identifier	taxonID	type	CVterm	format	language	title	description	additionalInformationURL	UsageTerms	rights	Owner	contributor	creator	bibliographicCitation
03A787FE7B35FFF7FF7BACABFA5E570C.text	03A787FE7B35FFF7FF7BACABFA5E570C.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Corynoppia Balogh 1983	<div><p>Generic diagnosis of  Corynoppia</p><p>With character states of  Oppiidae (Norton and Behan-Pelletier 2009) . Measurements: Length about 200–380. Integument. Body nearly smooth or tuberculate/granulate (partially or entirely); sometimes covered by specific tuberculate, ridge- or band-like cerotegument; lateral side of body between bothridium and acetabula densely tuberculate. Prodorsum. Rostrum rounded. Costular-transcostular complex reduced or slightly visible, nearly trapezoid. Lateral arch-like prodorsal ridge usually present. Interbothridial region with two longitudinal ridges and two or three pairs of muscle sigillae between them; interbothridial tubercle absent; postbothridial tubercle usually present. Rostral seta medium-sized, setiform or phylliform; lamellar seta medium-sized, phylliform, located closer to in than to ro; interlamellar seta short, setiform or medium-sized, phylliform; bothridial seta long, clavate/unilaterally fusiform/nearly setiform, barbed or with shortly ciliated; exobothridial seta short, setiform. Notogaster. Anterior margin narrowed medially. Humeral region usually with humeral tubercle and crista. Ten pairs of setae: c short, setiform or phylliform; other setae medium-sized, phylliform.  Gnathosoma . Subcapitulum diarthric. Palp setation: 0-2-1-3-9(+ω); solenidion long, bacilliform, pressed to surface, located in middle of dorsal part of tarsus. Chelicera chelate-dentate. Epimeral and lateral podosomal regions. Epimere IV of normal length; border IV present. Epimeral setal formula: 3-1-3-3; all setae setiform. Ventrosejugal tubercle absent. Pedotectum I represented by small lamina. Discidium present. Anogenital region. Anogenital setal formula: 5-1-2-3; adanal setae setiform or phylliform; other anogenital setae setiform. Adanal seta ad 1 posterior, ad 2 posterior or posterolateral, ad 3 anterolateral to anal plate; distance ad 3 – ad 3 longer than ag–ag and ad 2 – ad 2. Adanal lyrifissure located close and parallel to anal aperture. Legs. Tibia I with anterodorsal projection. Tarsus II with two solenidia. Sexual dimorphism: sometimes present in morphology of adanal setae and leg segments.</p><p>Note: Roman letters refer to normal setae; Greek letters to solenidia; single prime (′) marks setae on the anterior and double prime (′′) setae on the posterior side of a given leg segment; parentheses refer to a pair of setae.</p><p>General remarks on  Corynoppia species</p><p>Subías (2004, online version 2011) included  Striatoppia weigmanni Murvanidze &amp; Behan-Pelletier, 2011 in  Corynoppia . The genera  Striatoppia Balogh 1958 and  Corynoppia are morphologically similar to each other, but the former differs from the latter in having two strong longitudinal prodorsal ridges between rostral and lamellar setae (versus ridges completely absent) and the presence (versus absence) of striae on the notogaster (usually also in anogenital region).  Striatoppia weigmanni has no ridges between rostral and lamellar setae as well as true striae on the body (only cerotegumental bands visible), therefore, the placement of this species in  Corynoppia seems correct.</p><p>Subías (2022, online version 2024) supported the initial generic placement of  Striatoppia margaritata Mahunka, 1969, but Ermilov and Rybalov (2025) transferred it in  Corynoppia due absence of the longitudinal prodorsal ridges between rostral and lamellar setae as well as striae on the body.</p><p>Key to known species of  Corynoppia</p><p>1. Bothridial seta clavate, with small, rounded distally head..................................................... 2</p><p>- Bothridial seta unilaterally fusiform, with elongate, narrowed distally head, or nearly setiform........................ 3</p><p>2. Sexual dimorphism in morphology of the adanal seta ad 1 present (long, slightly dilated mediobasally, with setiform distal part in males versus comparatively short, broadly phylliform in females); notogastral seta c narrowly phylliform, other notogastral setae frequently bifid; body length: 325–338..........  Corynoppia margaritata (Mahunka, 1969) . Distribution: Tanzania.</p><p>- Sexual dimorphism in morphology of the adanal seta ad 1 absent (phylliform in males and females); notogastral seta c setiform, other notogastral setae not bifid; body length: 345–360..................................................................................  Corynoppia dhatiensis Ermilov, 2025 (in Ermilov &amp; Rybalov 2025). Distribution: Ethiopia.</p><p>3. Interlamellar seta medium-sized, broadly phylliform......................................................... 4</p><p>- Interlamellar seta short, setiform......................................................................... 5</p><p>4. Adanal seta ad 2 setiform; notogastral seta c narrowly phylliform, inserted medial to humeral tubercle; two pairs of muscle sigillae between interlamellar prodorsal ridges; notogaster and anogenital region with cerotegumental bands; body length: 250–280.......................  Corynoppia weigmanni (Murvanidze &amp; Behan-Pelletier, 2011) . Distribution: Caucasus.</p><p>- Adanal seta ad 2 broadly phylliform; notogastral seta c setiform, inserted on humeral tubercle; three pairs of muscle sigillae between interlamellar prodorsal ridges; notogaster and anogenital region without cerotegumental bands; body length: 270– 300.....................................................  Corynoppia gamoensis sp. nov. Distribution: Ethiopia.</p><p>5. Adanal seta ad 2 setiform............................................................................... 6</p><p>- Adanal seta ad 2 broadly phylliform....................................................................... 8</p><p>6. Rostral seta broadly phylliform; three pairs of muscle sigillae between interlamellar prodorsal ridges; body length: 311......................................................  Corynoppia maritima Pérez-Íñigo jr., 1991. Distribution: Spain.</p><p>- Rostral seta nearly setiform; two pairs of muscle sigillae between interlamellar prodorsal ridges...................... 7</p><p>7. Aggenital seta and adanal seta ad 3 located in one transverse row (ad 3 lateral to ag); body length: 315–338...............  Corynoppia kosarovi kosarovi (Jeleva, 1962) (in Csiszár &amp; Jeleva 1962; see also Mahunka &amp; Paoletti 1984; Baran &amp; Gökyeşil 2015). Distribution: Mediterranean, Iran, Panama.</p><p>- Aggenital seta and adanal seta ad 3 located in one diagonal row (ad 3 posterolateral to ag); body length: 325–335............................................  Corynoppia kosarovi matritensis (Pérez-Íñigo, 1967) . Distribution: Mediterranean.</p><p>8. Adanal seta ad 3 setiform............................................................................... 9</p><p>3</p><p>- Adanal seta ad narrowly phylliform..................................................................... 11</p><p>9. Rostral seta nearly setiform; bothridial seta nearly setiform; body length: 345–380............................................................................  Corynoppia hispanica Subías &amp; Shtanchaeva, 2011 . Distribution: Spain.</p><p>- Rostral seta broadly phylliform; bothridial seta with unilaterally fusiform head................................... 10</p><p>10. Notogastral seta c narrowly phylliform; body covered by rugose-like cerotegument; body length: 270–285...................................................  Corynoppia papillisetigera Iturrondobeitia &amp; Saloña, 1998 . Distribution: Spain.</p><p>- Notogastral seta c setiform; body not covered by rugose-like cerotegument; body length: 249–274................................................................  Corynoppia foliatoides Subías &amp; Rodríguez, 1986 . Distribution: Spain.</p><p>11. Rostral seta broadly phylliform; body length: 265–330...........................  Corynoppia foliata (Mihelčič, 1957) (see also Arillo &amp; Subías 1996; Pérez-Íñigo 1971; Subías &amp; Arillo 2001). Distribution: Spain.</p><p>- Rostral seta nearly setiform............................................................................ 12</p><p>12. Anogenital region partially striate; rostral region without ridges; body length: 207....................................................................  Corynoppia andulau sakaryaensis Baran &amp; Gökyeşil, 2015 . Distribution: Turkey.</p><p>- Anogenital region not striate; rostral region with two short ridges; body length: 252–268..........................................................................  Corynoppia andulau andulau Mahunka, 2001 . Distribution: Borneo.</p></div>	https://treatment.plazi.org/id/03A787FE7B35FFF7FF7BACABFA5E570C	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2025): Taxonomic contribution to knowledge of the oribatid mite genera Corynoppia, Ramusella and Wallworkella (Acari, Oribatida, Oppiidae). Zootaxa 5621 (3): 353-370, DOI: 10.11646/zootaxa.5621.3.4, URL: https://doi.org/10.11646/zootaxa.5621.3.4
03A787FE7B36FFF4FF7BA847FBA8563D.text	03A787FE7B36FFF4FF7BA847FBA8563D.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Corynoppia gamoensis Ermilov 2025	<div><p>Description of  Corynoppia gamoensis sp. nov.</p><p>(Figures 1, 2)</p><p>Type material.  Holotype (female) and  three paratypes (one male and two females): southern Ethiopia, South Ethiopia Regional State, Gamo Zone, 6°01’36.7’’N, 37°35’42.7’’E, 1170 m a.s.l., 5 km East from the town Arba Minch, 1 km from the lake Abai, mineral soil (0–5 cm) in a mixed forest on the terrace (first layer of forest:  Celtis sp.,  Ficus sycomorus,  Schefflera sp.,  Acacia sp.; second layer of forest: Lemon sp.,  Coffee arabica) on the brown (black-brown) soil with light-medium loam soil texture, 7.XII.2018, dry season (L.B. Rybalov).</p><p>Five paratypes (three males and two females): southern Ethiopia, South Ethiopia Regional State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.597137&amp;materialsCitation.latitude=6.024639" title="Search Plazi for locations around (long 37.597137/lat 6.024639)">Gamo Zone</a>, 6°01’28.7’’N, 37°35’49.7’’E, 1179 m a.s.l., 5 km East from the town Arba Minch, 20–40 m from the lake Abai, mineral soil (0–5 cm) in a mixed deciduous forest on the alluvial organic soil, wet place, around water spots, 8.XII.2018, dry season (L.B. Rybalov)  .</p><p>The holotype is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz, Germany; eight paratypes are deposited in the collection of the University of Tyumen, Museum Zoology, Tyumen, Russia. All specimens are preserved in 70% solution of ethanol with a drop of glycerol.</p><p>Diagnosis. Body length: 270–300. Prodorsum with sparse tubercles; notogaster and anogenital region not tuberculate. Interbothridial region with three pairs of muscle sigillae. Rostral seta medium-sized, nearly setiform, barbed; lamellar and interlamellar setae medium-sized, broadly phylliform, barbed; bothridial seta long, with unilaterally fusiform and shortly ciliate head. Humeral tubercle developed. Notogastral seta c short, setiform, roughened, inserted on humeral tubercle; other notogastral setae medium-sized, broadly phylliform (sometimes some setae slightly bifid), barbed. Epimeral, genital and aggenital setae setiform, nearly smooth; both anal setae and adanal seta ad 3 setiform, slightly barbed; adanal setae ad 1 and ad 2 broadly phylliform (sometimes slightly bifid), barbed. Anterodorsal part of leg tibia IV with broad dilatation in males.</p><p>Description of adult. Measurements. Body length: 300 (holotype), 270–300 (male paratypes), 285–300 (female paratypes); notogastral width: 157 (holotype), 135–150 (male paratypes), 143–157 (female paratypes).</p><p>Integument (Figs 1A, C; 2B). Body color light brown to grayish. Body nearly smooth, but prodorsum and subcapitular mentum sparsely tuberculate; region between bothridium and acetabula I–III with typical dense tubercles; leg femora I–IV and trochanter III, IV partially densely tuberculate; all tubercles small (their diameter up to 2).</p><p>Prodorsum (Figs 1A, C). Rostrum broadly rounded. Costular-transcostular complex elongate trapezoid, poorly developed. Interbothridial longitudinal ridges and three pairs of muscle sigillae between them well visible. Lateral ridge medium-sized, slightly arch-like. Rostral seta (26–28) nearly setiform, barbed; lamellar and interlamellar setae (17–19) broadly phylliform, narrowed distally, barbed; exobothridial seta (5) setiform, roughened; bothridial seta (52–56) with long, smooth stalk and shorter, unilaterally fusiform and shortly ciliate head. Postbothridial tubercle present; slight tubercle-like thickening located opposite to postbothridial tubercle.</p><p>Notogaster (Figs 1A, C; 2A). Anterior margin of notogaster slightly convex and shortly interrupted medially. Humeral region with tubercle and crista. Notogastral seta c (5) setiform, roughened, inserted in medial part of humeral tubercle; other setae (22–26) broadly phylliform, narrowed distally, barbed; sometimes some setae slightly bifid. Opisthonotal gland opening and all lyrifissures well visible.</p><p>Gnathosoma (Figs 2B–D). Subcapitulum size: 60–64 × 45–49; subcapitular setae a (13–15) and h (13–15) setiform, roughened; m (15) setiform, barbed; all adoral setae (6) setiform, smooth. Palp length: 45–49; with typical setation: 0-2-1-3-9(+ω); postpalpal seta (4) spiniform, smooth. Chelicera length: 60–64; setae (cha: 19; chb: 13–15) setiform, barbed.</p><p>Epimeral and lateral podosomal regions (Figs 1B, C). Epimeres with typical epimeral setal formula (3-1-3-3); setae (11–15) setiform, nearly smooth. Discidium triangular, with rounded top.</p><p>Anogenital region (Figs 1B, C; 2A). With typical anogenital setal formula (5-1-2-3); genital (7–9) and aggenital (11–15) setae setiform, nearly smooth; both anal setae (19–22) and adanal seta ad 3 (22) setiform, slightly barbed; adanal setae ad 1 (22) and ad 2 (22) broadly phylliform, narrowed distally (sometimes slightly bifid), barbed. Adanal lyrifissure distinct, in typical position for genus (close and parallel to lateral side of anal plate).</p><p>Legs (Figs 2E–H). Claw of each leg smooth. Porose area on all leg segments not visible.Anterodorsal projection well developed on tibia I. Trochanter III with posterior tooth. Sexual dimorphism in morphology of tibia IV present: in males, anterodorsal part of segment with broad dilatation (Fig. 2H) versus in females, segment without dilatation. Formulas of leg setation and solenidia: I (1-5-2-4-20) [1-2-2], II (1-5-2-4-16) [1-1-2], III (2-3-1-3-15) [1-1-0], IV (1-2-2-3-12) [0-1-0]; homology of setae and solenidia indicated in Table 1; seta s on tarsus I eupathidial; some setae narrowly or broadly phylliform; solenidion ω 2 on tarsus I rod-like; φ 1 on tibia I subflagellate; other solenidia nearly thickened, rounded distally.</p><p>Remarks.  Corynoppia gamoensis sp. nov. is similar is similar to  C. weigmanni (Murvanidze &amp; Behan-Pelletier, 2011) from Caucasus in the presence of the unilaterally fusiform bothridial seta and medium-sized, broadly phylliform interlamellar seta. However, the new species differs from the latter by the morphology of the adanal seta ad 2 (phylliform versus setiform), the morphology and location of the notogastral seta c (setiform, inserted on humeral tubercle versus phylliform, inserted medial to humeral tubercle), the number of the interbothridial muscle sigillae (three pairs versus two pairs), and the absence (versus presence) of the cerotegumental bands on the notogaster and in the anogenital region.</p><p>Etymology. The species name  gamoensis refers to the place of origin, Gamo Zone.</p></div>	https://treatment.plazi.org/id/03A787FE7B36FFF4FF7BA847FBA8563D	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2025): Taxonomic contribution to knowledge of the oribatid mite genera Corynoppia, Ramusella and Wallworkella (Acari, Oribatida, Oppiidae). Zootaxa 5621 (3): 353-370, DOI: 10.11646/zootaxa.5621.3.4, URL: https://doi.org/10.11646/zootaxa.5621.3.4
03A787FE7B32FFF8FF7BACF9FE075490.text	03A787FE7B32FFF8FF7BACF9FE075490.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ramusella (Insculptoppia) arbaminchensis Ermilov 2025	<div><p>Description of  Ramusella (Insculptoppia) arbaminchensis sp. nov.</p><p>(Figures 3, 4)</p><p>Type material. Holotype (male) and  six paratypes (two males and four females): southern Ethiopia, South Ethiopia Regional State, Gamo Zone, 6°01’36.7’’N, 37°35’42.7’’E, 1170 m a.s.l., 5 km East from the town Arba Minch, 1 km from the lake Abai, mineral soil (0–5 cm) in a mixed forest on the terrace (first layer of forest:  Celtis sp.,  Ficus sycomorus,  Schefflera sp.,  Acacia sp.; second layer of forest: Lemon sp.,  Coffee arabica) on the brown (black-brown) soil with light-medium loam soil texture, 7.XII.2018, dry season (L.B. Rybalov).</p><p>One paratype (one male): southern Ethiopia, South Ethiopia Regional State, Gamo Zone, 6°01’28.7’’N, 37°35’49.7’’E, 1179 m a.s.l., 5 km East from the town Arba Minch, 20–40 m from the lake Abai, mineral soil (0–5 cm) in a mixed deciduous forest on the alluvial organic soil, wet place, around water spots, 8.XII.2018, dry season (L.B. Rybalov).</p><p>The holotype is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz, Germany; seven paratypes are deposited in the collection of the University of Tyumen, Museum Zoology, Tyumen, Russia. All specimens are preserved in 70% solution of ethanol with a drop of glycerol.</p><p>Diagnosis. Body length: 255–300. Costular line short, thin; transcostular line absent. Interbothridial region with three pairs of muscle sigillae. Rostral and interlamellar setae medium-sized, setiform, slightly barbed; lamellar and exobothridial setae short, setiform, roughened; bothridial seta long, with clavate head bearing four (sometimes three) long, setiform, barbed branches and three to five short ciliae. Notogastral seta c represented by alveolus; la, lm, lp, h 1 – h 3 long, p 1 – p 3 short, all setiform, sparsely barbed or nearly smooth. Surface of chelicera with ridges. Epimeral and anogenital setae setiform, nearly smooth or roughened. Ventrosejugal region with one pair of tubercles. Lateral longitudinal carina in podosomal region with tooth.</p><p>Description of adult. Measurements. Body length: 255 (holotype), 255–285 (male paratypes), 270–300 (female paratypes); notogastral width: 135 (holotype), 135–150 (male paratypes), 150–157 (female paratypes).</p><p>Integument (Fig. 3C). Body color light brown. Body nearly smooth; region between bothridium and acetabula I–III with typical dense tubercles (their diameter up to 2).</p><p>Prodorsum (Figs 3A, C). Rostrum rounded. Costular line short, thin; transcostular line absent. Interbothridial region with three pairs of muscle sigillae. Lateral ridge absent. Rostral (30–34) and interlamellar (30–37) setae setiform, slightly barbed; lamellar (15–19) and exobothridial (11) setae setiform, roughened; bothridial seta (49–52) with long, smooth stalk and shorter, clavate head bearing four (sometimes three) long, setiform, barbed branches and three to five short cilia. Postbothridial and sejugal tubercles not visible. Sejugal porose area (5–7) present, circular.</p><p>Notogaster (Figs 3A, C; 4A). Anterior margin of notogaster convex medially. Notogastral seta c represented by alveolus; other setae (la, lm, lp, h 1 – h 3: 34–45; p 1 – p 3: 17–22) setiform, sparsely barbed or nearly smooth. Opisthonotal gland opening and all lyrifissures well visible.</p><p>Gnathosoma (Figs 4B–D). Subcapitulum size: 60–64 × 45–49; subcapitular setae a (9–11) and h (13–15) setiform, roughened; m (13–15) setiform, barbed; all adoral setae (6) setiform, smooth. Palp length: 41–49; with typical setation: 0-2-1-3-9(+ω); postpalpal seta (4) spiniform, smooth. Chelicera length: 60–64; seta cha (17–19) setiform, dorsally ciliate; chb (11–13) setiform, barbed; surface of chelicera with four ridges (Fig. 4D).</p><p>Epimeral and lateral podosomal regions (Figs 3B, C). Epimeres with typical epimeral setal formula (3-1-3- 3); setae (1a, 2a, 3a: 7; 1c, 3b, 4a, 4b: 11) setiform, nearly smooth; others (1b, 4c: 17–19; 3c: 22–26) setiform, roughened. Ventrosejugal region with one pair of strong tubercles (each tubercle located posterior to insertion of 2a). Lateral longitudinal carina with tooth (well visible in ventral aspect; Fig. 3B). Discidium triangular, with pointed top.</p><p>Anogenital region (Figs 3B, C; 4A). With typical anogenital setal formula (5-1-2-3); genital (5) and anal (11–13) setae setiform, nearly smooth; aggenital and adanal setae (17–22) setiform, roughened. Adanal lyrifissure distinct, in typical position for genus (close and parallel to lateral side of anal plate).</p><p>Legs (Figs 4E–H). Claw of each leg smooth. Porose area on all leg segments not visible. Trochanter III with two or three posterior teeth. Formulas of leg setation and solenidia: I (1-5-2-4-20) [1-2-2], II (1-5-2-4-16) [1-1-2], III (2-3-1-3-15) [1-1-0], IV (1-2-2-3-12) [0-1-0]; homology of setae and solenidia indicated in Table 1; seta s on tarsus I eupathidial; setae a ′′and pv ′′on tarsus IV modified (brush-like); solenidia ω 1 on tarsi I, ω 1 and ω 2 on tarsus II and σ on genu III nearly bacilliform, φ 1 on tibia I and φ on tibia IV subflagellate; other solenidia setiform or rod-like.</p><p>Remarks.  Ramusella (Insculptoppia) arbaminchensis sp. nov. is similar to  R. (I.) rybalovi Ermilov, Shtanchaeva &amp; Subias, 2022 from Ethiopia in the presence of the barbed branches on the bothridial seta as well as long dorsal notogastral setae and strong tubercles in ventrosejugal region. However, the new species differs from the latter by the morphology of the bothridial seta (with well developed head versus without head), the absence (versus presence) of the transcostula, postbothridial tubercle and prodorsal lateral ridge, and in the presence (versus absence) of the tooth on lateral carina in the podosomal region.</p><p>Etymology. The species name  arbaminchensis refers to the town Arba Minch, five kilometers from which the new species was collected.</p></div>	https://treatment.plazi.org/id/03A787FE7B32FFF8FF7BACF9FE075490	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2025): Taxonomic contribution to knowledge of the oribatid mite genera Corynoppia, Ramusella and Wallworkella (Acari, Oribatida, Oppiidae). Zootaxa 5621 (3): 353-370, DOI: 10.11646/zootaxa.5621.3.4, URL: https://doi.org/10.11646/zootaxa.5621.3.4
03A787FE7B3DFFFBFF7BAE1BFC3A5010.text	03A787FE7B3DFFFBFF7BAE1BFC3A5010.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Ramusella	<div><p>General remarks on  Ramusella species</p><p>The nominate subgenus of  Ramusella and the subgenus R. (Insculptoppia) differs from each other mainly by the morphology of the rostral setae (Subías 1980): geniculate in R. ( Ramusella) versus not geniculate in R. (Insculptoppia). The analysis of literature on species within these subgenera has revealed an incorrect systematic placement of some species.</p><p>According to original descriptions, supplementary descriptions and existing figures,  R. baichengensis (Zhao &amp; Wen, 1992),  R. confusa Arillo &amp; Subías, 1990,  R. cordobensis (Balogh &amp; Mahunka, 1968),  R. filamentosa (Mahunka, 1985),  R. filigera (Mahunka, 1985),  R. gyrata (Mahunka &amp; Paoletti, 1984),  R. hainardorum (Mahunka, 1992),  R. persica Akrami, Behmanesh &amp; Subías, 2015,  R. rybalovi Ermilov, Shtanchaeva &amp; Subías, 2022, and  R. tasetata Subías, 1980 were considered by Subías (2022, online version 2024) within the nominate subgenus, however, their rostral setae are arch-like or nearly straight (not geniculate), therefore, all these species must be included in R. (Insculptoppia). Hugo-Coetzee (2016) described rostral setae as geniculate in  R. filamentosa and  R. filigera, but supplementary descriptions of these species did not include figures.</p><p>Ramusella tobiasi Hugo-Coetzee, 2016 was described by Hugo-Coetzee (2016) within the nominate subgenus, however, Subías (2022, online version 2024) included it in R. (Insculptoppia). The species has geniculate rostral setae, therefore, its initial subgeneric placement position must be maintained.</p></div>	https://treatment.plazi.org/id/03A787FE7B3DFFFBFF7BAE1BFC3A5010	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2025): Taxonomic contribution to knowledge of the oribatid mite genera Corynoppia, Ramusella and Wallworkella (Acari, Oribatida, Oppiidae). Zootaxa 5621 (3): 353-370, DOI: 10.11646/zootaxa.5621.3.4, URL: https://doi.org/10.11646/zootaxa.5621.3.4
03A787FE7B3EFFFFFF7BAD07FE3D54D6.text	03A787FE7B3EFFFFFF7BAD07FE3D54D6.taxon	http://purl.org/dc/dcmitype/Text	http://rs.tdwg.org/ontology/voc/SPMInfoItems#GeneralDescription	text/html	en	Wallworkella Balogh 1983	<div><p>Generic diagnosis of  Wallworkella</p><p>With character states of  Oppiidae (Norton and Behan-Pelletier 2009) . Measurements: Length about 240–390.</p><p>Integument. Body nearly smooth; sometimes tuberculate band present instead mediobasal part of costula; lateral side of body between bothridium and acetabula densely tuberculate. Prodorsum. Rostrum tripartite, sometimes rounded or concave. Costulae and transcostula fused, forming arch-like or trapezoid complex; rarely, complex indistinct or costulae developed partially. Lateral arch-like prodorsal ridge usually present. Interbothridial region usually with two or three pairs of muscle sigillae. Interbothridial and postbothridial tubercles absent but prodorsobasal part sometimes with short ridges or tubercle-like thickenings. Rostral, lamellar, interlamellar, and exobothridial setae well developed, setiform; ro and le short to long, le and in short; le inserted closer to in than to ro. Bothridial seta long, ciliate (its mediodistal part with well developed unilaterally lanceolate expansion) or pectinate (its mediodistal part without expansion or with indistinct expansion). Notogaster. Anterior margin convex or straight medially.</p><p>Humeral region without tooth and crista. Ten pairs of setiform setae: c short; other setae short to long.  Gnathosoma .</p><p>Subcapitulum diarthric. Palp setation: 0-2-1-3-9(+ω); solenidion long, bacilliform, pressed to surface, located in mediodistal part of dorsal part of tarsus. Chelicera chelate-dentate. Epimeral and lateral podosomal regions.</p><p>Epimere IV of normal length; border IV present. Epimeral setal formula: 3-1-3-3; all setae setiform. Ventrosejugal tubercle absent. Pedotectum I represented by small lamina. Discidium present. Anogenital region. Anogenital setal formula: 6-1-2-3; all setae setiform. Adanal seta ad 1 posterior, ad 2 lateral, ad 3 anterolateral to anal plate, distance ad 3 – ad 3 longer than ag–ag and ad 2 – ad 2. Adanal lyrifissure located close and parallel to anal aperture. Legs. Tibia I without anterodorsal projection. Tarsus II with two solenidia.</p><p>Description of  Wallworkella southensis sp. nov.</p><p>(Figures 5, 6)</p><p>Type material.  Holotype (female) and  four paratypes (one male and three females): southern Ethiopia, South Ethiopia Regional State, <a href="https://tb.plazi.org/GgServer/search?materialsCitation.longitude=37.595192&amp;materialsCitation.latitude=6.026861" title="Search Plazi for locations around (long 37.595192/lat 6.026861)">Gamo Zone</a>, 6°01’36.7’’N, 37°35’42.7’’E, 1170 m a.s.l., 5 km East from the town Arba Minch, 1 km from the lake Abai, mineral soil (0–5 cm) in a mixed forest on the terrace (first layer of forest:  Celtis sp.,  Ficus sycomorus,  Schefflera sp.,  Acacia sp.; second layer of forest: Lemon sp.,  Coffee arabica) on the brown (black-brown) soil with light-medium loam soil texture, 7.XII.2018, dry season (L.B. Rybalov)  .</p><p>The holotype is deposited in the collection of the Senckenberg Museum of Natural History, Görlitz, Germany; four paratypes are deposited in the collection of the University of Tyumen, Museum Zoology, Tyumen, Russia. All specimens are preserved in 70% solution of ethanol with a drop of glycerol.</p><p>Diagnosis. Body length: 240–255. Rostrum rounded. Costulae and transcostula not visible. Lateral ridge long. Rostral seta medium-sized, setiform, slightly barbed; lamellar and interlamellar setae short, setiform, roughened; bothridial seta long, with unilaterally lanceolate head bearing setiform apex and three long, setiform, smooth branches and one short (distal) cilium unilaterally. Anterior margin of notogaster straight. Dorsal notogastral setae comparatively short, setiform, sparsely barbed or nearly smooth. Epimeral and anogenital setae short, setiform, smooth (except roughened 3c and 4c). Discidium with broadly rounded top.</p><p>Description of adult. Measurements. Body length: 255 (holotype), 240 (male paratype), 240–255 (female paratypes); notogastral width: 120 (holotype), 105 (male paratype), 113–120 (female paratypes).</p><p>Integument (Fig. 5C). Body color light brown. Body nearly smooth; region of costula and region between bothridium and acetabula I–III with typical dense tubercles (their diameter up to 2).</p><p>Prodorsum (Figs 5A, C). Rostrum rounded. Costular-transcostular complex absent, but its vague outlines are sometimes partially hardly visible. Interbothridial muscle sigillae not visible. Lateral ridge long. Rostral seta (19–22) setiform, slightly barbed; lamellar (7–9), interlamellar (11–13) and exobothridial (7–9) setae setiform, roughened; bothridial seta (45–49) with long, smooth stalk and shorter, unilaterally lanceolate head bearing setiform apex and three long, setiform, smooth branches and one short (distal) cilium unilaterally.</p><p>Notogaster (Figs 5A, C; 6A). Anterior margin of notogaster straight. Notogastral seta c (4–6) setiform, smooth; other setae (la, lm, lp, h 1 – h 3, p 2, p 33: 17–19; p 1: 11–13) setiform, sparsely barbed or nearly smooth. Opisthonotal gland opening and lyrifissures ia, im, ip well visible (ih, ips not found).</p><p>Gnathosoma (Figs 6B–D). Subcapitulum size: 56–60 × 41–45; all subcapitular setae (11) setiform, nearly smooth; all adoral setae (4) setiform, smooth. Palp length: 37; with typical setation: 0-2-1-3-9(+ω); postpalpal seta (4) spiniform, smooth. Chelicera length: 52–56; setae (cha: 17–19; chb: 11) setiform, barbed.</p><p>Epimeral and lateral podosomal regions (Figs 5B, C). Epimeres with typical epimeral setal formula (3-1-3-3); setae 3c (11–15) and 4c (9–11) setiform, roughened; others (7) setiform, smooth. Discidium with broadly rounded top.</p><p>Anogenital region (Figs 5B, C; 6A). With typical anogenital setal formula (6-1-2-3); genital (4–6), aggenital (11–15), anal (11–15), and adanal (11–15) setae setiform, smooth. Adanal lyrifissure distinct, in typical position for genus (close and parallel to lateral side of anal plate).</p><p>Legs (Figs 6E–H). Claw of each leg smooth, with tooth ventrobasally. All leg femora with hardly visible ventral porosity. Trochanter III with posterior tubercle. Formulas of leg setation and solenidia: I (1-5-2-4-20) [1-2-2], II (1-5-2-4-16) [1-1-2], III (2-3-1-3-15) [1-1-0], IV (1-2-2-3-12) [0-1-0]; homology of setae and solenidia indicated in Table 1; seta s on tarsus I eupathidial; setae a ′′and pv ′′on tarsus IV modified (brush-like); solenidia ω 1 on tarsi I, ω 1 and ω 2 on tarsus II and σ on genu III nearly bacilliform, φ 1 on tibia I and φ on tibia IV subflagellate; other solenidia setiform or rod-like.</p><p>Remarks.  Wallworkella southensis sp. nov. is similar to  W. tetraciliata (Ermilov, 2018) (in Ermilov &amp; Rybalov 2018) from Ethiopia in having rounded rostrum and ciliate bothridial seta. However, the new species differs from the latter by smaller body size (length: 240–255 versus 298–315), the indistinct (versus well developed) costular-transcostular complex, the relative length of some prodorsal setae (ro longer then in versus ro shorter than in), the morphology of anterior margin of the notogaster (straight versus convex), and the length of the dorsal notogastral setae (short, lm shorter than 1/2 of distance lm–lp versus comparatively long, lm longer than 1/2 of distance lm– lp).</p><p>Etymology. The species name  southensis refers to the place of origin, South Ethiopia Regional State.</p><p>General remarks on  Wallworkella species</p><p>Balogh and Balogh (1990) synonymized  Wallworkella longiramosa (Woas, 1986) with  Wallworkella cervifer (Mahunka, 1983) . As noted by Ermilov (2024), perhaps, the synonymy is correct, however, according to the original descriptions (Mahunka 1983; Woas 1986) of these species, there are some differences between these species, therefore, both species are given in the key below as independent species.</p><p>Key to known species of  Wallworkella</p><p>1. Rostrum not tripartite................................................................................. 2</p><p>- Rostrum tripartite..................................................................................... 4</p><p>2. Bothridial seta pectinate (its mediodistal part without expansion or with indistinct expansion); rostrum usually concave medially; body length: 266–282......  Wallworkella minima (Ermilov, Rybalov &amp; Hundama, 2014) . Distribution: Ethiopia.</p><p>- Bothridial seta ciliate (its mediodistal part with well developed unilaterally lanceolate expansion); rostrum rounded....... 3</p><p>3. Transcostula and costula not visible; rostral seta longer than interlamellar seta; anterior margin of notogaster straight; dorsal notogastral setae short (lm shorter than 1/2 of distance lm–lp); body length: 240–255.......................................................................................  Wallworkella southensis sp. nov. Distribution: Ethiopia.</p><p>- Transcostula and costula developed; rostral seta shorter than interlamellar seta; anterior margin of notogaster convex; dorsal notogastral setae medium-sized (lm longer than 1/2 of distance lm–lp); body length: 298–315................................................  Wallworkella tetraciliata (Ermilov, 2018) (in Ermilov &amp; Rybalov 2018). Distribution: Ethiopia.</p><p>4. Bothridial seta ciliate (its mediodistal part with well developed unilaterally lanceolate expansion)..................... 5</p><p>- Bothridial seta pectinate (its mediodistal part without expansion or with indistinct expansion)........................ 6</p><p>5. Median apex of rostrum very narrow, blunt; notogastral setae la and lm similar in length; body length: 274..................................................................  Wallworkella secata (Mahunka, 1999) . Distribution: Uganda.</p><p>- Median apex of rostrum triangular; notogastral seta la distinctly shorter than lm; body length: 312...................................................................  Wallworkella parasensillus (Mahunka, 1999) . Distribution: Uganda.</p><p>6. Specific ridge-like structure (similar to an arch with triangular narrowness anteriorly) present in front of transcostula...... 7</p><p>- Specific ridge-like structure absent in front of transcostula.................................................... 8</p><p>7. Lateral prodorsal ridge short; notogastral seta lm not reaching insertion of lp; longitudinal interbothridial ridges absent; body length: 390.......................................  Wallworkella longiramosa (Woas, 1986) . Distribution: Salvador.</p><p>- Lateral prodorsal ridge long; notogastral seta lm reaching insertion of lp; two short longitudinal interbothridial ridges present; body length: 270–335...  Wallworkella cervifer (Mahunka, 1983) (see also Ermilov 2024). Distribution: Neotropical region.</p><p>8. Median apex of rostrum swollen in the middle; branches of bothridial seta barbed; dorsal notogastral setae medium-sized (lm longer than 1/2 of distance lm–lp); body length: 255–284.....................................................................  Wallworkella trimucronata (Wallwork, 1961) (see also Mahunka 1999). Distribution: Afrotropical region.</p><p>- Median apex of rostrum not swollen in the middle; branches of bothridial seta smooth; dorsal notogastral setae short (lm about 1/2 of distance lm–lp)................................................................................. 9</p><p>9. Median apex of rostrum distinctly shorter than lateral ones; anal plate with longitudinal ridge; body length: 275–302.................................  Wallworkella directa (Mahunka &amp; Mahunka-Papp, 2007) . Distribution: Afrotropical region.</p><p>- Median apex of rostrum little shorter than lateral ones; anal plate without longitudinal ridge; body length: 256–284........  Wallworkella machadoi (Balogh, 1958) (=  Arcoppia granulata Mahunka, 1986) (see also Mahunka 1999). Distribution: Afrotropical region, Ecuador.</p></div>	https://treatment.plazi.org/id/03A787FE7B3EFFFFFF7BAD07FE3D54D6	Public Domain	No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.		MagnoliaPress via Plazi	Ermilov, Sergey G.	Ermilov, Sergey G. (2025): Taxonomic contribution to knowledge of the oribatid mite genera Corynoppia, Ramusella and Wallworkella (Acari, Oribatida, Oppiidae). Zootaxa 5621 (3): 353-370, DOI: 10.11646/zootaxa.5621.3.4, URL: https://doi.org/10.11646/zootaxa.5621.3.4
