphincter. Pedal disc broad, usually elongated in the transverse plane, often grasping. Column commonly with thick mesoglea, smooth or with small mesogleal papillae distally arranged in longitudinal rows. Cinclides absent or very rare (1 – 3), distally over the directives endocoels, with the external opening mounted on a papilla. Tentacles usually marginal on wide oral disc, their aboral sides sometimes thickened. Oral disc sometimes lobed. Mesenteries not divisible into macro- and micro-cnemes. At least six pairs of perfect and fertile mesenteries. Retractor muscles diffuse, weak. Longitudinal muscles of the tentacles ectodermal. Acontia highly reduced, being absent in many preserved specimens, when present with only basitrichs. Cnidom: spirocysts, basitrichs, pmastigophores B 1, and holotrichs.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7850FFADFF28F9B6FE9DFEF9.taxon	diagnosis	Diagnosis (adapted from Carlgren 1949, modifications on bold): Amphianthidae with well-developed base often elongate in the transversal axis, often attached to gorgonians, sponges or worm-tubes. Column rather low, smooth or with longitudinal rows of mesogleal papillae in its distal part. Cinclides, if present, few, occurring only over the directive chambers. Sphincter well developed, mesogleal. Tentacles possibly more numerous than the mesenteries at the base or the same number; fairly short, with or without mesogleal thickenings at the base on their outer sides. Longitudinal muscles of tentacles ectodermal. Radial muscles of oral disc ectodermal to ecto-mesogleal. Two distinct siphonoglyphs. Perfect pairs of mesenteries 6 – 12. Retractors and parietobasilar muscles weak. Gonads on all stronger mesenteries except possibly the directives. Acontia few or perhaps sometimes absent. Cnidom: spirocysts, basitrichs, p - mastigophores B 1.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7850FFADFF28F9B6FE9DFEF9.taxon	type_taxon	Type species: Actinia abyssicola Moseley, 1877	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7851FFA9FF28F972FEA0FE69.taxon	description	(Figs. 2, 3; Tables 1, 2)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7851FFA9FF28F972FEA0FE69.taxon	materials_examined	MATERIAL EXAMINED. Holotype: MNRJ 9080 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83 (04 ° 28 ' 21 '' S, 36 ° 24 ' 45 '' W), May 4, 2011, 1950 m. Paratypes: MNRJ 9078 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83 - 2 (04 ° 27 ' 01 '' S, 36 ° 25 ' 36 '' W), May 21, 2011, 2123 m. MNRJ 9091 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 84 (04 ° 25 ' 49 '' S, 36 ° 37 ' 22 '' W), May 6, 2011, 1964 m. MOUFPE-CNI 875 (one specimen), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 84 (04 ° 25 ' 49 '' S, 36 ° 37 ' 22 '' W), May 6, 2011, 1964 m. LC 153 (one specimen attached on Diacria sp.), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 85 (04 ° 21 ' 21 '' S, 36 ° 44 ' 16 '' W), May 6, 2011, 2057 m.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7851FFA9FF28F972FEA0FE69.taxon	diagnosis	DIAGNOSIS Body short, cylindrical, with wide, grasping base. Tubercles in 24 longitudinal rows at distal scapus. Tentacles 96, in four cycles. Number of tentacles is the same as number of mesenteries at base. Mesenteries of first three cycles perfect, except the directives. Acontia mostly inconspicuous, may be absent in some individuals.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7851FFA9FF28F972FEA0FE69.taxon	description	DESCRIPTION EXTERNAL MORPHOLOGY. In preservation, body cylindrical, short, 90 - 150 mm in height (100 mm in holotype) and 80 - 250 mm in diameter (250 mm in holotype). Color pale orange to pale pink / purple in preserved specimens. Base wide, variable in shape: circular in outline or more oval when grasping shells or rocks (Fig. 2 A). Column divisible into scapus and scapulus. Mesoglea thick (Fig. 2 B). Distal portion of scapus with distinct tubercles arranged in 24 longitudinal rows, 3 or 4 per row. Longitudinal ridges between each tubercule row, 12 in total (Fig. 2 C). Proximal part with weak remains of brownish cuticle, soft depressions and reduced tubercules. All tubercules dome-shaped, varying only in size with smaller tubercles proximally. Scapulus concealed by retraction of oral disc. Two cinclides distally, on opposite sides of column, on mesogleal papillae above directive endocoels (Fig. 2 C). Oral disc with central oval mouth, two visible siphonoglyphs (Fig. 2 B). Tentacles smooth, short and filiform, some with thickening at base (Fig. 2 F), 96 in total and arranged in 4 - 5 cycles (Fig. 2 H). INTERNAL ANATOMY. Two siphonoglyphs, each attached to a directive pair of mesenteries. Sphincter mesogleal, strong, alveolar, nearer endoderm than ectoderm, occupying most of mesoglea proximal to oral disc (Fig. 2 E). Longitudinal muscles of tentacles ectodermal. Mesenteries slender, visible as rows through column. On holotype, four cycles of mesenteries hexamerously arranged 6 + 6 + 12 + 24, total of 48 pairs (Fig. 2 D) near base; and 48 mesenteries near mouth (Fig. 2 H). First cycle perfect. Mesenterial arrangement constant among examined individuals. All cycles well-developed with exception of last cycle (Fig. 2 I), with mesenteries shorter, barely breaking through the mesoglea, and some with tiny mesenterial filaments. Mesenterial filaments more developed on older mesenteries. Retractors diffuse, often weak, stronger near the mesenterial base (Fig. 2 G). Parietobasilar muscles poorly developed. Gametogenic tissue present in the first three cycles of mesenteries except directives. Acontia inconspicuous and poorly developed. CNIDOM. Spirocysts, basitrichs, and microbasic p - mastigophores B 1 (Fig. 3, Table 2).	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7851FFA9FF28F972FEA0FE69.taxon	materials_examined	TYPE LOCALITY. Potiguar Basin, Rio Grande do Norte, Brazil. BIOLOGICAL CHARACTERISTICS. Sediment found on the stations ranged from rocks to mud. Paratype LC 153 attached to snails of the genus Diacria. Some of the paratypes were attached to rocks, with small fragments of rocks and broken shells attached to the pedal disc.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7851FFA9FF28F972FEA0FE69.taxon	etymology	ETYMOLOGY. From latin “ prima ” = the first, so named because this is the first species of the genus recorded on the South Atlantic and in the XXI century.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7851FFA9FF28F972FEA0FE69.taxon	discussion	REMARKS / COMPARISON. Of the 11 valid species within the genus Stephanauge, seven are reported in the Atlantic Ocean (Table 1). Stephanauge prima sp. nov. possesses a unique set of characters that differentiates it from its counterparts in this region. Stephanauge prima sp. nov. primarily differs from S. nexilis and S. abyssicola in possessing more tentacles and mesenteries and bearing no gametogenic tissue on the younger cycle of mesenteries (only mesenteries of first cycle are fertile in S. nexilis; only mesenteries of first cycle are sterile in S. abyssicola). Stephanauge inornata, and S. spongicola also have fewer tentacles (60 in S. inornata vs. maximum of 80 in S. spongicola vs. 96 in S. prima sp. nov.) and neither has tubercles on the distal part of the column. Stephanauge prima sp. nov. can be distinguished from S. annularis, because the latter has a smooth column and all cycles of mesenteries fertile, except the directive ones (vs. a tuberculated column and the directives and fourth cycle of mesenteries sterile in S. prima sp. nov.). Stephanauge impedita has similar morphological features to S. prima sp. nov., but has fewer cycles of tentacles (3 vs. 4 in S. prima sp. nov.) and longer tentacles in the outer vs inner cycle, and lacks acontia. The remaining species, S. acanellae, found on Acanella sp. branches, has more numerous tentacles (140 vs. 96 in S. prima sp. nov.) and cycles of mesenteries (5 vs. 4 in S. prima sp. nov.), and has no tubercles on the column and no trace of acontia.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7855FFA6FF28FC34FB63FF41.taxon	description	(Fig. 4; Table 3)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7855FFA6FF28FC34FB63FF41.taxon	materials_examined	MATERIAL EXAMINED. MNRJ 9077 (two specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 72 (04 ° 40.1817 ’ S, 036 ° 23.8647 ’ W), May 7, 2011, 908 m. MOUFPE-CNI 878 (two specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 72 (04 ° 40.1817 ’ S, 036 ° 23.8647 ’ W), May 7, 2011, 908 m. LC 157 (one specimen) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 72 - 2 (04 ° 40.2981 ' S, 036 ° 23.7001 ’ W), May 20, 2011, 960 m. MNRJ 9083 (two specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 73 (04 ° 37.6640 ' S, 036 ° 30.5400 ’ W), May 5, 2011, 957 m. LC 158 (one specimen) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 73 (04 ° 37.6640 ' S, 036 ° 30.5400 ’ W), May 5, 2011, 957 m. LC 159 (three specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 74 (04 ° 34.1484 ' S, 036 ° 41.6035 ’ W), May 5, 2011, 902 m. MOUFPE-CNI 877 (two specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 75 (04 ° 28.8002 ' S, 036 ° 52.5554 ' W), May 3, 2011, 915 m. LC 154 (two specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 84 (04 ° 25.8308 ’ S, 036 ° 37.3678 ’ W), May 6, 2011, 1964 m. LC 155 (two specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 85 (04 ° 21.3580 ’ S, 036 ° 036 ° 44.2730 ’ W), May 4, 2011, 2057 m.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7855FFA6FF28FC34FB63FF41.taxon	description	SHORT DESCRIPTION. Individuals 5.6 to 8.7 mm in height and 16.6 to 24.1 mm in diameter. Number of tentacles corresponds to the number of mesenteries: 48 pairs of mesenteries and 48 tentacles arranged in four cycles. Most of the sampled specimens were attached to Acanella sp. and branches of Chrysogorgia tuberculata Cordeiro, Castro & Pérez, 2015. CNIDOM. Spirocysts, basitrichs, holotrichs, and microbasic p - mastigophores B 1 (Fig. 4 I: a – e, Table 3).	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7855FFA6FF28FC34FB63FF41.taxon	distribution	GEOGRAPHIC DISTRIBUTION. North Pacific Ocean 2000 - 5285 m (Hertwig 1888; Carlgren 1949), North Atlantic 1075 – 4851 m (Riemann-Zürneck 1987; Iken et al. 2001; OBIS 2023). This is the first record for A. bathybium in the South Atlantic (Potiguar basin, RN, Brazil) 900 – 2000 m.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7855FFA6FF28FC34FB63FF41.taxon	discussion	REMARKS. The individuals analyzed agree well with the previous descriptions of A. bathybium (e. g., Hertwig 1882; 1888; Riemann-Zürneck 1987). The morphological descriptions and cnidom of the individuals from the Potiguar Basin are highly similar to those provided by Riemman-Zürneck (1987) in her redescription of the type specimen of A. bathybium. Although the types and average sizes for cnidae that we recorded match fairly well with what Riemman-Zürneck (1987) described, we found an extra type of holotrich in the tentacle and smaller microbasic p - mastigophores B 1 in the filament. The holotrich is rare and was not found in all specimens.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7855FFA9FF28FDE0FD8FFC1D.taxon	diagnosis	Diagnosis (see Carlgren 1949)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7855FFA9FF28FDE0FD8FFC1D.taxon	type_taxon	Type species: Amphianthus bathybium Hertwig, 1882	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785AFFA7FF28FB1AFD46FF6D.taxon	description	(Fig. 4; Table 4)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785AFFA7FF28FB1AFD46FF6D.taxon	materials_examined	MATERIAL EXAMINED. MNRJ 9083 (six specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 85 (04 ° 21.3580 ' S, 036 ° 44.2730 ’ W), May 8, 2011, 2057 m. MOUFPE-CNI 876 (ten specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 63 (04 ° 41.7490 ’ S, 036 ° 31.1670 ’ W), May 8, 2011, 375 m. LC 160 (one specimen) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 63 (04 ° 41.7490 ’ S, 036 ° 31.1670 ’ W), May 8, 2011, 375 m. LC 161 (seven specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 85 (04 ° 21.3580 ’ S, 036 ° 036 ° 44.2730 ’ W), May 4, 2011, 2057 m.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785AFFA7FF28FB1AFD46FF6D.taxon	description	SHORT DESCRIPTION. Small individuals, 3 to 9 mm in height and 4 to 11 mm in diameter. Body short, flat, or globose with 24 rows of small tubercles. Around 70 tentacles, relatively long, in four alternating cycles. Up to two cinclides, each one associated with a directive endocoels. Mesenteries irregularly arranged in four cycles, more numerous at base than at margin. Acontia not numerous. CNIDOM. Spirocysts, basitrichs, holotrichs, and microbasic p - mastigophores B 1 (Fig. 4 I: f – j, Table 4).	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785AFFA7FF28FB1AFD46FF6D.taxon	distribution	GEOGRAPHIC DISTRIBUTION. Amphianthus michaelsarsi is known only from its type material, ten individuals from the Michael Sars Expedition (1910), studied by Carlgren (1934), and from Molodtsova et al. (2008), both from the Mid-Atlantic Ridge in the North Atlantic. The present work expands its geographical distribution to the South Atlantic (Potiguar basin, RN, Brazil).	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785AFFA7FF28FB1AFD46FF6D.taxon	discussion	REMARKS. The morphological investigation of A. michaelsarsi from Potiguar Basin matches the description from North Atlantic surveys made by Carlgren (1934), with minor differences in cnidom. Carlgren (1934) does not differentiate the nematocysts of the mesenterial filaments, which we observed to be basitrichs and microbasic p - mastigophores B 1. We found holotrichs in the tentacles and some slightly larger basitrichs in the acontia of our individuals not reported by Carlgren (1934).	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785BFFA7FF28FB65FE8AFA22.taxon	diagnosis	Diagnosis (see Gusmão & Rodríguez 2021)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785BFFA7FF28FB65FE8AFA22.taxon	type_taxon	Type species: Actinernus saginata Verrill, 1882 Valid species: Actinoscyphia aurelia (Stephenson, 1918); Actinoscyphia groendyki Eash-Loucks & Fautin, 2012; Actinoscyphia plebeia (McMurrich, 1893); Actinoscyphia saginata (Verrill, 1882); Actinoscyphia verrilli (Gravier, 1918)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785BFFA4FF28FA29FCF4FE15.taxon	description	(Fig. 4, Table 5)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785BFFA4FF28FA29FCF4FE15.taxon	materials_examined	MATERIAL EXAMINED. MNRJ 9092 (two specimens), South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83 (04 ° 28.3642 ’ S, 036 ° 24.7602 ’ W), May 4, 2011, 1950 m. MOUFPE-CNI 874 (two specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83 (04 ° 28.3642 ’ S, 036 ° 24.7602 ’ W), May 21, 2011, 1950 m. LC 156 (10 specimens) South Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 83 - 2 (04 ° 40.1817 ’ S, 036 ° 23.8647 ’ W), May 21, 2011, 2123 m.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785BFFA4FF28FA29FCF4FE15.taxon	description	SHORT DESCRIPTION. Column 35.6 to 56.7 mm in height, oral disc diameter 72.4 to 114.5 mm, base diameter 21.6 to 43.5 mm. Oral disc bilobed. Tentacles with mesogleal swellings at their base, total approximately 160, in two cycles. Mesenteries in five cycles, only first one perfect. Acontia absent. CNIDOM. Spirocysts, basitrichs, holotrichs, and microbasic p - mastigophores B 1 (Fig. 4 I: k – o, Table 5).	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785BFFA4FF28FA29FCF4FE15.taxon	distribution	GEOGRAPHIC DISTRIBUTION. Actinoscyphia saginata was originally known from the Atlantic coast of North America (Verrill 1882) and later described from France, Morocco and Brazil (Riemann-Zürneck 1978; Gusmão & Rodríguez 2021), at depths ranging from 978 to 2177 m. Our findings extend its distributional records northwards off the Brazilian coast, suggesting a more continuous Atlantic range for this species.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D785BFFA4FF28FA29FCF4FE15.taxon	discussion	REMARKS. In Riemann-Zürneck’s (1978) review of Actinoscyphia, the cnidom was interpreted as an important characteristic to distinguish species. Following the identification key proposed by Eash-Loucks & Fautin (2012), Actinoscyphia saginata has approximately 160 long tentacles, which differentiates it from closer species, such as Actinoscyphia aurelia, with around 275 tentacles and more than five mesentery cycles. The attributes of our specimens fit well to the original description of Actinoscyphia saginata by Verrill (1882) to later treatments by Riemann-Zürneck (1978) and Gusmão & Rodríguez (2021).	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7858FFA4FF28FB11FE7CFA24.taxon	diagnosis	Diagnosis (see Gusmão & Rodríguez 2021)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7858FFA4FF28FB11FE7CFA24.taxon	type_taxon	Type species: Actinauge nodosa var. coronata Verrill, 1883	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7858FFA2FF28FA21FC6EFF41.taxon	description	(Fig. 4; Table 6)	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7858FFA2FF28FA21FC6EFF41.taxon	materials_examined	MATERIAL EXAMINED. MNRJ 9079 (one specimen). Atlantic Ocean, Potiguar Basin, Rio Grande do Norte, Brazil, station MT 72 (04 ° 40 ' 10 '' S, 36 ° 23 ' 51 '' O), May 7, 2011, 897 - 908 m.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7858FFA2FF28FA21FC6EFF41.taxon	description	DESCRIPTION. Small, elongated body, almost perfectly cylindrical, column 15.1 mm in height and 6 mm in diameter at mid-column, divisible into scapus and short scapulus (Fig. 4 B). Tentacles without mesogleal thickenings at base, arranged in five cycles, 72 in total. Mesenteries hexamerously arranged in four cycles. Acontia present, but rare. CNIDOM. Spirocysts, basitrichs, microbasic p - mastigophores B 1 (Fig. 4 I: p – s, Table 6).	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7858FFA2FF28FA21FC6EFF41.taxon	distribution	GEOGRAPHIC DISTRIBUTION. Chondrophellia coronata is known from the North Atlantic, Gulf of Mexico, and from the North and South Pacific (Carlgren 1942; Doumenc & Van Präet 1988; McMurrich 1893; Molodtsova et al. 2008). In Brazil, it was recorded off the coast of Rio de Janeiro, Southwest region (Gusmão & Rodríguez 2021). The collection of specimens at the Potiguar Basin, in the northeast of the country, indicates a broader or even Panamerican distribution for this species.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
039DBD0D7858FFA2FF28FA21FC6EFF41.taxon	discussion	REMARKS. The description of C. coronata from the Potiguar Basin is in agreement with former descriptions of this species. We found an additional larger microbasic p - mastigophore B 1 in the filaments not reported by other authors but suspect that it is the result of contamination from acontia.	en	Targino, Alessandra K. G., De Melo, Yago A., Gomes, Paula B. (2025): Deep-sea anemones (Cnidaria, Actiniaria) from off the northeastern coast of Brazil: new records and description of Stephanauge prima sp. nov. Zootaxa 5569 (2): 328-344, DOI: 10.11646/zootaxa.5569.2.6, URL: https://doi.org/10.11646/zootaxa.5569.2.6
