taxonID	type	description	language	source
03ADA257FFBBFFEF8EAAF9B1D30CFA75.taxon	discussion	Remarks. The morphological characters of males indicate that Anommatium is closely related to Pantolyta. The aberrant appearance of Anommatium female is associated with geophilous life mode and has a distinct adaptive value (Macek, 1989 b). Most features of Anommatium female mentioned by Macek (1989 b) are not unique and independently evolved in other lineages of diapriid wasps, except for two ones, a reduced number of antennal segments (13 or 14) and palpal formula (maxillary palpus one- segmented; labial palpus strongly reduced, represented by a small tubercle). The mentioned two differences are found to be distinctive, supporting a separate taxonomic status of the genera Anommatium and Pantolyta. The monotypic genus Erasikea Szabó, 1961 was based on a single female specimen. In the generic diagnosis, Szabó (1961) mentioned that this genus shares some diagnostic characters (i. e. rounded head, missing ocelli, 14 - segmented antennae, mesosoma narrower than head, absent wings and tegulae) with Anommatium ashmeadi. Other morphological characters of Erasikea list- ed in the original description are (1) the mesonotum with notauli, (2) posterior two-thirds of the propodeum crooked, hollowed out like a spoon, deeply cut out in the middle, with a small tooth on each side, and (3) the petiole flattened dorsoventrally. We have examined the holotype of Erasikea nutrix and found that actually the notauli on the mesonotum are missing, and the propodeum and petiole are malformed, causing the misinterpretation of characters 2 and 3. Thus, A. ashmeadi is found to be conspecific with E. nutrix, syn. nov., and the genus Erasikea, syn. nov., is a junior synonym of Anommatium.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFBBFFEE8D0DF9FAD6C3F805.taxon	description	(Fig. 3)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFBBFFEE8D0DF9FAD6C3F805.taxon	materials_examined	Holotype of Erasikea nutrix. Female; Greece, “ Holotypus / Erasikea / nutrix / Szabo 1961 ”, “ Antr. Jovis Mt. Ida. 906 ”, “ Creta Biro ”, [Crete I., Mt. Ida, 1906, L. Biro leg.] (HNMH). Other material examined. Moldova: Kishinev [ChiȘinĂu], 11. VI. 1967, V. Tobias leg., 2 males; Kotovsk [Hincesti], 4. VI. 1967, V. Tobias leg., 1 male; StrĂȘeni, 21. VII. 1961, V. Talitskiy leg., 1 male. Ukraine, Ivano-Frankivsk Prov., Nadvirna Distr., vicinity of Virikhta, 21. VII. 1989, A. Kotenko leg., 1 male. Russia: Kaliningrad Prov.: vicinity of Svetlogorsk, 1 – 12. VIII. 1999, V. Kolyada leg., 1 female (ZMUM); Curonian Spit National Park, 55 ° 09 ′ 17 ″ N 20 ° 51 ′ 27 ″ E, Rybachiy Vill., 24. VII – 4. VIII. 2006, V. Kolyada, 4 males (ZMUM); Republic of Karelia, west coast of Ladoga Lake, Koyonsary I., 28 – 29. VII. 2005, K. Tomkovich leg., 1 male; Krasnodar Terr.: NW of Sochi, Lazarevskoye, 20 – 24. V. 1973, V. Tobias leg.; same locality, 14. VI. 1984, Udalova leg., 2 males; Yaroslavl’ Prov., Berditsyno [date unknown], N. Kokujev collection, 4 males; Moscow: Bitsa Park, 4. VIII. 1993 and 10. VII. 1994, V. Kolyada leg., 3 females, 22 males (ZMUM); Krylatskoye, Rublevskiy forest, 1 – 15. VI. 2004 and VII. 2004, V. Kolyada leg., 10 males, 4 females (ZMUM); Moscow Prov.: vicinity of Pushkino, 5. VIII. 1993, V. Kolyada leg., 7 males (ZMUM); Malakhovka, 26. VII – 1. VIII. 1994 and 23. VII. 1995, M. Mostovski, 17 males (ZMUM); Vladimir Prov. [locality unknown], mixed forest, 26. VII. 1977 [collector unknown], 1 male; Kirov Prov., Bol’sheromanovo Vill., 2 – 17. VIII. 1994, V. Kolyada leg., 13 males, 2 females (ZMUM); Chuvash Republic, Asanovo Vill., 4. VIII. 1966, M. Kozlov leg., 1 male. Abkhazia: Khipsta River, 43 ° 18 ′ 4.2 ″ N 40 ° 41 ′ 31.3 ″ E, 29 – 30. VII. 2015, V. Chemyreva leg., 1 male; Lashipse River, 43 ° 29 ′ 29.6 ″ N 40 ° 39 ′ 41.9 ″ E, 7 – 8. VIII. 2015, V. Chemyreva leg., 5 males; Bzipi River, 43 ° 21 ′ 50.1 ″ N 40 ° 29 ′ 44.8 ″ E, 11 – 14. VIII. 2015, V. Chemyreva leg., 1 male. Variation. Female. Body length 1.7 – 2.3 mm. Antennae with 13 or 14 segments. Median propodeal keel complete and distinct to completely reduced. Male. Body length 1.5 – 2.3 mm. Brown to dark brown; dark specimens with propodeum paler than remainder of body; clypeus coloured as adjacent parts of head or paler. Marginal vein 1.2 – 2.0 times as long as distance from it to basal vein; petiole 0.8 – 1.1 times as wide as long. Distribution. * Abkhazia, Albania, Czech Republic, Germany, Hungary, Italy, * Moldova, Poland, * Russia (European part), Slovakia, Sweden, * Ukraine, former Yugoslavia. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFBAFFEE8D1CFF2ED2C3FD1A.taxon	discussion	Remarks. We examined a syntype of B. radiatula and found that this species actually belongs to the genus Pantoclis. Hence, we propose here a new combination Pantoclis radiatula, comb. nov.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFBAFFE98D1CFD14D5FFFAE2.taxon	description	Acropiesta Förster, 1856: 129, 131, 135; syn. nov. (Type species: Acropiesta flavicauda Ashmead, 1893).	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFBAFFE98D1CFD14D5FFFAE2.taxon	discussion	Remarks. Macek (1993, 1998) revised all known European species of Pantolyta and Acropiesta and discussed the morphological features for distinguishing these two genera. According to Macek, Acropiesta species are characterised by the following characters: (1) mandibles asymmetrical, not prominent (in rest), (2) mouth conus indistinct, (3) labrum almost hidden, stripe-like to level-semicircular, (4) apodemae of aedeagus slightly inflexed anteriorly and not sclerotised, and (5) body length more than 2.5 mm. The genus Pantolyta possesses (1) mandibles almost symmetrical, prominent (in rest), (2) mouth conus pronounced, (3) labrum prominent, triangular, (4) apodemae of aedeagus strongly inflexed anteriorly and sclerotised, and (5) body length less than 2.5 mm. Actually, species of the former genus Acropiesta have asymmetrical mandibles (the left one bidentate, the right one tridentate), but similar mandibles are known also in the genus Pantolyta (Fig. 1 A). This indicates a considerable variation in the mandibular structures, ranging from asymmetrical to symmetrical state. Pantolyta elegans Chemyreva et Kolyada, 2019 recently described from Japan and the Russian Far East and P. melniki sp. nov. have an intermediate shape of the mouth conus and semicircular labrum (Fig. 2 B, C). At the same time, all other features mentioned above (characters 1, 4 and 5) testify that P. elegans should be classified as a species of Pantolyta sensu stricto, and P. melniki sp. nov., as a species of Acropiesta. The differences in the male genitalia, i. e. in the degree of sclerotisation and the shape of the apodemae, are also not significant enough to separate these two genera. Moreover, the variation of the latter character is not enough studied. Some undescribed species from Japan possess all characters listed above for the former genus Acropiesta but have the body length about 2.0 mm (Chemyreva, pers. obs.). The body length is found to be more variable in species from the East Palaearctic than in European ones. Thus, we conclude that Acropiesta Förster, 1856, syn. nov., is a junior synonym of Pantolyta Förster, 1856, and the following species of the former genus Acropiesta are transferred here to Pantolyta: P. flavicauda (Ashmead, 1893), comb. nov.; P. flaviventris (Thomson, 1858), comb. nov.; P. flexinervis (Macek, 1998), comb. nov.; P. macrocera (Thomson, 1858), comb. nov.; P. micans (Macek, 1998), comb. nov.; P. nigrocincta (Kieffer, 1909), comb. nov.; P. nitida (Thomson, 1858), comb. nov.; P. pseudosciarivora (Macek, 1998), comb. nov.; P. pulchella (Whittaker, 1930), comb. nov.; P. radialis (Hellén, 1964), comb. nov.; P. rufiventris (Kieffer, 1909), comb. nov.; P. sciarivora (Kieffer, 1907), comb. nov.; P. semirufa (Fouts, 1924), comb. nov.; P. seticornis (Kieffer, 1910), comb. nov.; and P. subaptera (Ashmead, 1893), comb. nov.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFBDFFEA8EAAFA4CD6C3F9A5.taxon	description	(Figs 1 B, 4 B, 6 C, 8 A, 10 D, 11 F, 13 J, 14 A) Holotype. Female; Russia, Primorskiy Terr., Lazovskiy Nature Reserve, 43 ° 15 ′ 17 ″ N 134 ° 07 ′ 59 ″ E, yellow pan trap, 5 – 25. VII. 2005, K. Makarov leg. (ZIN). Paratypes (all in ZIN). Russia: Samara Prov.: Stavropol'skiy Distr., Zhiguli Nature Reserve, near Bakhilova Polyana, 3. VII. 2011, Chemyreva leg., 1 female; Kinel’ Distr., near Domashka, forest along Samara River, 29 – 31. V. 2019, 16 – 18. VI. 2019 and 2 – 4. VII. 2019, Chemyreva leg., 1 male, 4 females; Primorskiy Terr.: same locality and data as for holotype, 2 females; same locality as for holotype, VII. 2007, K. Makarov leg., 3 females; same locality as for holotype, 9 – 17. VII. 2008, A. Khalaim leg., 1 male; Sakhalin Prov., Kunashir I., vicinity of Sernovodsk, 15 – 16. VII. 1973, D. Kasparyan leg., 1 male. Azerbaijan, Lenkoran’, 3 – 7. V. 1971, V. Tobias leg., 1 male. Description. Female (holotype). Body length 3.4 mm; fore wing length 2.3 mm; antenna length 1.9 mm. Coloration. Head, A 13 – A 15, mesosoma and petiole dark brown; T 2 – T 6 and S 2 – S 5 brown; A 1 – A 12, mandibles, veins and metasoma (excluding petiole) pale brown; palpi, legs, T 7 – T 8 and S 6 yellowish brown. Head not nasiform in dorsal view, 0.77 times as long as wide, wider than mesosoma, smooth, with few scattered long setae. Temples sharply receding behind eye in dorsal view. Occipital flange narrow, sculptured. Head in lateral view 1.26 times as high as long. Malar space 0.71 times as long as largest diameter of eye. Eye weakly pubescent. Antennal shelf in frontal view with fine coriaceous sculpture below toruli. Face smooth, with long scattered setae. Head in frontal view with mouth conus indistinct; genae converging towards mouthparts in frontal view. Clypeus transverse, 0.67 times as high as wide. Tentorial pits small. Labrum small, semicircular, hardly discernible. Mandibles asymmetrical (Fig. 1 B). Antenna slender, weakly broadened apically (Fig. 4 B). Ratios of length to width of A 1 – A 15 in dorsal view: A 1 (70: 12); A 2 (20: 10); A 3 (24: 9); A 4 (18: 9); A 5 (16: 10); A 6 (16: 10); A 7 (16: 10); A 8 (15: 10); A 9 (15: 10); A 10 (15: 12); A 11 (15: 12); A 12 (14: 12); A 13 (14: 13); A 14 (15: 14); A 15 (34: 16). Mesosoma slightly compressed, 1.1 times as high as wide, in dorsal view 1.8 times as long as wide. Sides of pronotum bare, pronotal collar with scattered long setae; epomia strongly prominent; pronotal pit bare, with epomia inside. Mesoscutum 1.19 times as long as wide, convex, with few scattered semierect setae. Notauli deep and complete, not sculptured. Anterior scutellar pit deep and rounded. Scutellum large, widened posteriorly. Axillar depression scarcely pubescent, with verriculate tubercle anteriorly (Fig. 11 F). Mesopleuron scarcely pubescent ventrally and anteriorly, with subalar bridge posterodorsally and epicnemial carina anteroventrally; epicnemial pit deep and pubescent; mesopleural pit completely absent (Fig. 10 D). Metascutellum narrow, pubescent; dorsellum distinct, with three short longitudinal keels. Propodeum transverse, 0.68 times as long as wide, bare dorsally and pubescent laterally, medi- an propodeal keel bifurcate from base; upper and lower posterior propodeal projections forming distinct teeth (Figs 10 D and 11 F). Sides of propodeum entirely pubescent, with three longitudinal keels (Fig. 10 D). Legs slender. Fore wing with closed radial cell (Fig. 14 A). Distance from marginal vein to basal vein / marginal vein length / radial cell length as 15 / 16 / 16. Petiole of metasoma cylindrical, as long as wide, with coarse rugose sculpture, longitudinal keels and several rows of long erect setae laterally and ventrally; petiole ventrally with several bunches of setae; posterior margin of petiole weakly arcuate (Fig. 11 F). T 2 anteriorly with short striation, median groove the longest; T 3 – T 6 narrow, with few setae; T 7 and T 8 long, strongly compressed laterally. S 2 with short grooves at base and with numerous erect scattered setae; S 3 – S 5 narrow, with scattered setae; S 6 distinctly elongate and compressed, with a few scattered setae. Male. Body length 2.9 – 3.2 mm. Similar to female except following characters: antenna filiform, slightly longer than body; antennomeres cylindrical, with uniform short pubescence; A 1 slender, 0.55 times as long as head width; A 3 deeply emarginate, with keel extending to one-third of A 3 length (Fig. 6 C); ratios of length to width of A 1 – A 5 in lateral view: A 1 (50: 12); A 2 (14: 11); A 3 (50: 12); A 4 (44: 11); A 5 (44: 11); radial cell longer than marginal vein; petiole elongate, 1.57 as long as wide (Fig. 13 J); T 7 transverse, not compressed; S 2 – S 7 with numerous semierect scattered setae; T 8 and S 8 subtriangular, rounded apically. Variation. Female. Body length 3.2 – 3.7 mm. Median propodeal keel bifurcate from base to posterior 0.65; branches of median propodeal keel adjacent to each other or distinctly separated. Distance from marginal vein to basal vein, length of marginal vein and length of radial cell equal to each other, or sometimes marginal vein longer. Petiole as long as wide or slightly elongate. Comparison. Pantolyta dichromia sp. nov. can be distinguished from all known species of Pantolyta by the combination of the following characters: the head is not nasiform (Fig. 8 A) and has no mouth conus, the genae converge toward mouthparts (Fig. 1 B); the female antennae are slender, weakly broadened apically, A 3 – A 10 are yellowish, with two or three apical segments dark brown (Fig. 4 B); V. G. Chemyreva & V. A. Kolyada. Taxonomy of Acropiesta, Anommatium, Erasikea and Pantolyta A 3 is deeply emarginated, with a keel extending to one-third of A 3 length (Fig. 6 C); pronotal pit is bare, with short epomia inside; the mesopleuron possesses subalar and epicnemial bridges, the mesopleural pit is completely absent (Fig. 10 D); the median keel is bifurcated from the base; upper and lower posterior propodeal projections form large teeth (Figs 10 D and 11 F); the fore wing has a closed radial cell. Etymology. The name derives from the Ancient Greek adjective “ δί- χρωµος ” (bicolor), in reference to the bicolourous antennae of female. Distribution. Azerbaijan, Russia (European part, Primorskiy Territory, Kunashir Island). Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFBEFFE58EA5F989D4C6FA39.taxon	description	(Figs 5 A, 6 A, 11 H, 15 A – C, 16 B)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFBEFFE58EA5F989D4C6FA39.taxon	materials_examined	Material examined. Ukraine: Zakarpattia Prov., Carpathian Biosphere Reserve, 7 – 8. VIII. 1989, A. Kotenko leg., 3 males; Ternopil Prov., Berezhany, 31. V. 1952, Mesnyaev leg., 2 males; Kyiv, Holosiivskyi Forest, 27. VI. 1993, A. Kotenko leg., 1 female; Kyiv Prov., Irpin, 27. V. 1984, A. Kotenko leg., 1 male. Russia: Kaliningrad Prov., Curonian Spit National Park, 55 ° 09 ′ 16.83 ″ N 20 ° 51 ′ 27.03 ″ E, Rybachiy Vill. ,, 24. VII – 4. VIII. 2006, V. Kolyada leg., 1 female (ZMUM); Leningrad Prov., Semrino, 22. VII. 1972, D. Kasparyan leg., 1 male; Moscow: Krylatskoe, Rublevskiy forest, VI. 2004, V. Kolyada leg., 3 females (ZMUM); same locality, VII. 2004, V. Kolyada leg., 1 female (ZMUM); Bitsa Park, 23. VI. 1993 and 1 – 10. VII. 1994, V. Kolyada leg., 2 females (ZMUM); Moscow Prov.: Malinki Vill., 18. V. 1973, E. Antonova leg., 2 females (ZMUM); Pavlovskaya Sloboda Vill., 3. VII. 1993, V. Kolyada leg., 2 males (ZMUM); Karachay-Cherkess Republic, Teberda, 10 – 15. VII. 1976, D. Kasparyan leg., 1 female; Chechen Repiblic, Itum- Kale Vill., 8. VI. 1972, D. Kasparyan leg., 1 female; Komi Republic, 40 km W of Labytnangi, 17. VIII. 1972, D. Kasparyan leg., 1 male; Tyumen Prov., Krasnosel’kup, 11 – 13. VIII. 1992, D. Kasparyan leg., 1 female; Krasnoyarsk Terr., Eniseysk Distr., Nazimovo, 11 – 13. VII. 1988, D. Kasparyan leg., 1 female; Sakhalin Prov.: Sakhalin I., vicinity of Novoaleksandrovka, V. G. Chemyreva & V. A. Kolyada. Taxonomy of Acropiesta, Anommatium, Erasikea and Pantolyta 5. VII and 7. X. 1973, D. Kasparyan leg., 1 male, 1 female; Noyba River, 10 km from Bykovo, 10 – 20. VIII. 1991, V. Zherikhin leg., 1 male (ZMUM); Primorskiy Terr., Shkotovskiy Distr., Peyshula, 25. VII. 1972, V. Kuslitzky leg. 1 male; 10 km SE of Ussuriysk, 12 – 19. VII. 2001, S. Belokobylskij leg., 1 female; Lazovskiy Nature Reserve, 9 – 17. VII. 2008, A. Khalaim leg., 1 female. Crimea: Crimean Nature Reserve, 21. VI. 1976 and 20. VI. 1978, A. Kotenko leg., 3 males; same locality, 28. VI. 1976, S. Kononova leg., 3 males. Georgia, Arsiani Mts, Goderdzis Pass, 3. VII. 1967, D. Kasparyan leg., 1 male. Variation. Female with antenna yellowish brown to dark brown; A 3 1.0 – 1.2 times as long as A 15; petiole weakly transverse to quadrate. Male with antenna shorter to longer than body; metasoma posterior to petiole 0.92 – 1.1 times as long as head, mesosoma and petiole combined; petiole 1.5 – 2.15 times as long as wide. Both sexes with radial cell 1.75 – 2.45 times as long as marginal vein; marginal vein 0.85 – 1.1 times as long as distance from it to basal vein. Distribution. Western, northern and eastern Europe, * Georgia, Russia (European part, * Siberia, * Far East), * Ukraine. Hosts. Trichosia caudata (Walker, 1848) (Diptera: Sciaridae) (Tuomikoski, 1957; Hellén, 1964) and Sciara ligniperda (Brischke, 1891) (Diptera: Sciaridae) living in rotten alder wood (Brischke, 1891).	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB1FFE48D0DFA35D6C3F912.taxon	description	(Figs 5 B, 6 G, 9 A, 11 D, 13 A, 14 E)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB1FFE48D0DFA35D6C3F912.taxon	materials_examined	Material examined. Russia, Sakhalin Prov.: Sakhalin I., Cape Ostryy, 5 – 21. VII. 2009, A. Khalaim leg., 1 female; Kunashir I., Golovnin Caldera, 25. VII. 2011, E. Tselikh leg., 1 female. Variation. Female antenna with three to six segments between A 6 – A 12 weakly thickened. Marginal vein in both sexes 1.25 – 1.50 times as long as distance from it to basal vein; radial cell 1.47 – 1.74 times as long as marginal vein; petiole 1.0 – 1.3 times as long as wide. Distribution. Czech Republic, Slovakia, * Russia (Far East). Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB0FFE78EA5F96BD6D3F805.taxon	description	(Figs 4 H, 7 F, 11 G, 14 D, 15 E – F)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB0FFE78EA5F96BD6D3F805.taxon	materials_examined	Material examined. Moldova: Cotovscoe Vill., 4. VI. 1967, V. Tobias leg., 5 males; Kishinev [ChiȘinĂu], 11. VI. 1967, V. Tobias leg., 1 male. Ukraine: Kyiv, Holosiivskyi forest, 27. VI. 1993, A. Kotenko leg., 2 males; Kyiv Prov., vicinity of Fastiv, 12. VII. 1980, A. Kotenko leg., 2 males. Russia: Kaliningrad Prov., Curonian Spit National Park, 55 ° 09 ′ 16.83 ″ N 20 ° 51 ′ 27.03 ″ E, V. G. Chemyreva & V. A. Kolyada. Taxonomy of Acropiesta, Anommatium, Erasikea and Pantolyta Fig. 7. Pantolyta, female. Head, frontal view. A, P. radialis; B, P. micans; C, P. nigrocincta; D, P. simplicior sp. nov.; E, P. sciarivora; F, P. macrocera. Rybachiy Vill. ,, 24. VII – 4. VIII. 2006, V. Kolyada leg., 7 males, 2 females (ZMUM). Armenia, Syunik Prov., Kapan Distr., 1 – 10. VII. 1971, V. Kuslitzky leg., 1 male. Azerbaijan, Qusar, 27. V. 1972, D. Kasparyan leg., 1 male. Variation. Male. Head, mesosoma and petiole reddish brown to dark brown; marginal vein 1.0 – 1.45 times as long as distance from it to basal vein; radial cell as long as marginal vein (rarely) or longer, at most 1.57 times as long as marginal vein; petiole 1.9 – 2.5 times as long as wide. Distribution. * Armenia, * Azerbaijan, Austria, Czech Republic, Finland, Germany, Hungary, Ireland, * Moldova, * Russia (European part), Slovakia, Sweden, * Ukraine, United Kingdom. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB3FFE08D0DFA8AD369F927.taxon	description	(Figs 2 C, 4 C, E, 6 D, 8 B, 9 D, 10 F, 11 B, 13 I, 14 C) Holotype. Female; Russia, Sakhalin Prov., Kunashir I., vicinity of Grozovoe, 8 – 15. VIII. 2008, I. Melnik leg. (ZIN). Paratype. Republic of Korea, Gangwon Prov., Odaesan National Park, Dongsan-Li, near Woljeongsa, fir forest, Malaise trap, 19. VII – 18. VIII. 2003 [collector unknown], 1 male (ZIN). Description. Female. Body length 3.5 mm; fore wing length 2.7 mm; antenna length 2.1 mm. Coloration. Head, mesosoma and petiole dark brown; antenna, mandibles, venation and metasoma (except petiole) pale brown; legs and palpi yellowish brown. V. G. Chemyreva & V. A. Kolyada. Taxonomy of Acropiesta, Anommatium, Erasikea and Pantolyta Fig. 8. Pantolyta and Anommatium, female (A – E) and male (F). Head, dorsal view. A, P. dichromia sp. nov.; B, P. melniki sp. nov.; C, P. sciarivora; D, P. radialis; E, P. nigrocincta; F, A. ashmeadi. Head not nasiform, in dorsal view 1.25 times as wide as long (Fig. 8 B), wider than mesosoma, smooth, with few scattered long setae. Temples just behind eyes parallel, broadly receding posteriorly. Occipital flange narrow, foveolate (Fig. 8 B). Head in lateral view 1.21 times as high as long. Malar space 0.90 times as long as largest diameter of eye. Eye pubescent. Antennal shelf in frontal view with fine coriaceous sculpture below toruli. Face smooth, with long scattered setae. Head in frontal view without mouth conus; genae slightly convex and converging toward mouthparts. Clypeus transverse, 0.67 times as high as wide. Tentorial pits large and transverse. Labrum small, semicircular, hardly discernible (Fig. 2 C). Mandibles simple. Antenna slender, weakly broadened apically (Fig. 4 C, E). Ratios of length to width of A 1 – A 15 in dorsal view: A 1 (40: 8); A 2 (12: 7); A 3 (18: 7); A 4 (12: 7); A 5 (11: 7); A 6 (11: 7); A 7 (10: 7); A 8 (10: 7); A 9 (10: 7); A 10 (10: 8); A 11 (9: 9); A 12 (9: 9); A 13 (9: 9); A 14 (9: 10); A 15 (16: 11). Mesosoma slightly compressed, 1.08 times as high as wide, in dorsal view 1.83 times as long as wide. Sides of pronotum bare, pronotal collar with scattered long setae; epomia strongly prominent; pronotal pit bare, with epomia inside (Fig. 9 D). Mesoscutum slightly elongate, 1.13 times as long as wide, convex, with few scattered semierect setae. Notauli deep and complete, sculptured. Anterior scutellar pit deep and rounded. Scutellum large, widened posteriorly. Axillar depression scarcely pubescent, with two verriculate tubercles (Fig. 11 B). Mesopleuron scarcely pubescent ventrally and anterodorsally, with subalar bridge posterodorsally and epicnemial bridge anteroventrally; epicnemial pit deep and pubescent; mesopleural pit developed as shallow depression (Fig. 10 F). V. G. Chemyreva & V. A. Kolyada. Taxonomy of Acropiesta, Anommatium, Erasikea and Pantolyta Fig. 9. Pantolyta, female. Pronotum, lateral view. A, P. flexinervis; B, P. sciarivora; C, P. seticornis; D, P. melniki sp. nov.; E, F, P. radialis. Metascutellum narrow, pubescent; dorsellum distinct, with three short longitudinal keels. Propodeum transverse, 0.38 times as long as wide, entirely pubescent, with simple median keel; upper and lower posterior propodeal projections forming large teeth (Figs 11 B and 13 I). Sides of propodeum entirely pubescent, with three longitudinal keels (Fig. 10 F). Legs slender. Fore wing with closed radial cell (Fig. 14 C); distance from marginal vein to basal vein / marginal vein length / radial cell length as 12 / 19 / 18. Petiole of metasoma transverse 0.75 times as long as wide, with coarse rugose sculpture and few scattered setae laterally (Fig. 11 B); petiole ventrally with several bunches of setae. T 2 anteriorly with short striation, median groove the longest; T 3 – T 6 narrow, scarcely pubescent; T 7 and T 8 long, strongly compressed laterally. S 2 with short grooves at base and with numerous erect scattered setae; S 3 – S 5 narrow, with scattered setae; S 6 distinctly elongate and compressed, with few scattered setae. Male. Body length 2.9 mm. Similar to female except following characters: antenna filiform; antennomeres cylindrical, with uniform short pubescence; A 1 slender, 5.0 times as long as wide and 0.64 times as long as head width; A 3 deeply emarginate, with keel extending to 0.6 of A 3 length (Fig. 6 D); ratios of length to width of A 1 – A 5 in lateral view: A 1 (40: 8); A 2 (14: 8); A 3 (29: 9); A 4 (20: 7); A 5 (18: 7); petiole elongate, 1.38 times as long as wide (Fig. 13 I); T 7 transverse, not compressed; S 2 – S 7 with numerous semierect scattered setae; T 8 and S 8 subtriangular, rounded apically. Fig. 10. Pantolyta, female (A – E) and male (F). Mesosoma and anterior part of metasoma, lateral view. A, P. micans; B, P. nigrocincta; C, P. rufiventris; D, P. dichromia sp. nov.; E, P. simplicior sp. nov.; F, P. melniki sp. nov. Green arrow – upper posterior propodeal projection; white arrow – lower posterior propodeal projection; red arrows – median propodeal keel. Comparison. Pantolyta melniki sp. nov. can be distinguished from all known species of Pantolyta by the combination of the following characters: the head is not nasiform (Fig. 8 B) and has no mouth conus (Fig. 2 C); the genae are weakly convex in frontal view (Fig. 2 C); the female antenna is slender, weakly broadened apically, A 15 is wider than A 14 in dorsal view (Fig. 4 C, E); A 3 in male has a deep emargination and a keel reaching mid-length of the segment (Fig. 6 D); the pronotal pit is bare, with a distinct vertical keel inside (Fig. 9 D); the mesopleuron possesses subalar and epicnemial bridges and the mesopleural pit developed as a shallow depression (Fig. 10 F); the propodeum has a simple median keel (Fig. 11 B); upper and lower posterior propodeal projections form large teeth (Figs 11 B and 13 I); the fore wing has a closed radial cell, marginal vein longer than the distance from marginal vein to basal vein and almost as long as radial cell; the petiole is transverse. Etymology. This new species is named in honour of the entomologist Igor Melnik (Moscow), who collected the holotype of this species. Distribution. Russia (Kunashir Island), Republic of Korea. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB4FFE38D1CF917D6D3F805.taxon	description	(Figs 5 C, 6 K, 7 B, 10 A, 11 E, 13 B)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB4FFE38D1CF917D6D3F805.taxon	materials_examined	Material examined. Russia: Moscow Prov., Serpukhov Distr., Prioksko-Terrasnyy National Reserve, V. G. Chemyreva & V. A. Kolyada. Taxonomy of Acropiesta, Anommatium, Erasikea and Pantolyta Fig. 11. Pantolyta, female. Mesosoma and base of metasoma, dorsal view. A, P. seticornis; B, P. melniki sp. nov.; C, P. nitida; D, P. flexinervis; E, P. micans; F, P. dichromia sp. nov.; G, P. macrocera; H, P. flaviventris. 2. VI. 2018, I. Panina leg., 1 male; Krasnodar Terr., NW of Sochi, Lazarevskoye, 20 – 24. V. 1973, V. Tobias leg., 1 male, 1 female; Sverdlovsk Prov., Achit Distr., VII. 1972, Sorokina leg., 1 male. Abkhazia, Pitsunda, 6 – 11. V. 1983, R. Dbar leg., 1 male. Variation. Head, mesosoma and petiole yellowish brown to dark brown. Radial cell 0.65 – 1.3 times as long as marginal vein; marginal vein 1.1 – 1.4 times as long as distance from it to basal vein. Distribution. Austria, Czech Republic, Germany, Netherlands, * Russia (European part), Slovakia, Sweden. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB7FFE28D0DFA7CD6C0F9A9.taxon	description	(Figs 4 D, F, 6 I, 7 C, 8 E, 10 B, 12 F, 13 C, 14 B)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB7FFE28D0DFA7CD6C0F9A9.taxon	materials_examined	Material examined. Russia: Novgorod Prov., Pestovo Distr., Tychkino, 8 – 10. VII. 1990, V. Tobias leg., 1 female; Moscow Prov., Serpukhov Distr., Prioksko-Terrasnyy National Reserve, 10. VI. 2016, N. Smirnova leg., 1 male; Krasnodar Terr., Gelendzhik, 10. VI. 1954, L. Arnoldi leg., 1 female; Primorskiy Terr., Lazovskiy Nature Reserve, 9 – 17. VII. 2008, A. Khalaim leg., 1 female. V. G. Chemyreva & V. A. Kolyada. Taxonomy of Acropiesta, Anommatium, Erasikea and Pantolyta Fig. 12. Pantolyta, female. Mesosoma and base of metasoma, dorsal view; female. A, P. radialis; B, P. pseudosciarivora; C, P. rufiventris; D, P. simplicior sp. nov.; E, P. sciarivora; F, P. nigrocincta. Variation. Female with genae in frontal view weakly to distinctly convex. Male with medial furrow at base of T 2 strongly reduced to distinct. Both sexes with radial cell 0.95 – 1.3 times as long as marginal vein; marginal vein 0.95 – 1.25 times as long as distance from it to basal vein. Distribution. Austria, Czech Republic, Germany, Hungary, Poland, * Russia (European part, Far East), Sweden. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB6FFE28EA5F986D369FAFF.taxon	description	(Figs 5 D, 6 B, 11 C, 13 D, 14 G)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB6FFE28EA5F986D369FAFF.taxon	materials_examined	Material examined. Russia: Novgorod Prov., Pestovo Distr., Tychkino, 4 – 17. VI. 1998, V. Tobias leg., 1 male; Moscow, Krylatskoe, Rublevskiy forest, 1 – 15. VI. 2004, V. Kolyada leg., 2 males, 1 female. Variation. Head, mesosoma and petiole yellowish brown to dark brown; marginal vein 0.65 – 0.90 times as long as distance from it to basal vein. Distribution. Czech Republic, Denmark, Germany, Slovakia, * Russia (European part), Sweden. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB6FFFD8D1CFA71D2EDF89D.taxon	description	(Figs 5 E, 6 J, 12 B, 13 F, 14 H)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFB6FFFD8D1CFA71D2EDF89D.taxon	materials_examined	Material examined. Crimea, 3 km S of Pereval’noe Vill., 29. V. 1995, V. Kolyada leg., 1 male. Variation. Male. Largest diameter of eye 1.30 – 1.55 times as long as malar length. Distribution. Czech Republic, Germany, Ireland, * Russia (European part), Sweden, United Kingdom. Hosts. Unknown. Fig. 13. Pantolyta, male. Petiole, dorsal view. A, P. flexinervis; B, P. micans; C, P. nigrocincta; D, P. nitida; E, P. rufiventris; F, P. pseudosciarivora; G, P. seticornis; H, P. radialis; I, P. melniki sp. nov.; J, P. dichromia sp. nov.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFA8FFFC8EA5FF64D6C3FDFA.taxon	description	(Figs 5 F, 6 F, 7 A, 8 D, 9 E – F, 12 A, 13 H, 14 I)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFA8FFFC8EA5FF64D6C3FDFA.taxon	materials_examined	Material examined. Russia, Leningrad Prov., Roshchino, 25. VI. 1980, V. Tryapitsyn leg., 1 male. Variation. Epomia in pronotal pit distinct to strongly reduced (Fig. 9 E, F). Propodeum with median keel distinctly bifurcate from mid-length or from base. Distribution. Austria, Czech Republic, Finland, * Russia (European part), Slovakia. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFA8FFFC8EA5FD74D369FC4E.taxon	description	(Figs 4 A, 6 E, 10 C, 12 C, 13 E)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFA8FFFC8EA5FD74D369FC4E.taxon	materials_examined	Material examined. Estonia, Torma Distr., 17. VI. 1989, G. Dlusskiy leg., 1 female, 2 males. Moldova, Kishinev [ChiȘinĂu], 11. VI. 1967, V. Tobias leg., 1 male. Ukraine: Odessa Prov., Lisne Vill., Staromanzyrs’kyy Zakaznyk, 13. VI. 1974, D. Kasparyan leg, 1 female; Zaporizhzhia Prov., Melitopol’ Distr., Staroberdyanskoe forestry, 24. V. 1974, D. Kasparyan leg., 1 female, 1 male; Poltava Prov., Myrhorod Distr., 19. V. 1984, A. Kotenko leg., 2 males; Zhytomyr Prov., Korostyshiv Distr., vicinity of Ozerne, 2. VIII. 1980, A. Kotenko leg., 1 male; Kherson Prov., Black Sea Biosphere Reserve, 8. V. 1972, V. Alekseev leg., 2 males (ZMUM). Russia: Kaliningrad Prov., Curonian Spit National Park, 55 ° 09 ′ 16.83 ″ N 20 ° 51 ′ 27.03 ″ E, Rybachiy Vill. ,, 24. VII – 4. VIII. 2006, V. Kolyada leg., 1 female (ZMUM); Novgorod Prov., Pestovo Distr., Tychkino, 11 – 15. VI. 1992, V. Tobias leg., 1 female; Krasnodar Terr., NW of Sochi, Lazarevskoye, 2 – 10. V. 1973, V. Tobias leg., 2 males; Moscow: Krylatskoe, Rublevskiy forest, 1 – 15. VI. 2004, V. Kolyada leg., 65 females, 59 males (ZMUM); same locality, 15 – 30. VI. 2004, V. Kolyada leg., 35 females (ZMUM); same locality, VII. 2004, V. Kolyada leg., 4 females (ZMUM); Bitsa Park, 11. VII. 1993 and 1 – 10. VII. 1994, V. Kolyada leg., 2 females (ZMUM); Losinyy Ostrov, 20. VII. 1993, V. Kolyada leg., 1 female (ZMUM); Moscow Prov., Pavlovskaya Sloboda Vill., 3. VII. 1993, V. Kolyada leg., 2 females (ZMUM); Ul’yanovsk Prov., 3 km N of Ul’yanovsk, 14. VII. 1958, V. Tobias leg., 1 male; Samara Prov.: Kinel’ Distr., vicinity of Domashka, 12. VI. 2012, V. Chemyreva leg., 8 males, 10 females; Samara City, Druzhba Park, 8 – 16. VI. 2002, E. Beloslutsev leg., 2 females; Kirov Prov., Bol’sheromanovo, 4 – 7. VIII. 1994, V. Kolyada leg., 4 males (ZMUM); Primorskiy Terr., Lazovskiy Nature Reserve, 9 – 17. VII. 2008, A. Khalaim leg., 4 females. Variation. Female with A 9 – A 13 weakly transverse to weakly elongate; A 15 as wide as, or distinctly wider than A 14; petiole from transverse to slightly elongate, 1.25 times as long as wide. Male with antenna yellow to dark brown, with keel and emargination on A 3 reaching 0.34 – 0.45 of segment length; petiole 1.8 – 2.2 times as long as wide. Both sexes with radial cell 1.36 – 2.5 times as long as marginal vein; marginal vein 0.86 – 1.44 times as long as distance from it to basal vein; median propodeal keel double throughout, its branches parallel, weakly divergent posteriorly or arcuate, very close to each other or separated by distance equal to length of one-third of propodeum. Distribution. Western, northern and eastern Europe, * Estonia, * Moldova, Russia (European part, * Far East), * Ukraine. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFA8FFFE8D1CFC20D6ACF99A.taxon	description	(Figs 5 G, 6 H, 7 E, 8 C, 9 B, 12 E, 14 J, 15 D, 16 A)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFA8FFFE8D1CFC20D6ACF99A.taxon	materials_examined	Material examined. Estonia: Torma Distr., 17. VI. 1989, G. Dlusskiy leg., 2 males; Vorbuse, 58 ° 25 ′ 31.8 ″ N 26 ° 38 ′ 4.6 ″ E, 16. VI. 2017, V. Soon leg., 1 male (TUZ; no. 102123). Ukraine: Kyiv Prov., Irpin, 27. V. 1984, A. Kotenko leg., 1 male; Kharkiv Prov., Krasnokutsk, 17. VII. 1992, Storozheva leg., 1 female. Russia: Leningrad Prov., Kamenka, 1. VIII. 1972, D. Kasparyan leg., 1 male; Moscow: Krylatskoe, Rublevskiy forest, 1 – 15. VI. 2004, V. Kolyada leg., 8 females (ZMUM); same locality, 15 – 30. VI. 2004, V. Kolyada leg., 4 females (ZMUM); Bitsa Park, 11. VIII. 1993 and 17. VI. 1994, V. Kolyada leg., 2 males (ZMUM); vicinity of Luzhniki, 15. VI. 1947, D. Panfilov leg., 2 males (ZMUM); Moscow Prov., Malakhovka, 10. VI and 26 – 30. VII. 1994, M. Mostovski leg., 5 males (ZMUM); Ryazan’ Prov., Oka Nature Reserve, 17. VI. 1965 [collector unknown], 1 male; Vladimir Prov., 26. VII. 1977 [collector V. G. Chemyreva & V. A. Kolyada. Taxonomy of Acropiesta, Anommatium, Erasikea and Pantolyta Fig. 14. Pantolyta. Venation of fore wing. A, P. dichromia sp. nov.; B, P. nigrocincta; C, P. melniki sp. nov.; D, P. macrocera; E, P. flexinervis; F, P. simplicior sp. nov.; G, P. nitida; H, P. pseudosciarivora; I, P. radialis; J, P. sciarivora. unknown], 1 male; Stavropol’ Prov., Bol’shaya Laba River, 26. VI. 1972, D. Kasparyan leg., 1 male; Kirov Prov., Bol’sheromanovo, 4 – 15. VIII. 1994, V. Kolyada leg., 1 female, 13 males (ZMUM); Chelyabinsk Prov., Il’men’ Nature Reserve, near Il’men’ Lake, 13 – 18. VII. 1958, V. Tobias leg., 2 males; Sverdlovsk Prov., Nizhniy Tagil, VI. 1971, Lebedkina leg., 1 female; Sakhalin Prov.: Sakhalin I., Schmidt Peninsula, 30 km N of Okha Mt., 11. VIII. 2003, A. Lelej leg., 1 female; Shikotan I., Malokuril’skoe, 20 – 21. VIII. 1973, D. Kasparyan leg., 1 male; Primorskiy Terr.: Partizansk Distr., Molchanovka, 1. VII. 1972, V. Kuslitzky leg., 1 male; Khasan Distr., vicinity of Nezhino, 6. VI. 1972, V. Zherikhin leg., 1 female (ZMUM). Azerbaijan, Lenkoran’, 3 – 7. V. 1971, V. Tobias leg., 1 female. Variation. Female with antenna stout to relatively slender (1.25 – 1.45 times as long as head, mesosoma and petiole combined); petiole subquadrate to distinctly elongate. Both sexes with median propodeal keel very variable, from simple to bifurcate or duplicate, branches close to distant from each other; radial cell 1.5 – 2.2 times as long as marginal vein; marginal vein 0.9 – 1.3 times as long as distance from it to basal vein. Distribution. Western, northern and eastern Europe, Azerbaijan, * Russia (European part, West and East Siberia, Far East); Canada, USA. Hosts. Sciara sp. (Diptera: Sciaridae) (Kieffer, 1907). Fig. 15. Pantolyta. A – C, P. flaviventris (A, lateral view, female; B, lateral view, male; C, metasoma, dorsal view, male); D, P. sciarivora (metasoma, dorsal view, male). E, F, P. macrocera (E, lateral view, female; F, antenna, lateral view, male). Scale bars: 1 mm.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFAAFFF98EA5F98AD6D3FE00.taxon	description	(Figs 2 D, 5 H, 6 L, 9 C, 11 A, 13 G)	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFAAFFF98EA5F98AD6D3FE00.taxon	materials_examined	Material examined. Ukraine, Kyiv, Holosiivskyi forest, 27. VI. 1993, A. Kotenko leg., 1 male. Russia: Kaliningrad Prov., Curonian Spit National Park, 55 ° 09 ′ 16.83 ″ N 20 ° 51 ′ 27.03 ″ E, Rybachiy Vill., 24. VII – 4. VIII. 2006, V. Kolyada leg., 2 males, 4 females (ZMUM); Moscow, Bitsa Park, 1 – 10. VI. 1994, V. Kolyada leg., 3 males, 2 females (ZMUM); Moscow Prov., Serpukhov Distr., near Luzhki, 7. VI. 2018, I. Panina leg., 1 male. Variation. Head, mesosoma and petiole yellowish brown to dark brown; propodeum with medi- an keel single or (rarely) weakly bifurcate in distal quarter, strongly projecting to shallow; marginal vein slightly longer to somewhat shorter than distance from it to basal vein; petiole transverse to weakly elongate. Distribution. Austria, Czech Republic, Italy, Poland, * Russia (European part), Ukraine. Hosts. Unknown.	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
03ADA257FFADFFFA8EAAFE6ED2DAF805.taxon	description	(Figs 4 G, 7 D, 10 E, 12 D, 14 F) Holotype. Female; Russia, Sakhalin Prov., Kunashir I., Tret’yakovo, 5. VII. 1973, I. Kerzhner leg. (ZIN). Description. Female. Body length 2.5 mm; fore wing length 2.1 mm; antenna length 1.5 mm. Coloration. Head, mesosoma and metasoma (except petiole) brown; antenna, mandible, venation, legs, petiole and palpi pale brown. Head not nasiform, in dorsal view transverse, 0.77 times as long as wide, distinctly wider than mesosoma, smooth, with few scattered long setae. Temples just behind eyes parallel, broadly receding posteriorly. Occipital flange very narrow, without sculpture. Head in lateral view 1.2 times as high as long. Malar space 0.8 times as long as maximum diameter of eye. Eye pubescent. Face (including antennal shelf) smooth, with long scattered setae. Head in frontal view without mouth conus; genae weakly convex and converging toward mouthparts. Clypeus transverse, 0.67 times as high as wide. Tentorial pits large. Labrum small, semicircular, hardly discernible. Mandibles simple (Fig. 7 D). Antenna slender, weakly broadened apically (Fig. 4 G). Ratios of length to width of A 1 – A 15 in dorsal view: A 1 (48: 8); A 2 (18: 7); A 3 (26: 6); A 4 (18: 6); A 5 (16: 6); A 6 (14: 6); A 7 (14: 6); A 8 (13: 7); A 9 (12: 7); A 10 (12: 8); A 11 (12: 9); A 12 (11: 9); A 13 (11: 9); A 14 (11: 9); A 15 (20: 9). Mesosoma slightly compressed, 1.9 times as high as wide, in dorsal view 1.82 times as long as wide. Sides of pronotum smooth with few scattered long setae (setae denser on pronotal collar); epomia strongly reduced; pronotal pit bare and smooth (Fig. 10 E). Mesoscutum as long as wide, convex, with few scattered semierect setae. Notauli deep and complete, not sculptured. Anterior scutellar pit deep, transverse, 0.55 times as long as wide. Scutellum large, widened posteriorly. Axillar depression weakly pubescent, with two verriculate tubercles posteriorly (Fig. 12 D). Fig. 16. Pantolyta, male. Fore tibia. A, P. sciarivora; B, P. flaviventris. Mesopleuron scarcely pubescent, subalar bridge posterodorsally and epicnemial bridge anteroventrally strongly reduced; epicnemial pit deep and pubescent; mesopleural pit developed as a shallow depression (Fig. 10 E). Metascutellum narrow, smooth and weakly pubescent; dorsellum distinct, with three short longitudinal keels. Propodeum transverse, 0.53 times as long as wide, pubescent laterally; median propodeal keel strongly reduced; upper and lower posterior propodeal projections forming very small teeth (Fig. 12 D). Sides of propodeum entirely pubescent, with one longitudinal keel. Legs slender. Fore wing with closed radial cell (Fig. 14 F); distance from marginal vein to basal vein / marginal vein length / radial cell length as 10 / 10 / 17. Petiole of metasoma cylindrical, 1.17 times as long as wide, with weakly visible striate sculpture and few scattered setae laterally; petiole ventrally with several bunches of setae; posterior margin of petiole not arcuate (Fig. 12 D). T 2 anteriorly with short and hardly visible striation, median groove the longest; T 3 – T 6 narrow, with few scattered setae laterally; T 7 and T 8 long, strongly compressed laterally. S 2 with short grooves at base and with numerous erect scattered setae; S 3 – S 5 narrow, with scattered setae; S 6 distinctly elongate and compressed, with few scattered setae. Male. Unknown. Comparison. Pantolyta simplicior sp. nov. can be distinguished from all known species of Pantolyta by the combination of the following characters: the head is not nasiform and has no mouth conus (Fig. 7 D); the genae are convex in frontal view; the female antenna is slender, weakly broadened apically, with A 3 – A 7 more than twice as long as wide (Fig. 4 G); pronotal pit is bare and smooth; subalar and epicnemial bridges on mesopleuron are strongly reduced; the mesopleural pit is as a shallow depression (Fig. 10 E); median propodeal keel is strongly reduced; upper and lower posterior propodeal projections form very small lateral teeth (Fig. 12 D); the fore wing has a closed radial cell, marginal vein as long as the distance from marginal vein to basal vein and distinctly shorter than radial cell; the petiole is weakly elongate. Etymology. The species name is the Latin adjective simplicior (simpler). Distribution. Russia (Kunashir Island). Host. Unknown. A key to the Palaearctic species of Pantolyta and Anommatium Note. In addition to 15 species considered in this paper, the key includes the following seven species considered in Chemyreva & Kolyada (2019): Pantolyta atrata Förster, 1861, P. elegans Chemyreva et Kolyada, 2019, P. hadrosoma Macek, 1993, P. marginalis Kieffer, 1909, P. nixoni Macek, 1993, P. pallida Kieffer, 1908, and P. stylata Macek, 1993. Females	en	Chemyreva, V. G., Kolyada, V. A. (2021): Taxonomy of the genera Acropiesta, Anommatium, Erasikea and Pantolyta (Diapriidae: Belytinae) with review of species occurring in Russia. Zoosystematica Rossica (China) 30 (1): 137-162, DOI: 10.31610/zsr/2021.30.1.137, URL: https://doi.org/10.31610/zsr/2021.30.1.137
