taxonID	type	description	language	source
03A58799FFB0FFB882E5A7211DFAFDA6.taxon	description	(Figs. 1 – 3, 10, 13 – 14, 17, 20, 23)	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB0FFB882E5A7211DFAFDA6.taxon	materials_examined	Type locality. Peru, Junin Department, Satipo Province, near Rio Venado village, 11 ° 11 ' 47 '' S, 74 ° 46 ' 10 '' W. Type material. Holotype, male: “ Peru, Junin dep., / Satipo prov., h ~ 1100 – 1400 m / near Rio Venado / 11 ° 11´04 ´´ S 074 ° 45´59 ´´ W / vii. 2019 leg. D. Quispe ” (in ISEA).	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB0FFB882E5A7211DFAFDA6.taxon	description	Description of holotype (male). Dorsum (Fig. 1). Body length 2.9 mm, elongate, robust, shiny, reddish brown, glabrous. Head (Fig. 10) relatively large, broad, trapezoidal, convex, in posterior half shiny, without microreticulation; in anterior half matt, with distinct microreticulation; with relatively dense, low but distinct tubercles in anterior half. Clypeus with margin bordered, distinctly sinuate anteriorly, widely rounded laterally, not notched before genae, clypeal border without macrosetae. Genae obtuse, weakly exceeding eyes, without macrosetae. Frontal suture not noticeable, without gibbosity. Punctation of clypeus double in posterior half, fine and coarse; all punctures quite dense, irregularly distributed, variable in size; in anterior half punctation simple, fine, quite dense, irregularly distributed. Epipharynx (Fig. 14) transverse, with sides rounded, anterior concavely arcuate, without corypha with two very short, thick celtes in the middle. Acanthopariae, acropariae, prophobae without chaetae. Chaetopariae with belt of dense, long, thick chaetae; adelochaetae similarly like chaetopariae but shorter; chaetopedia with dense belts of thick chaetae, as long as adelochaetae. Epitorma not observed. Tormae thin, quite long. Pronotum transverse, posteriorly approximately as wide as base of elytra, widest at posterior lobe, convex, shiny, without microreticulation, with double punctation; larger punctures variable in size, four to six times larger than smaller, dense, quite regularly distributed; smaller punctures dense, quite regularly distributed. Margins without macrosetae; posterior margin bordered by a groove covered by dense, coarse punctures; anterior margin not bordered; lateral margins bordered only near hind angles, on explanate surface. Anterior and posterior angles weakly divergent; sides near hind angles with small but distinct tooth anterior to emargination in addition to a basal tooth; margin between teeth very weakly sinuate in dorsal and lateral view (Figs. 20, 23). Scutellar shield small, triangular, with ogival sides, without punctures, shiny, without microreticulation. Elytra with complete, broad basal groove, elongate, convex, weakly widened posteriorly, shiny, without microreticulation, without macrosetae; without humeral denticles; with 10 striae and 10 intervals. Striae distinctly, densely punctate with medium sized, “ 8 ” - shaped punctures; punctures very gently indenting margins of intervals. All striae slightly shortened before apex, eighth and ninth shortened before base. Intervals shiny, evenly convex, with quite regularly distributed, simple, fine punctation along middle. Surface of the humeral callus (Fig. 17) shiny, with only a trace of microreticulation, with very sparse fine punctation. Legs. Profemora bordered anteriorly and posteriorly, very weakly shiny, with dense, very irregularly distributed, sparse punctation; mesofemora and metafemora bordered only posteriorly, shiny, without microreticulation, finely and sparsely punctate. Protibiae distinctly tridentate laterally, basal most tooth further from middle tooth, than middle tooth is from apical tooth, lateral edge not serrulate basally; dorsal surface smooth, shiny, with few fine punctures bearing long macrosetae; apical spur long, moderately broad, gently downwardly and outwardly bent, with acute apex. Mesotibiae and metatibiae without transverse carinae, fimbriate apically with row of short spinules of unequal length. Apex of metatibiae without accessory spine. Metatibiae superior apical spur slightly longer than basal metatarsomere, latter nearly as long as the next three metatarsomeres combined. Claws very small, short, thin, gently arcuate. Macropterous. Venter (Fig. 2). Metaventral plate shiny, flat, with distinct, deep, wide longitudinal line at middle; surface with fine, sparse punctures, on sides with coarse punctures. Abdominal ventrites shiny, anterior margin distinctly fluted, without microreticulation, with belt of dense, coarse, mainly longitudinal punctures. Pygidium with posterior half matt, eroded, with distinct microreticulation, additionally with distinct longitudinal, shiny median carina. Aedeagus (Figs 13). Parameres somewhat shorter than phallobase, elongate, in lateral view with small membranous area in the middle and with somewhat larger membranous process before the apex. Aedeagus similar in shape to other Odontolochini and as well a lot of members of Psammodiini and Eupariini.	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB0FFB882E5A7211DFAFDA6.taxon	etymology	Etymology. Toponymic; an adjective derived from the name Peru, where the new species was collected.	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB0FFB882E5A7211DFAFDA6.taxon	diagnosis	Diagnosis. Saprolochus peruvianus can be distinguished from all other known species by the combination of characters: body length less than 3 mm; central part of clypeus with small and large punctures, irregularly, moderately densely distributed; pronotum with lateral border reduced posteriorly, emarginate with one or two small teeth, lateral margin between teeth weakly sinuate in dorsal and lateral view; lateral pronotal border with tooth anterior to basal emargination in addition to basal tooth; lateral border of pronotum at most with indistinct row of punctures the same size as other large punctures; larger punctures of pronotum distinctly larger posteriorly than anteriorly; humeral callus of elytra shiny, with only a trace of microreticulation, with very sparse fine punctation; elytral striae not increasingly wider and deeper toward sides, elytral intervals shiny, without microreticulation; punctation of elytral intervals arranged in very indistinct rows. Also see Discussion and Table 1.	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB5FFB382E5A5F01C26F88A.taxon	description	(Figs. 4 – 6, 11, 15, 18, 21, 24)	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB5FFB382E5A5F01C26F88A.taxon	materials_examined	Type locality. Venezuela: Bolivar, 6 km S San Isidro. Type material. Holotype, female: “ Venezuela: Bolivar / 6 km S San Isidro (88 km) / 25. vi. - 11. vii. 1987 / S. & J. Peck, lowland / rainforest, ex: f. i. t., 87 – 45 ” (in CMNC).	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB5FFB382E5A5F01C26F88A.taxon	description	Description of holotype (female). Dorsum (Fig. 6). Body length 3.0 mm, elongate, robust, shiny, reddish brown, glabrous. Head (Fig. 11) relatively large, broad, trapezoidal, convex, in posterior half shiny, without microreticulation; in anterior half weakly shiny, with quite distinct microreticulation; additionally with relatively dense, low but distinct tubercles in anterior half. Clypeus with margin bordered, distinctly sinuate anteriorly, widely rounded laterally, not notched before genae, clypeal border without macrosetae. Genae obtuse, weakly exceeding eyes, without macrosetae. Frontal suture not noticeable, without gibbosity. Punctation of clypeus double in posterior half; larger punctures dense, irregularly distributed, variable in size, coarse; smaller punctures fine sparse, irregularly distributed; anterior half with punctation simple, fine, quite dense, irregularly distributed. Epipharynx (Fig. 15) transverse, with sides rounded, anterior concavely arcuate, without corypha with two very short, thick celtes in the middle. Acanthopariae, acropariae, prophobae without chaetae. Chaetopariae with belt of dense, long, thick chaetae; adelochaetae similarly like chaetopariae but shorter; chaetopedia with dense belts of thick chaetae, as long as adelochaetae. Epitorma not observed. Tormae thin, quite long. Pronotum transverse, posteriorly slightly narrower than base of elytra, widest at posterior lobe, convex, shiny, without microreticulation, with double punctation; larger punctures variable in size, four to six times larger than smaller punctures, dense, quite regularly distributed; smaller punctures dense, quite regularly distributed. All margins without macrosetae. Posterior margin bordered with a groove covered by belt of dense, coarse punctures; anterior margin not bordered; lateral margins anteriorly slightly explanate, entirely bordered with belt of dense, coarse, elongate punctures. Anterior and posterior angles weakly divergent; sides near posterior angles with distinct tooth anterior to emargination in addition to basal tooth; margin between teeth distinctly sinuate in dorsal and lateral view (Figs. 21, 24). Scutellar shield small, triangular, without punctures, shiny, with weak microreticulation. Elytra with complete, broad basal groove, elongate, convex, weakly widened posteriorly, quite shiny, with weak but distinct microreticulation, without macrosetae; without humeral denticles; with 10 striae and 10 intervals. Striae distinctly, quite densely punctate with medium sized, “ 8 ” - shaped punctures; punctures very gently indenting margins of intervals. All striae slightly shortened before apex, eighth and ninth shortened before base. Intervals shiny, evenly convex, with fine and simple punctation regularly distributed along the middle. Surface of the humeral callus (Fig. 18) matt, with distinct microreticulation, with sparse fine punctation, and sparsely distributed shiny tubercles. Legs. Profemora bordered anteriorly and posteriorly, very weakly shiny, with dense, very irregularly distributed, sparse punctation; mesofemora and metafemora bordered only posteriorly, shiny, without microreticulation, finely and sparsely punctate. Protibiae distinctly tridentate laterally, basal tooth further from middle tooth than middle tooth is from apical tooth, edge not serrulate basally; dorsal surface smooth, shiny, with few fine punctures bearing quite long macrosetae; apical spur long, moderately broad, gently downwardly and outwardly bent, with acute apex. Mesotibiae and metatibiae without transverse carinae, fimbriate apically with row of short spinules of unequal length. Apex of metatibiae without accessory spine. Metatibiae superior apical spur sub-equal in length to basal metatarsomere, latter nearly as long as two of next metatarsomeres combined. Claws very small, short, thin, gently arcuate. Macropterous. Venter (Fig. 5). Metaventral plate shiny, flat, with distinct, deep, wide longitudinal medial line; surface with fine, sparse punctures, on sides with coarse punctures. Abdominal ventrites shiny, anterior margin distinctly fluted, without microreticulation, with belt of dense, coarse, mainly rounded punctures. Pygidium with posterior half matt, eroded, with very distinct microreticulation, additionally with distinct longitudinal, shiny median carina.	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB5FFB382E5A5F01C26F88A.taxon	etymology	Etymology. Toponymic; an adjective derived from the name Venezuela, where the new species was collected.	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB5FFB382E5A5F01C26F88A.taxon	diagnosis	Diagnosis. Saprolochus venezuelensis can be distinguished from all other known species by the combinations of characters: body length 3.0 mm; pronotum with lateral margin reduced posteriorly, emarginate with one or two small teeth; lateral border of pronotum with distinct row of punctures much larger than others; lateral pronotal margin with tooth anterior to basal emargination in addition to basal tooth; larger punctures of pronotum slightly larger posteriorly than anteriorly; humeral callus of elytra matt, rough with distinct microreticulation, with very sparse fine punctation, and sparsely distributed shiny tubercles; elytral striae increasingly wider and deeper toward sides, elytral intervals quite shiny, but with distinct microreticulation. Also see Discussion and Table 1.	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB8FFB282E5A5B81B0BFD42.taxon	description	(Figs. 8 – 9, 12, 16, 19, 22, 25 – 26)	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB8FFB282E5A5B81B0BFD42.taxon	materials_examined	Material examined. Female: Colombia: Leticia, Amazonas, 700 m., 9. vii. 1970, leg. H. & A. Howden (in ISEA). Supplementary description. Body length of studied specimen: 2.9 mm. Posterior angles of pronotum, between anterior and basal tooth, with very weak lobe in dorsal view (Fig. 22), weakly sinuate in lateral view (Fig. 25). Surface of elytral humeral callus (Fig. 19) shiny, with weak microreticulation, with very dense, fine, irregularly distributed, punctures frequently merging.	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
03A58799FFB8FFB282E5A5B81B0BFD42.taxon	discussion	Comment. The supplementary description includes the shape of unique features useful when distinguishing S. tambopatae from newly described species. The specimen studied is labelled “ paratype ” (Fig. 26). However, it is not a true paratype - for reasons unknown to the authors, it was not included in the description. Thus, this is the first report of this species from Colombia.	en	Minkina, Łukasz, Skelley, Paul, Gama, Emanuel R. R. (2025): Two new species of the genus Saprolochus Stebnicka & Galante, 2007 (Coleoptera: Scarabaeidae: Aphodiinae: Odontolochini). Zootaxa 5689 (1): 165-175, DOI: 10.11646/zootaxa.5689.1.7, URL: https://doi.org/10.11646/zootaxa.5689.1.7
